Quaternary — Open Access Journal

At 37° 24′ N 22° 8′ E, the Megalopolis Basin lies in the central Peloponnese Peninsula, southwestern Greece. In the Megalopolis Basin at ~ 350 m amsl, the Paleolithic site, Marathousa 1, sits within a palustrine/lacustrine clastic package between Lignite Seams III and II, that both likely correlate with interglacial periods. At Marathousa 1, immediately below Lignite Seam III, lies a clayey-silty sand layer with a horizon rich in molluscs ranging from ~ 20–40 cm thick. About 0.8–1.3 m below the shell-rich horizon (SRH), lacustrine silty to muddy sands rich in organic matter yielded Paleolithic lithic artefacts associated with Middle Pleistocene fauna, some with cut marks and possible bone knapping, found within palustrine/lacustrine clastic deposits. Since ESR (electron spin resonance) can date teeth and molluscs aged > 2 Ma, two bivalve samples, AM66 and AM65, five subsamples from a cervid molar, AT39, and one subsample from another cervid molar, AT68, were independently dated by ESR from Marathousa 1. To calculate the ages, time-averaged cosmic and time- and volumetrically-averaged sedimentary dose rates were calculated using past water depths and sedimentation rates as determined from paleontological and geological criteria. Found in the SRH in Layer UA2, AM66 and AM65 averaged 488 ± 37 ka, which correlates with MIS 13a. Because the bivalves sat stratigraphically above the artefacts and mammalian fossils, their ages constrain the ESR ages for the teeth deposited below. Lying on the unconformity at the base of Layer UA3c with UA4, and its correlative unconformity at the Layer UB4c/UB5 boundary, sat the dated teeth from large mammals. Because the bones in the Palaeoloxodonantiquus skeleton lay in quasi-anatomical association, the likelihood for fossil reworking on the Layer UB3c/UB4 surface is low. Isochron analysis suggests that using a U uptake model with p = 2 provides the most accurate ages for AT39. With p = 2, AT39 dates to 503 ± 13 ka, while AT68 dates to 512 ± 34 ka. Nonetheless, two to three more teeth and molluscs should be dated to confirm these ages, when more samples suitable for ESR dating are found. Both tooth ages correlate well with early MIS 13, an interglacial period with cooler mean global temperatures compared to MIS 11 or 9. Assuming that the archaeological site formed in one event, rather than as a palimpsest, the data suggest that hominins processed elephant and other faunal carcasses along the shores of a shallow lake or marsh in the Megalopolis Basin at 503 ± 12 ka. Between the two horizons dated here, their sedimentation rate averaged 4.8 ± 1.8 to 7.8 ± 2.9 cm/ka. Full article

Rivers are central to debate about the Anthropocene because many human activities from antiquity focused on channels and floodplains. A literature compilation for the onset of human modification of rivers identifies six stages that represent key innovations focused in the Near East and adjoining areas: (1) minimal effects before about 15,000 cal yr BP, with the use of fire and gathering of plants and aquatic resources; (2) minor effects from increased cultivation after about 15,000 cal yr BP, with plant and animal domestication after about 10,700 cal yr BP; (3) agricultural era after about 9800 cal yr BP, with legacy sediments, widespread fire use, the first dams and irrigation, and mud-brick manufacture; (4) irrigation era from about 6500 cal yr BP, with large-scale irrigation, major cities, the first large dam, urban water supplies, expanded groundwater use, river fleets, and alluvial mining; (5) engineering era with embankments, dams, and watermills after about 3000 cal yr BP, especially in the Chinese and Roman empires; and (6) technological era after about 1800 CE. Anthropogenic river effects were more varied and intense than commonly has been recognised, and they should be considered routinely in interpreting Late Pleistocene and Holocene fluvial archives. Full article

Accurate species identification in fossil remains is a complex task but is a key component for developing good inferences on many, if not all, fundamental questions in macroecology and macroevolution. In the Quaternary, arvicolines are very abundant remains in archeological and paleontological sites in Western Europe and their identification is often based on the first lower molar. The common vole Microtus arvalis (Pallas, 1778) and the field vole Microtus agrestis (Linnaeus, 1761) are commonly found in those deposits. These two species are genetically and ecologically divergent. Nonetheless, their lower molars, on which species identification is done, exhibit a large morphological variation that can potentially lead to some confusion and misinterpretation. Moreover, molecular data suggest that present-day M. agrestis populations are a complex of divergent lineages, some of them being recognized nowadays as valid species. On the basis of extant populations representing a large part of the present-day geographical distribution of these two species, we developed a classification model based on geometric morphometrics of the first lower molar. Our statistical model was then applied on four fossil sites selected to evaluate the relevance of taxonomic determination found in species lists. The model using landmarks describing the overall shape of the first lower molar classifies the two species with the smallest prediction error together with very high individual posterior probabilities. The obtained classification is much better than those arising from shapes of any specific molar part such as the anterior loop, asymmetry or peculiar triangle shape. Discrepancies with expert classification on fossils suggest that existing faunal lists should always be considered cautiously for these two species. Our morphometric model provides a first step towards a rationalized way of revising past collections and expertise for future small mammal assemblages. It will thus help us better understand the paleobiogeographical expansion of these two key species in Quaternary faunas. Full article

Coulet des Roches is a natural karst trap in Southern France. Its infilling dates back to the end of the Pleniglacial (Last Glacial Maximum, LGM) and the end of the Tardiglacial (Last Glacial, LG). Three mustelid species have been identified in this infilling: the common polecat (Mustela putorius, minimum number of individuals (MNI) = 4), the stoat (Mustela erminea, MNI = 14) and the weasel (Mustela nivalis, MNI = 48). The common polecat remains are metrically and morphologically indistinguishable from recent European specimens. The smallest mustelids are mainly represented by average-sized specimens, which are slightly smaller than extant species. A partial weasel skeleton of an extremely small pygmy weasel, regarded as a typical glacial element, was also discovered. Sexual dimorphism is strongly pronounced. Seventeen horses have been identified, corresponding to the chrono subspecies Equusferusgallicus. The analysis of the muzzles and metapodials shows overall adaptation to cool and dry weather conditions. The ibexes are typical of Capra ibex (MNI = 15). The p3 morphology is similar to the LGM populations located on the southern side of the Durance River, with an important dilation of the metaconid, except for the oldest LGM specimen. This dilation is older on the southern side of the Durance River, as it occurs at the end of MIS 3. This difference could reflect the barrier role of the Durance River. Micromammals are abundant (mostly related to rodents and shrews; 18 genera/species; MNI = 470). The paleoecological study highlighted important and rapid climatic and environmental fluctuations throughout the sequence. As a result of climatic fluctuations, the plains constituted a corridor for the migration of temperate species to Provence during cold periods (“southern refuge zone”) and their re-immigration to Western Europe during temperate episodes. In a related and complementary way, the highland areas not only played a natural role as a geographical barrier, but also constituted a refuge zone during interglacial episodes for some micromammal species originating from northern and eastern parts of Europe (“cryptic southern refugia”). Full article

The article attests the presence of the genus Rucervus in the paleontological record of Europe and presents the description of new species of large-sized deer Rucervus radulescui sp. nov. from the Early Pleistocene of Valea Grăunceanului (Southern Romania) and Rucervus gigans sp. nov. from the late Early Pleistocene of Apollonia-1 (Greece). The described cervid species represent two different evolutionary radiations of Rucervus that are grouped into the extinct subgenus Arvernoceros that represents the northern evolutionary radiation and the nominotypical subgenus that is regarded as the southern evolutionary radiation and represented today by only one species Rucervus duvaucelii. The evolutionary radiation and dispersals of Rucervus are regarded in the paleobiogeographic context of faunal exchanges between southeastern Europe, Caucasus, and Near East during the Early Pleistocene and the westward dispersal of early hominins in Eurasia. Full article

Alluvial-fan successions record changes in hydrological processes and environments that may reflect tectonic activity, climate conditions and changes, intrinsic geomorphic changes, or combinations of these factors. Here, we focus on the evolution of a stream-dominated fan in a tectonic depression of the Xining basin of China, laid down under a semi-arid climate in the northeastern Tibetan Plateau (NETP). The fan succession is composed of three facies associations, from bottom to top: (1) matrix to clast-supported, poorly sorted, planar cross-stratified to crudely stratified sheets of coarse-grained sediments; (2) horizontal laminated sand, laminated layers of reddish fine silt and yellow coarse silt with stacked mounds of sand; and (3) clay-rich deposits with incipient paleosols. The succession shows rapid sediment aggradation from high-energy to low-energy alluvial fans and finally to a floodplain. The dating results using optically stimulated luminescence (OSL) method show that a gravelly, high-energy fan was deposited during MIS 6, after which a low-energy fan, mainly composed of sand and silt, was deposited and finally covered by flood loam during the MIS 6–5 transition and the warmer last interglacial. Stacked sand mounds are interpreted from their sediment structure and grain-size distribution as shrub-coppice dunes in low-energy fan deposits. They may be considered as a response to the interaction of alluvial and aeolian processes in a semi-arid environment. Full article

Collisional mountain belts commonly develop intramontane basins from mechanical and isostatic subsidence during orogenic development. These frequently display a relict top surface, evidencing a change interval from basin infilling to erosion often via capture or overspill. Such surfaces provide markers that inform on orogenic growth patterns via climate and base level interplay. Here, we describe the top surface from the Sorbas Basin, a key intramontane basin within the Betic Cordillera (SE Spain). The surface is fragmentary comprising high elevation hilltops and discontinuous ridges developed onto the variably deformed final basin infill outcrop (Gochar Formation). We reconstruct surface configuration using DEM interpolation and apply ¹⁰Be/²⁶Al cosmonuclides to assess surface formation timing. The surface is a degraded Early Pleistocene erosional pediment developed via autogenic switching of alluvial fan streams under stable dryland climate and base level conditions. Base-level lowering since the Middle Pleistocene focused headwards incision up interfan drainages, culminating in fan head capture and fan morphological preservation within the abandoned surface. Post abandonment erosion has lowered the basin surface by 31 m (average) and removed ~5.95 km³ of fill. Regional basin comparisons reveal a phase of Early Pleistocene surface formation, marking landscape stability following the most recent Pliocene-Early Pleistocene mountain building. Post-surface erosion rate quantification is low and in accordance with ¹⁰Be denudation rates typical of the low uplift Betic Cordillera. Full article

External impact on the development of fluvial systems is generally exerted by changes in sea level, climate and tectonic movements. In this study, it is shown that a regional to local differentiation of fluvial response may be caused by semi-direct effects of climate change and tectonic movement; for example, vegetation cover, frozen soil, snow cover and longitudinal gradient. Such semi-direct effects may be responsible for specific fluvial activity resulting in specific drainage patterns, sedimentation series and erosion–accumulation rates. These conclusions are exemplified by the study of the fluvial archives of the Tis(z)a catchment in the Pannonian Basin in Hungary and Serbia from the middle of the last glacial to the Pleistocene–Holocene transition. Previous investigations in that catchment are supplemented here by new geomorphological–sedimentological data and OSL-dating. Specific characteristics of this catchment in comparison with other regions are the preponderance of meandering systems during the last glacial and the presence of very large meanders in given time intervals. Full article

(1) Background: Over the last decades, important fossil records of Late Pliocene mammals and reptiles have been unearthed in Milia, Grevena (N Greece). This assemblage shows a remarkable composition and diversity, including the partial skeletons of mastodons that reached record-breaking sizes, abundant rhinos—the first occurrence of this species in Greece—and some new species; (2) Methods: We perform a thorough quantitative analysis of the recovered assemblage, presenting the information of the various collection spots in Milia, and calculate various biodiversity indices for each spot; (3) Results: Our research has revealed the vast majority of expected taxa in the assemblage. We argue that the various sub-localities in Milia could be grouped into a larger, composite assemblage representing a short period. We analyze the diversity changes through the various localities in Milia and highlight potential barriers that could affect the distribution of taxa; (4) Conclusions: We re-affirm the Early Villafranchian affinities of the fauna. In particular, Milia should date at MN16a, with a surprising presence of some more archaic, Ruscinian taxa. The fossils of Milia depict a Late Pliocene paradise in the Southern Balkans; a paradise, unfortunately, lost with the onset of the dramatic climate changes of the Quaternary. Full article

Recently collected fossil material in the Villafranchian locality of Sésklo, as well as a re-evaluation of a pre-existing, partly-published museum collection, allow the recognition of a lower faunal level in the locality, older than the main Equus-dominated fossil assemblage, dated in the Early Pleistocene (MNQ17). The lower level yielded, instead, an advanced hipparion, referred to the species Plesiohipparion cf. shanxiense, and a small number of associated taxa: an ostrich (Struthio cf. chersonensis), an unidentified proboscidean, the pig Sus arvernensis, two antelopes (Gazella cf. bouvrainae and Gazellospira torticornis), a large bovid (Bovini indet.), and a rhinoceros (Stephanorhinus sp.). The lower-level fauna is dated in the latest Pliocene (MN16) and indicates a rather open and dry palaeoenvironment. The faunal sequence in Sésklo shows that the hipparion did not co-occur with the stenonid horse, at least in this region. Previous reports on sympatry of these taxa may result from faunal mixing, requiring re-examination of the available samples. Full article

Eight anatomically and taxonomically different finds are presented in this paper, and they belong to four taxa: woolly mammoth (Mammuthus primigenius), giant deer (Megaloceros giganteus), red deer (Cervus elaphus), and dog (Canis familiaris). All specimens represent allochthonous Late Pleistocene and Holocene animal remains, and all were dredged during the gravel exploitation at the Sekuline site near Molve (Podravina region, SW Pannonian basin, NE Croatia). Mammoth remains (bone and tusk fragments) were radiocarbon dated, and these are the first absolute dates on mammoths in Croatia. One upper last left deciduous premolar (dP4 sin.) also belongs to the same species. Ascribed to a dog is one well-preserved skull with a peculiar abscess scar on the maxillary bone as the result of an inflammatory process on the carnassial (P4) premolar. The Late Pleistocene cervid remains are giant deer, while the other cervid finds were determined to be red deer of the Holocene age. Morphometrical and taphonomical data are presented for each specimen. Such fossil and recent bone/tooth aggregates are characteristic of fluvial deposits and selective collecting. Although lacking stratigraphic provenance, these finds help to fulfil the gaps in palaeoenvironmental, palaeoecological, and palaeoclimate reconstructions of Podravina and its neighbouring areas. Full article

Although the interpretation of Quaternary records of interrelated environmental–ecological–human processes is necessarily complex, it is often addressed using too-simple deterministic approaches. This paper suggests a holistic framework called EHLFS (Environmental–Human–Landscape Feedbacks and Synergies) to tackle Quaternary complexity. The EHLFS scheme is a multiple-working-hypotheses framework, able to account for the particular nature of Quaternary research, and is used in combination with the strong inference method of hypothesis testing. The resulting system is called the strong fuzzy EHLFS approach. This approach is explained in some detail and compared with the more extended simplistic determinisms—namely the environmental determinism and the human determinism—as well as with dual determinisms or deterministic approaches based on two contrasting and apparently contradictory and excluding hypotheses or theories. The application of the strong EHLFS methodology is illustrated using the Late Holocene ecological and cultural history of Easter Island since its initial human settlement, a topic that has traditionally been addressed using simplistic and dual deterministic approaches. The strong fuzzy EHLFS approach seems to be a robust framework to address past complex issues where environment, humans and landscape interact, as well as an open system able to encompass new challenging evidence and thorough changes in fundamental research questions. Full article

Small mammal remains obtained from the European localities dated to the Eemian (Mikulino) age have been analyzed for the first time at a regional scale based on the present biogeographical regionalization of Europe. The regional faunas dated to the warm interval in the first part of the Late Pleistocene display notable differences in fauna composition, species richness, and diversity indices. The classification of regional faunal assemblages revealed distinctive features of small mammal faunas in Eastern and Western Europe during the Eemian (=Mikulino, =Ipswichian) Interglacial. Faunas of the Iberian Peninsula, Apennine Peninsula, and Sardinia Island appear to deviate from the other regions. In the Eemian Interglacial, the maximum species richness of small mammals (≥40 species) with a relatively high proportion of typical forest species was recorded in Western and Central Europe and in the western part of Eastern Europe. The lowest species richness (5–14 species) was typical of island faunas and of those in the north of Eastern Europe. The data obtained make it possible to reconstruct the distribution of forest biotopes and open habitats (forest-steppe and steppe) in various regions of Europe. Noteworthy is a limited area of forests in the south and in the northeastern part of Europe. In these regions, it seems likely that under conditions of relatively high temperatures characteristic of the Last Interglacial and an insufficient moisture supply there could exist open forest stands or forest-steppe landscapes, as suggested by the presence of species indicative of forest-steppe and steppe north of the forest zone. The results obtained are useful in modeling changes in the mammal faunas as well as environmental changes in entire Europe due to global climatic changes (including the global warming recorded at present). Full article

The Late Glacial period is characterized by slow warming, punctuated by short, cold episodes, such as the Younger Dryas (i.e., GS1). The impact of this climatic event on the mammal community is still poorly documented in southwestern France. Here, a new radiocarbon date obtained directly on fossil remains of common hamster, Cricetus cricetus, confirms its presence in southwestern France during the Younger Dryas (GS1). This observation currently suggests that C. cricetus could be an accurate chronological indicator of this event in southwestern France. In this particular case, it also demonstrates an attritional death, polluting the deposit, these remains having been found in the Combe-Cullier layer, attributed to an earlier period. Full article

Tusks of very young mammoths and other proboscideans are extremely rare in the fossil record. This article presents all deciduous tusks and newly-developing permanent tusks of woolly mammoths which are known from Dutch localities and the North Sea. Four deciduous tusks are from The Netherlands and one is from Germany. Five specimens of newly-developing permanent tusks are also discussed. In this article, we present several characteristics and details of those tusks such as presence/absence of roots, and cementum and enamel layers that should facilitate an identification when new findings are made. We also describe a dental element, a digitelle, that is easily mistaken for a tusk of a juvenile woolly mammoth. Full article

During the last field campaigns in the mammal fossiliferous site Apollonia 1 (Macedonia, Greece), new carnivoran material has been discovered. The new collection added two new carnivoran taxa, Homotherium latidens and Panthera gombaszögensis. The new canid material and the revision of the old one (a) suggest the presence of two Canis species, C. etruscus and C. apolloniensis; (b) confirm the presence of the hypercarnivore Lycaon lycaonoides, and (c) allow for re-classifying the vulpine material to Vulpes praeglacialis. The taxonomic status of the species C. apolloniensis and Meles dimitrius is discussed. The composition and diversity of the Apollonia carnivoran assemblage are estimated and compared to those of various Greek and European Villafranchian ones. The results suggest close similarity to the Venta Micena (Spain) and Dmanisi (Georgia) carnivoran assemblages. The biochronological evidence indicates that Apollonia 1 is younger than Venta Micena and older than Untermassfeld (Germany), suggesting an age of 1.3–1.0 Ma. The study of the carnivoran guild structure of Apollonia 1 in comparison to the modern ones from known environments, as well as their functional morphology, suggest an open habitat, agreeing with previous interpretations. Full article

The analysis of the cervid fossil remains from the late Pleistocene fossiliferous deposit Lagoa dos Porcos (in the region of the National Park “Serra da Capivara”, Piauí, Brazil) proves the presence of at least two species: a small deer, belonging to the genus Mazama, and a larger one (Morenelaphus sp.). The latter taxon is recognized for the first time not only in this area, but in the whole Piauí State, enlarging the paleogeographic distribution of the genus. This study also points out the difference between the cervid fauna found in the karstic area of Park and Lagoa dos Porcos, which lies in the alluvial plain. Full article