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Fossil Studies

Fossil Studies - formerly Fossils - is an international, peer-reviewed, open access journal on all aspects of palaeontology published quarterly online by MDPI.

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All Articles (71)

  • Communication
  • Open Access

First Record of Penelope sp. (Aves, Cracidae) in the Late Pleistocene of Northern Uruguay

  • Washington W. Jones,
  • Andrés Rinderknecht and
  • Martín Ubilla
  • + 3 authors

We report the first fossil record of the family Cracidae in Uruguay from the Sopas Formation of the Late Pleistocene (Paso del Buey Negro locality, Arapey Chico River, Salto Department). The ulnar morphology of the genus Penelope does not allow definitive species-level identification. However, on the basis of the current geographic distribution, linear dimensions, articular features, and muscle impressions, it is highly likely that this is a specimen of Penelope obscura. At Paso del Buey Negro, numerous mammalian fossils have been recorded, including rodents, perissodactyls, artiodactyls, and xenarthrans, as well as freshwater turtles. This locality is thought to have had a heterogeneous environment with open and forested areas associated with watercourses and a temperate to subtropical climate similar to the region’s current climate. The fossil record of the genus Penelope is significant because these birds are clearly dependent on humid forest, gallery forest, and forest-edge environments, generally at low altitudes, and therefore they are proper paleoenvironmental proxies.

Foss. Stud.

6 November 2026

Pleistocene records of the genus Penelope in South America. (1) Talara tar seeps, northern Peru, Penelope cf. purpurascens [17]; (2) Penelope sp. Gruta dos Brejoes, Bahia, Brazil [14]; (3) Penelope sp. Lapa da Escrivania, Minas Gerais, Brazil [11]; (4) Penelope cf. obscura, Paso del Buey Negro, northern Uruguay. Current distribution of Penelope obscura in purple; modified from Winkler et al. [7]. Inset right: map of the locality Paso del Buey Negro, Salto Department, Uruguay.
  • Article
  • Open Access

The role of vertebrate fossils in the origin of ancient Greek myths remains an intriguing and open question. A comparative paleontological and historical assessment between Aegean Greece and Magna Graecia (Southern Italy) is presented, focusing on the Late Miocene giraffid Samotherium. The Genus is characteristic of the Greco-Iranian biogeographic province, also recorded outside its primary range in Calabria. This region was part of Magna Graecia, the extensive Southern Italian territory settled by Greek colonists who established flourishing cities from the 8th century BC onward. Whereas specific Late Miocene fossil sites in Greece are linked to geomyths, such as the bone-rich red beds of Samos associated with the monstruous Neades, no such attributions are recorded in Calabria. Although the coastline near the Greek colony of Hipponion, the modern city of Vibo Valentia, is known as the “Costa degli Dei” (Coast of the Gods), vertebrate fossils are not associated with ancient myth, whereas invertebrate fossils supported local folk traditions. These discrepancies between Greece and Calabria are likely driven by contrasting taphonomic contexts and differences in fossil exposure, discoverability, and skeletal element representation.

Foss. Stud.

2 November 2026

Geographical position of the sites quoted in the text and the respective national capital cities.
  • Feature Paper
  • Article
  • Open Access

Newly discovered theropod footprints are described from collapsed rock blocks derived from the Lower Cretaceous Guantou Formation at the Shangantian tracksite in Dongyang City, Zhejiang Province, eastern China. The specimens are preserved in at least 18 natural molds on a single siltstone slab. The footprint assemblage is composed of minute-sized tridactyl tracks with lengths ranging between 6.8 cm and 8.8 cm. Morphologically, these tracks exhibit a relatively wide divarication angle between digits II and IV, a low length/width ratio, weak mesaxony, and a prominent, elongate metatarsophalangeal pad associated with digit IV aligned along the long axis of digit III. Based on these diagnostic traits, the new tracks are tentatively assigned to Jialingpus isp. The reduced mesaxony and incomplete expression of the medial metatarsophalangeal area behind digit II represent morphometric and preservation-driven variations characteristic of Cretaceous Jialingpus material. Kinematic estimates from three distinct single-step trackways indicate that the trackmakers were moving at slow walking speeds of 0.4–0.8 m/s (1.4–2.9 km/h). This discovery enriches the mid-Cretaceous dinosaur track diversity in Zhejiang Province and provides important ichnological insights into the evolution and morphometric variation of Jialingpus across the Jurassic–Cretaceous boundary.

Foss. Stud.

20 October 2026

Geographic location map of the Shangantian tracksite.
  • Article
  • Open Access

Reported trace fossils from the Clifton Formation, New Brunswick, Canada, are limited to a single occurrence of a temnospondyl trackway and invertebrate trackways identified in the Pennsylvanian continental strata. Our study builds on this with a description of a horizontal burrow interpreted to have been constructed by a fossorial tetrapod. Historically, limited focus has been placed on tetrapod burrow activity and its ichnotaxonomy, so our findings also have broader implications. We report on the first horizontal burrow collected from the Clifton Formation near the township of Stonehaven, with unique features that include transversely oriented angular mudstone intraclasts without a meniscate backfill. We propose a new ichnotaxon, Thrasmaichnus hindsi igen. et isp. nov., and interpret the trace as having been made by a fossorial tetrapod. Thrasmaichnus hindsi is the first described Pennsylvanian-age tetrapod burrow, whereas Pennsylvanian-age fossorial tetrapods are currently only known from body fossils.

Foss. Stud.

10 October 2026

(A) Map of Canada with New Brunswick highlighted in green; (B) map of the Maritimes Basin of Atlantic Canada highlighted in brown; study area denoted with a red box; (C) geological map of north-eastern New Brunswick. The study area at Clifton is highlighted with a red box; (D) orthographic drone image of the study site with sandstone-channel bodies annotated and sandstone-trace-fossil horizon denoted.

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Foss. Stud. - ISSN 2813-6284