Navigating the Blue Transition: A Dynamic Simulation of the Coupling and Coordination in China’s Marine Fishery Ecological–Social–Economic System
Abstract
1. Introduction
2. Conceptual Framework
2.1. Coupling Mechanism of the Economic–Social System
2.2. Coupling Mechanism of the Ecological–Social System
2.3. Coupling Mechanism of the Ecological–Economic System
3. Materials and Methods
3.1. Indicator System and Data Sources
3.2. Carbon-Emission and Biological-Carbon Accounting Procedures
3.3. Coupling-Coordination Degree Evaluation Model
3.4. System-Dynamics Model
3.4.1. System Boundaries and Assumptions
3.4.2. Causal-Loop Diagram of the System
3.4.3. Model Construction, Historical-Behavior Check, and Scenario Setting
4. Results
4.1. Historical Coupling-Coordination Patterns
4.2. Scenario Results
5. General Discussion
5.1. Summary of Findings
5.2. Policy Implications
5.3. Limitations and Future Research
Author Contributions
Funding
Data Availability Statement
Conflicts of Interest
Appendix A. Stock-and-Flow Diagram

References
- Gao, Y.; Fu, Z.; Yang, J.; Yu, M.; Wang, W. Spatial–Temporal Differentiation and Influencing Factors of Marine Fishery Carbon Emission Efficiency in China. Environ. Dev. Sustain. 2024, 26, 453–478. [Google Scholar] [CrossRef] [Scilit]
- Wang, Y.; Khan, S.U.; Kong, D.; Xu, R.; Zhang, Y. Navigating the Dual Currents: Advanced Insights into Carbon Emissions and Economic Growth in China’s Coastal Marine Fisheries. Mar. Pol. 2025, 177, 106671. [Google Scholar] [CrossRef] [Scilit]
- Kong, F.; Cui, W. Ecological Sustainability of Marine Fishery in Coastal Countries of the “Belt and Road”: Spatial–Temporal Features and Future Predictions. Environ. Dev. Sustain. 2026, 28, 12845–12871. [Google Scholar] [CrossRef] [Scilit]
- Thrush, S.F.; Hewitt, J.E.; Gladstone-Gallagher, R.V.; Savage, C.; Lundquist, C.; O’Meara, T.; Vieillard, A.; Hillman, J.R.; Mangan, S.; Douglas, E.J.; et al. Cumulative Stressors Reduce the Self-regulating Capacity of Coastal Ecosystems. Ecol. Appl. 2021, 31, e02223. [Google Scholar] [CrossRef] [Scilit] [PubMed]
- Alsaleh, M.; Abdul-Rahim, A.S. What Are the Influence of Fishery Activities and Their Implication on Marine Water Pollution? An Empirical Analysis. Environ. Dev. Sustain. 2026, 28, 15063–15094. [Google Scholar] [CrossRef] [Scilit]
- Song, M.; Fatima, S.; Ullah, E.; Haseeb, M.; Hossain, M.E. Environmental Impacts of Aquaculture, Marine Shipping, and Blue R&D in Nordic Countries. Environ. Dev. Sustain. 2025; early access. [CrossRef] [Scilit]
- Greer, K.; Zeller, D.; Woroniak, J.; Coulter, A.; Winchester, M.; Palomares, M.L.D.; Pauly, D. Global Trends in Carbon Dioxide (CO2) Emissions from Fuel Combustion in Marine Fisheries from 1950 to 2016. Mar. Policy 2019, 107, 103382. [Google Scholar] [CrossRef] [Scilit]
- Guan, H.; Chen, Y.; Zhao, A. Carbon Neutrality Assessment and Driving Factor Analysis of China’s Offshore Fishing Industry. Water 2022, 14, 4112. [Google Scholar] [CrossRef] [Scilit]
- Jia, D.; Liu, X.; Guan, X.; Guo, J.; Zhang, S.; Li, H.; Jin, Y.; Sun, J. Spatio-temporal Differences and Simulation Studies of the Carbon Budget from Fisheries in the Northern Marine Economic Circle of China. Front. Mar. Sci. 2024, 11, 1393659. [Google Scholar] [CrossRef] [Scilit]
- Feng, C.; Ye, G.; Jiang, Q.; Zheng, Y.; Chen, G.; Wu, J.; Feng, X.; Si, Y.; Zeng, J.; Li, P.; et al. The Contribution of Ocean-Based Solutions to Carbon Reduction in China. Sci. Total Environ. 2021, 797, 149168. [Google Scholar] [CrossRef] [Scilit] [PubMed]
- Liu, F.; Huang, Y.; Zhang, L.; Li, G. Marine Environmental Pollution, Aquatic Products Trade and Marine Fishery Economy—An Empirical Analysis Based on Simultaneous Equation Model. Ocean Coast. Manag. 2022, 222, 106096. [Google Scholar] [CrossRef] [Scilit]
- Phan, K.-T.; Hsu, Y.-L.; Chen, S.-H. Do Sustainable Development Goals (SDGs) Boost Green Productivity in National Marine Fisheries? International Evidence. Mar. Coast. Fish. 2024, 16, e10322. [Google Scholar] [CrossRef] [Scilit]
- Ren, W. Study on the Removable Carbon Sink Estimation and Decomposition of Influencing Factors of Mariculture Shellfish and Algae in China—A Two-Dimensional Perspective Based on Scale and Structure. Environ. Sci. Pollut. Res. 2021, 28, 21528–21539. [Google Scholar] [CrossRef] [Scilit] [PubMed]
- Williams, C.; Rees, S.; Sheehan, E.V.; Ashley, M.; Davies, W. Rewilding the Sea? A Rapid, Low Cost Model for Valuing the Ecosystem Service Benefits of Kelp Forest Recovery Based on Existing Valuations and Benefit Transfers. Front. Ecol. Evol. 2022, 10, 642775. [Google Scholar] [CrossRef] [Scilit]
- Le, J.; Wei, Y. Green Efficiency Measurement of Seaweed Culture in China under the Double Carbon Target. Sustainability 2023, 15, 7683. [Google Scholar] [CrossRef] [Scilit]
- Li, G.; Tan, C.; Zhang, W.; Zheng, W.; Liu, Y. Carbon Emission Efficiency, Technological Progress, and Fishery Scale Expansion: Evidence from Marine Fishery in China. Sustainability 2023, 15, 6331. [Google Scholar] [CrossRef] [Scilit]
- Zhang, Y.; Khan, S.U.; Wang, Y. The Future Is Digital: Can the Digital Economy Drive Marine Sustainability? Exploring Regional Impacts on Fisheries’ Carbon Emissions in Coastal China. J. Clean. Prod. 2025, 506, 145518. [Google Scholar] [CrossRef] [Scilit]
- Aranda-Garrido, N.; Martínez-Martínez, P.; Antón-Linares, I.; Abel-Abellán, I.; Encabo-Lucena, S.; Arroyo-Martínez, E.; Trives-Escudero, M.; Barberá-Cebrián, C.; Giménez-Casalduero, F. Profile of Marine Sport Fishers and Interannual Variation in Coastal Catches in Southeastern Spain. Fishes 2026, 11, 402. [Google Scholar] [CrossRef] [Scilit]
- Chen, X.; Di, Q.; Hou, Z.; Yu, Z. Measurement of Carbon Emissions from Marine Fisheries and System Dynamics Simulation Analysis: China’s Northern Marine Economic Zone Case. Mar. Policy 2022, 145, 105279. [Google Scholar] [CrossRef] [Scilit]
- Wang, Y.; Yang, Y.; Ren, Q. Comprehensive Estimation and Spatiotemporal Differentiation of Marine Fishery Carbon Sinks in China’s Coastal Provinces. Ocean Coast. Manag. 2025, 269, 107846. [Google Scholar] [CrossRef] [Scilit]
- He, L.; Du, X.; Zhao, J.; Chen, H. Exploring the Coupling Coordination Relationship of Water Resources, Socio-Economy and Eco-Environment in China. Sci. Total Environ. 2024, 918, 170705. [Google Scholar] [CrossRef] [Scilit] [PubMed]
- Wang, N.; Li, J.-M.; Zhou, Y.-F. Mechanism of Action of Marine Ecological Restoration on Ecological, Economic, and Social Benefits—An Empirical Analysis Based on a Structural Equation Model. Ocean Coast. Manag. 2024, 248, 106950. [Google Scholar] [CrossRef] [Scilit]
- Zhang, J.; Chen, J.; Gao, G.; Lv, M. Decoding Carbon Emissions in China’s Marine Fisheries: Trends, Drivers, and Pathways to Sustainability. J. Clean. Prod. 2025, 520, 146101. [Google Scholar] [CrossRef] [Scilit]
- Wang, B.; Han, L.; Zhang, H. The Impact of Regional Industrial Structure Upgrading on the Economic Growth of Marine Fisheries in China—The Perspective of Industrial Structure Advancement and Rationalization. Front. Mar. Sci. 2021, 8, 693804. [Google Scholar] [CrossRef] [Scilit]
- Liu, G.; Xu, Y.; Ge, W.; Yang, X.; Su, X.; Shen, B.; Ran, Q. How Can Marine Fishery Enable Low Carbon Development in China? Based on System Dynamics Simulation Analysis. Ocean Coast. Manag. 2023, 231, 106382. [Google Scholar] [CrossRef] [Scilit]
- Alsaleh, M.; Yang, Z. The Evolution of Information and Communications Technology in the Fishery Industry: The Pathway for Marine Sustainability. Mar. Pollut. Bull. 2023, 193, 115231. [Google Scholar] [CrossRef] [Scilit] [PubMed]
- González Laxe, F.; Bermúdez, F.M.; Palmero, F.M.; Novo-Corti, I. Governance of the Fishery Industry: A New Global Context. Ocean Coast. Manag. 2018, 153, 33–45. [Google Scholar] [CrossRef] [Scilit]
- Nguyen, T.V.; Hoang, N.K. How Economic Policies and Development Impact Marine Fisheries: Lessons Learned from a Transitional Economy. Ecol. Econ. 2024, 225, 108314. [Google Scholar] [CrossRef] [Scilit]
- Morosini, P. Industrial Clusters, Knowledge Integration and Performance. World Dev. 2004, 32, 305–326. [Google Scholar] [CrossRef] [Scilit]
- Yu, J.; Zhang, L. Evolution of Marine Ranching Policies in China: Review, Performance and Prospects. Sci. Total Environ. 2020, 737, 139782. [Google Scholar] [CrossRef] [Scilit] [PubMed]
- Yu, J.; Yan, T.; Ma, X. Evolution of Support Policies for the Development of Distant-Water Fishing Bases in China: Review, Performance, Challenges and Prospects. Mar. Policy 2025, 171, 106489. [Google Scholar] [CrossRef] [Scilit]
- Scroggins, R.E.; Fry, J.P.; Brown, M.T.; Neff, R.A.; Asche, F.; Anderson, J.L.; Love, D.C. Renewable Energy in Fisheries and Aquaculture: Case Studies from the United States. J. Clean. Prod. 2022, 376, 134153. [Google Scholar] [CrossRef] [Scilit]
- Feng, J.-C.; Sun, L.; Yan, J. Carbon Sequestration via Shellfish Farming: A Potential Negative Emissions Technology. Renew. Sustain. Energy Rev. 2023, 171, 113018. [Google Scholar] [CrossRef] [Scilit]
- Li, W.; Li, X.; Song, C.; Gao, G. Carbon Removal, Sequestration and Release by Mariculture in an Important Aquaculture Area, China. Sci. Total Environ. 2024, 927, 172272. [Google Scholar] [CrossRef] [Scilit] [PubMed]
- Zheng, S.; Yu, L. The Government’s Subsidy Strategy of Carbon-Sink Fishery Based on Evolutionary Game. Energy 2022, 254, 124282. [Google Scholar] [CrossRef] [Scilit]
- Wang, Y.; Yang, Y.; Hu, X. The Evolution and Effectiveness of China’s Marine Carbon Sink Fishery Policies. Ocean Coast. Manag. 2024, 259, 107470. [Google Scholar] [CrossRef] [Scilit]
- Thébaud, O.; Nielsen, J.R.; Motova, A.; Curtis, H.; Bastardie, F.; Blomqvist, G.E.; Daurès, F.; Goti, L.; Holzer, J.; Innes, J.; et al. Integrating Economics into Fisheries Science and Advice: Progress, Needs, and Future Opportunities. ICES J. Mar. Sci. 2023, 80, 647–663. [Google Scholar] [CrossRef] [Scilit]
- Wang, S.; Li, W.; Xing, L. A Review on Marine Economics and Management: How to Exploit the Ocean Well. Water 2022, 14, 2626. [Google Scholar] [CrossRef] [Scilit]
- Yan, A.; Guo, R.; Wang, Q.; Liu, Y.; Fu, X.; Yin, Q. The Development of Chinese Land-Raised Seafood Industry under Japanese Nuclear Wastewater Discharge into the Sea: Considering the Effects of Government Subsidy, Consumer Preference and Product Safety. Ocean Coast. Manag. 2025, 266, 107687. [Google Scholar] [CrossRef] [Scilit]
- Zhao, Y.; Li, Y. Blue Transition for Sustainable Marine Fisheries: Critical Drivers and Evidence from China. J. Clean. Prod. 2023, 421, 138535. [Google Scholar] [CrossRef] [Scilit]
- Hurd, C.L.; Law, C.S.; Bach, L.T.; Britton, D.; Hovenden, M.; Paine, E.R.; Raven, J.A.; Tamsitt, V.; Boyd, P.W. Forensic Carbon Accounting: Assessing the Role of Seaweeds for Carbon Sequestration. J. Phycol. 2022, 58, 347–363. [Google Scholar] [CrossRef] [Scilit] [PubMed]







| System | Dimension | Indicator | Unit | Attr. | References |
|---|---|---|---|---|---|
| Ecological System | Carbon Emission | X1: Carbon emissions from fishing vessels | 10 kt CO2 | − | [15,36] |
| X2: Carbon emissions from aquaculture electricity | 10 kt CO2 | − | |||
| X3: Carbon emissions from seafood processing | 10 kt CO2 | − | |||
| Carbon Sink | X4: Harvest-mediated biological-carbon removal indicator | 10 kt CO2 | + | ||
| X5: Carbon sink from shellfish & algae aquaculture | 10 kt CO2 | + | |||
| Social System | Marine Scientific Research | X6: Number of marine scientific research institutions | Number | + | [16,17,24,26] |
| X7: Number of marine scientific research personnel | People | + | |||
| X8: Number of R&D projects in marine institutions | Project | + | |||
| X9: Scientific and technological works by marine institutions | Article | + | |||
| X10: Number of technology patents from marine institutions | Item | + | |||
| Aquatic Technology Promotion | X11: Funding for aquatic technology promotion | 104 CNY | + | ||
| X12: Number of personnel in aquatic tech promotion | People | + | |||
| X13: Number of aquatic tech promotion institutions | Number | + | |||
| Environmental Governance | X14: Investment in environmental pollution control | 108 CNY | + | ||
| X15: Area of marine nature reserves | 104 km2 | + | |||
| Population Development | X16: Growth rate of marine fishery population | % | + | ||
| X17: Growth rate of marine fishery employees | % | + | |||
| Economic System | Resource Input | X18: Total investment in fixed assets | 108 CNY | + | [2,17,40] |
| X19: Mariculture area | kha | + | |||
| X20: Year-end number of motorized marine fishing vessels | 104 vessels | + | |||
| Economic Output | X21: Per capita output value of marine fisheries | 104 CNY/people | + | ||
| X22: Growth rate of marine fishery output value | % | + | |||
| X23: Proportion of marine to total aquatic products | Ratio | + | |||
| X24: Proportion of marine processed to total aquatic processed products | Ratio | + | |||
| X25: Output value of seafood processing | 108 CNY | + | |||
| X26: Output value of recreational fisheries | 108 CNY | + |
| Operation Type | Trawl | Seine | Gillnet | Stow Net | Angling | Other |
|---|---|---|---|---|---|---|
| Coefficient | 0.56 | 0.29 | 0.5 | 0.23 | 0.66 | 0.45 |
| Coordination Degree (D) | |||||
| Coordination Level | Moderate Dissonance | Slight Dissonance | Basic Coordination | Slight Coordination | Moderate Coordination |
| No. | Variable Name | Core Variable Equation | Unit |
|---|---|---|---|
| (1) | Regional GDP | INTEG (GDP Inflow–Marine Fishery Value Added, 230,872) | 108 CNY |
| (2) | Marine Capture Output | EXP(−0.025 + 0.228 × LN(IFA) + 1.574 × LN(MNV)) | 10 kt |
| (3) | Aquaculture Output | EXP(6.526 − 0.01 × LN(SPI) + 0.15 × LN(IFA) + 0.319 × LN(MA) − 0.404 × LN(MFE) + 0.152 × LN(ATF)) | 10 kt |
| (4) | Total Fishery Economic Output | (AQO_Value + MCO_Value + SPO_Value + RFO_Value)/Proportion | 108 CNY |
| (5) | Total Aquatic Product Output | 1.691 × MCO + 1.65 × AQO − 413.448 | 10 kt |
| (6) | Total Aquatic Processed Products | 0.422 × APO − 130.866 | 10 kt |
| (7) | Total Marine Processed Products | 0.401 × APO − 222 | 10 kt |
| (8) | Seafood Processing Output Value | (TMPP/TAPP) × APO_Value | 108 CNY |
| (9) | Marine Fishery Added Value | 1836.86 + 0.141 × TFE | 108 CNY |
| (10) | Recreational Fishery Output Value | 17 + 0.03 × IFA + 0.018 × AQO_Value + 0.05 × EPO + 0.2 × MNR | 108 CNY |
| (11) | Sci-tech & Promotion Index | 0.26 × ((RDP − 823)/4486) + 0.19 × ((SWI − 11562)/4980) + 0.16 × ((TPI − 4305)/1920) | Dmnl2 |
| (12) | Marine Scientific Research Personnel | 0.026 × Number of Marine Research Institutions − 1.535 | 104 people |
| (13) | R&D Projects in Research Institutions | 4345 × Marine Scientific Research Personnel − 2126 | Project |
| (14) | Scientific Works by Research Institutions | 5629 × Marine Scientific Research Personnel − 3275 | Article |
| (15) | Technology Patents of Research Institutions | 1922 × Marine Scientific Research Personnel − 2470 | Item |
| (16) | Harvest-Mediated Biological-Carbon Removal | (FCO × 2.18 + CCO × 1.25 + CeCO × 1.56) × (44/12) | 10 kt CO2 |
| (17) | Shellfish & Algae Aquaculture Biological-Carbon Component | (SAO × SAC) × (44/12) | 10 kt CO2 |
| (18) | Aquaculture Electricity Emissions | (SPO_pond × 370 + SPO_factory × 8660) × EEF / 1000 | 10 kt CO2 |
| (19) | Seafood Processing Emissions | SPO_Value × EEC × SCC × (44/12) | 10 kt CO2 |
| Variable Type | Historical Value Range | Status Quo | Restrictive Development | Economic Development | Environmental Protection | Integrated Coordination |
|---|---|---|---|---|---|---|
| Capture Policy Factor | 1 | 1 | 0.5000 | 2 | 0.5 | 1.5000 |
| Aquaculture Policy Factor | 1 | 1 | 0.5000 | 2 | 2 | 2 |
| Proportion of IFA | 0.0113–0.0212 | 0.0178 | 0.0113 | 0.0212 | 0.0191 | 0.0200 |
| Proportion of EPCI | 0.0053–0.0166 | 0.0105 | 0.0053 | 0.0053 | 0.0166 | 0.0157 |
| Area of Marine Nature Reserves | 2.9600–46.2600 | 11.3400 | 2.9600 | 2.9600 | 46.2600 | 43.9500 |
| Funding for Aquatic Tech Promotion | 4.9900–19.9900 | 13.2000 | 4.9900 | 19.9900 | 17.9900 | 18.9900 |
| Number of Marine Research Institutions | 137–216 | 161 | 137 | 216 | 194 | 205 |
| Personnel in Aquatic Tech Promotion | 11562–16542 | 14351 | 11562 | 16542 | 14888 | 15714 |
| Aquatic Tech Promotion Institutions | 4305–6225 | 5573 | 4305 | 6225 | 5602 | 5914 |
| Proportion of Marine Fishery Population | 0.0080–0.0100 | 0.0090 | 0.0080 | 0.0100 | 0.0080 | 0.0095 |
| Year-end No. of Motorized Vessels | 20.4000–29.7300 | 25.2500 | 20.4000 | 29.7300 | 20.400 | 28.2400 |
| Mariculture Area | 199.2200–231.7800 | 213 | 199.2200 | 231.7800 | 208 | 219.4500 |
| Year | MCO (Hist.) | MCO (Sim.) | MCO (Error %) | AQO (Hist.) | AQO (Sim.) | AQO (Error %) | SPOV (Hist.) | SPOV (Sim.) | SPOV (Error %) |
|---|---|---|---|---|---|---|---|---|---|
| 2010 | 1203.590 | 1222.170 | 1.540 | 1482.300 | 1401.110 | −5.480 | 1853.830 | 1847.020 | −0.370 |
| 2011 | 1241.940 | 1231.300 | −0.860 | 1551.330 | 1493.890 | −3.700 | 2101.800 | 2096.290 | −0.260 |
| 2012 | 1267.190 | 1225.780 | -3.270 | 1643.810 | 1642.100 | −0.100 | 2377.240 | 2398.300 | 0.890 |
| 2013 | 1264.380 | 1288.760 | 1.930 | 1739.250 | 1707.320 | −1.840 | 2581.800 | 2606.510 | 0.960 |
| 2014 | 1280.840 | 1292.530 | 0.910 | 1812.650 | 1772.220 | −2.230 | 2777.760 | 2799.530 | 0.780 |
| 2015 | 1314.780 | 1298.090 | −1.270 | 1875.630 | 1873.080 | −0.140 | 2873.550 | 2881.560 | 0.280 |
| 2016 | 1328.270 | 1261.420 | −5.030 | 1963.130 | 1898.900 | −3.270 | 3010.240 | 3015.420 | 0.170 |
| 2017 | 1112.420 | 1156.300 | 3.940 | 2000.700 | 2001.490 | 0.040 | 3184.950 | 3194.180 | 0.290 |
| 2018 | 1044.460 | 1082.170 | 3.610 | 2031.220 | 2014.780 | −0.810 | 3225.500 | 3197.800 | −0.860 |
| 2019 | 1000.150 | 978.677 | −2.150 | 2065.330 | 2067.050 | 0.080 | 3331.530 | 3285.850 | −1.370 |
| 2020 | 947.410 | 981.377 | 3.590 | 2135.280 | 2128.010 | −0.340 | 3210.080 | 3175.030 | −1.090 |
| 2021 | 951.460 | 941.742 | −1.020 | 2211.140 | 2148.180 | −2.850 | 3262.500 | 3242.220 | −0.620 |
| 2022 | 950.850 | 919.524 | −3.290 | 2275.700 | 2195.910 | −3.510 | 3353.790 | 3330.340 | −0.700 |
| Scenario | Integrated Coordination | Environmental Protection | Economic Development | Restrictive Development | Status Quo |
|---|---|---|---|---|---|
| Avg. Annual Increase (%) | 0.7610 | 0.3250 | 0.7490 | − 0.7600 | 0.1490 |
Disclaimer/Publisher’s Note: The statements, opinions and data contained in all publications are solely those of the individual author(s) and contributor(s) and not of MDPI and/or the editor(s). MDPI and/or the editor(s) disclaim responsibility for any injury to people or property resulting from any ideas, methods, instructions or products referred to in the content. |
© 2026 by the authors. Licensee MDPI, Basel, Switzerland. This article is an open access article distributed under the terms and conditions of the Creative Commons Attribution (CC BY) license.
Share and Cite
Du, J.; Liao, W.; Yan, B.; Dai, X. Navigating the Blue Transition: A Dynamic Simulation of the Coupling and Coordination in China’s Marine Fishery Ecological–Social–Economic System. Fishes 2026, 11, 501. https://doi.org/10.3390/fishes11090501
Du J, Liao W, Yan B, Dai X. Navigating the Blue Transition: A Dynamic Simulation of the Coupling and Coordination in China’s Marine Fishery Ecological–Social–Economic System. Fishes. 2026; 11(9):501. https://doi.org/10.3390/fishes11090501
Chicago/Turabian StyleDu, Jun, Wenhao Liao, Bo Yan, and Xinhui Dai. 2026. "Navigating the Blue Transition: A Dynamic Simulation of the Coupling and Coordination in China’s Marine Fishery Ecological–Social–Economic System" Fishes 11, no. 9: 501. https://doi.org/10.3390/fishes11090501
APA StyleDu, J., Liao, W., Yan, B., & Dai, X. (2026). Navigating the Blue Transition: A Dynamic Simulation of the Coupling and Coordination in China’s Marine Fishery Ecological–Social–Economic System. Fishes, 11(9), 501. https://doi.org/10.3390/fishes11090501
