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19 August 2022

29 Pages

New Records and Updated Checklist of the Pentatomoidea (Hemiptera: Heteroptera) of Greece

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1
Laboratory of Agricultural Zoology and Entomology, Agricultural University of Athens, Athens 118 55, Greece
2
Laboratory of Sericulture and Apiculture, Agricultural University of Athens, Athens 118 55, Greece
3
Laboratory of Agricultural Entomology, Institute for Olive Tree, Subtropical Plants and Viticulture, Hellenic Agricultural Organization ‘DEMETER’, Agrokipio, Chania 731 00, Greece
4
Laboratory of Genetics, Agricultural University of Athens, Athens 118 55, Greece
Insects2022, 13(8), 749;https://doi.org/10.3390/insects13080749
This article belongs to the Collection Hemiptera: Ecology, Physiology, and Economic Importance
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Simple Summary

An updated species checklist for all the Pentatomoidea species for Greece is provided. Eight species are recorded from Greece for the first time. The checklist is supported with distributional data notes for all the Pentatomoidea species of Greece.

Abstract

Eight species of the superfamily Pentatomoidea are recorded from Greece for the first time: Aelia germari Küster 1852, Eurygaster hottentotta (Fabricius 1775), Eysarcoris aeneus (Scopoli 1763), Neottiglossa lineolata (Herrich-Schaeffer 1830), Neottiglossa pusilla (Gallen 1789), Picromerus bidens (Linnaeus 1758), Podops (Podops) inunctus (Fabricius 1775) and Tarisa pallescens (Jakovlev 1871). A complete updated species checklist with distributional data notes for all the new species for Greece are provided.

1. Introduction

The superfamily Pentatomoidea constitutes one of the most important insect groups of the suborder Heteroptera. It includes 1080 genera and 5907 species belonging to 16 families of which the Cydnidae, Pentatomidae, Scutelleridae and Tessaratomidae are the most important; 94% of the species belong to these four families [1,2,3]. Pentatomidae is the fourth most numerous family within Heteroptera with more than 4700 species, commonly known as stink bugs [4]. It includes predominantly herbivorous species, some of them having high economic importance, such as the brown marmorated stink bug Halyomorpha halys (Stål) and the southern green stink bug Nezara viridula (L.).
Halyomorpha halys is a polyphagous stink bug native to China, Korea, Japan and Taiwan [5]. It was introduced into the United States in the mid-1990s, resulting in losses of apples, peaches, tomatoes and other important crops [6]. Recently, it has been detected in Europe for the first time in Switzerland in 2007, damaging stone fruits and legume pods [7]. Until recently, it has been recorded in many European countries including Greece [8], and it is considered a potential threat to agricultural productivity in all these regions [9,10] since it has a broad host range and is capable to disperse long-distance by flight [11].
There have been sporadic references on specific species of Pentatomoidea in Greece [8,12,13,14,15,16,17,18,19,20,21,22,23,24,25,26,27,28,29,30,31,32,33,34,35,36,37,38,39,40,41,42,43], and two overall studies, one in 1980 [44] and the other in 2019, which updated the previous one and included all new species [45]. Given the worldwide problem of the brown marmorated stink bug pest status [6] and its presence in Greece [8], understanding this family in Greece may assist in distinguishing this pest species from native ones, and provide more targeted response for future detections of this pest. The aim of the present study is to present new records and an updated checklist of the Pentatomoidea of Greece.

2. Materials and Methods

Insect material was collected using sweeping nets and glass containers, or by glass tube aspirators during 1974–2015. Insects were killed in glass tubes with ethyl-acetate or were directly stored in 70% ethyl-alcohol. Genitalia were prepared for observation under a Carl Zeiss Stemi 305 binocular stereoscope and Olympus CX23 binocular microscope by maceration in 10% potassium-hydroxide (KOH). All specimens were identified by the authors, according to keys and descriptions in references [46,47,48,49,50].
Material of the collected species was deposited in the collection established by the fourth author, which is kept in the Laboratory of Agricultural Zoology and Entomology of the Agricultural University of Athens.
Localities studied
Collection sites for each species are presented in the same fashion as Drosopoulos et al. [50]. The altitudes and distances reported for the collection sites may deviate ca. 50 m and ca. 500 m respectively.
The localities from which specimens were collected are the following:
  • NORTHWESTERN GREECE
    • Kerkyra: Tritsi; Liapades
    • Ioannina: Voutsaras (500 m); Pindos (Katara); Miliotades (600 m); Vrysochori (1000–1600 m); Aristi; Aoos-ΝΕ Konitsa; Vikos gorge (E Monodendri); Ioannina; Millies; Vouchorina; Korydallos; Votonossi
    • Florina: Ε Pisoderion (1650 m); Kalo-Nero (Vernon Mt; southern slopes; 1100 m); Florina; Megali Prespa (Psarades); Ladopotamos valley (Ν Kotas); Vigla; Pisoderion; Vevi; Varnous Mt. (Bela Voda)
    • Kastoria: Gavros (Ladopotamos valley); Aposkepos (800 m)
    • Kozani: 15 km SE Siatista; Vourinos Μt. (1000–1300 m)
    • Grevena: Anixis; Agii Theodori
    • Trikala: Orthovounion; Aghiofilo; Mourgani
    • Larisa: Olympus Μt. (southwestern slopes; Kryovryssi; 800 m)
    • Pieria: Olympus Μt. (eastern slopes; Stavros); Olympus Μt. (eastern slopes; Prionia; 1000–1250 m); Poroi (coastal swamp); Litochoro
    • Pella: Vryta; Ν Vegoritis Lake
  • WESTERN GREECE
    • Epirus: Arta (Louros Riv; near Agios Georgios); Vryssochori
    • Aetoloakarnania: Messolonghion; Messolonghion (Tourlida); Amphilochia (Anoixiatiko); Babini; Evinos Riv. (Evinochori); Astakos (Mytikas); Vonitsa (Loutrakion); Aetolikon; Trichonis (Kapsorachi)
    • Kephallinia: Aenos Mt. (600–1000 m); K. Katelios; Myrtos; Assos
  • NORTHEASTERN GREECE
    • Evros: Metaxades; Evros Delta
    • Samothraki: SW Kamariotissa (coastal swamp); Loutra
    • Xanthi: W Porto Lagos
    • Drama: Rodopi Μt. (fountain “Krya Vryssi” near Elatia); Rodopi Μt. (13 km SE Elatia); Rodopi Μt. (Vathyrhemma; 1450 m); Falakron Mt. (2000–2200 m); Falakron Μt. (1400–1600 m); Rodopi Μt. (ΝΕ Elatia in direction Vathyrhemma; 1300 m); Rodopi Μt. (Betula Forest; 25 km SE Elatia; 1100 m); Rodopi Μt. (ΝΕ “Virgin Wood”; supra Zagrantenia; 1800–1900 m); S Livaderon 600 m; Ν Drama 450 m; Potami; NW Silli (Prasinada; 800 m)
    • Thessaloniki: Cedron Hills; Plagiarion
  • CENTRAL GREECE
    • Attiki: Votanikos; Alimos; Kifissia; Amaroussion; Chalandri; Avlon; Mati; Marathon; Marathon (coastal swamp and salt marshes near Schinias); Parnis Μt. (1200 m); Parnis Mt. (Mola fountain); Ymitos Mt. (Kareas)
    • Voeotia: Arachova; E Agia Paraskevi; Aliartos; Tsoukalades; Avlis (Vathy)
    • Fokis: Monastirakion (Doris; coastal swamp); Skaloula (Doris; 600 m); Eratini (Doris); Kalion (Doris); Eleon; Galaxidi; Amfissa; Agios Nikolaos (Doris); NW Itea (Agia Euthymia); Ghiona Mt. (near refuge); Parnassos Mt. (National Park); Marathias
    • Fthiotis: Kalamakion; Oiti Mt. (1200–1600 m); Malessina; Kalamakion; Arkitsa
    • Karditsa: Artessiano
  • PELOPONNESOS
    • Korinthia: Killini Μt. (supra Trikala; 1550–1600 m); Akrokorinthos; Derveni
    • Achaia: Bouboukas; Kato Klitoria
    • Llia: Krestena; swamp near Loutra Kaiafa; Kyllini; SE Pyrgos (Paralia Zacharo)
    • Messinia: NE Kalamata (Artemissia); E Kyparissia (Dorion); Kazarma; Menina
    • Arkadia: Menalon Mt. (5 km W Vitina; 900–1000 m); SW Astros (Platanos); Tripolis; N Astros (Xeropigado); Parnon Μt. (Ν. Kosmas; 1100 m); Charadros; Parnon Μt. (supra Kastanitsa; 1100 m)
    • Lakonia: Mystras; Taygetos Mt. (600–1000m)
  • EAST AEGEAN ISLANDS
    • Ikaria: W Gyaliscari (coastal swamp); Ε Raches (600 m)
    • Samos: Pythagorion; Neochorion (5 km E Koumeika); S Psili Ammos (coastal swamp); Pyrgos (10 km E Koumeika); N Agios Konstantinos (coastal swamp); Chora
    • Chios: Chora; E Mastichochoria (Armolia); Central Mastichochoria (Agios Georgios)
    • Lesvos: Sykaminea (Skala); Andissa; Kalloni; Petra
  • CYCLADES ISLANDS
    • Naxos: SE Moutsouna, Apeiranthos
    • Paros: swamp near Paroikia; Monastirion
    • Santorini: Kamarion
    • Syros: Poseidonia
    • Tinos: Triantaros
  • DODECANESOS ISLANDS
    • Rhodos: Kremasti (coastal biotopes); Dimilia; Plymiri; Emponas; Nea Afantou; Apolakkia; Salakos; Petaloudes valley; Kalathos; Archipolis; Malonas
    • Kastellorizo: around village
  • CRETE
    • Heracleon: Agios Vassilios
    • Rethymnon: Myloi; Agioi Apostoloi; Oidi Mt. (600–1000 m)
    • Chania: Chryssopigi

3. Results

Results of this study showed that 92 species were found in 52 genera, belonging in 10 subfamilies, within 5 families of Pentatomoidea: Acanthosomatidae (Acanthosomatinae), Cydnidae (Cydninae, Sehirinae), Pentatomidae (Asopinae, Pentatominae, Podopinae), Plataspidae (Plataspinae) and Scutelleridae (Eurygastrinae, Odontoscelinae, Odontotarsinae). Eight of these species are new records for Greece: 6 species of the subfamily Pentatominae (Pentatomidae) and 2 species of Podopinae (Pentatomidae). The updated checklist with the new records is referred below in the text and in Table 1.
Checklist of species found in the present study (with bold and asterisk the new species record for Greece)
Family ACANTHOSOMATIDAE
Subfamily Acanthosomatinae
Genus Cyphostethus Fieber 1860 (Figure 1)
Figure 1. Map showing collecting sites of species of genera Cyphostethus and Elasmucha during the present study in Greece (AL = Albania, NM = North Macedonia, BG = Bulgaria, TR = Turkey).
  • Cyphostethus tristriatus (Fabricius 1860)
Doris [Agios Nikolaos (20.VII.1986), Skaloula (27.III.1979)], Olympos Mt. [Prionia (21.V.1981)]. Total: 3 specimens.
Genus Elasmucha Stål 1864 (Figure 1)
2.
Elasmucha grisea grisea (Linnaeus 1758)
Doris [Skaloula (2.V.1986)], Ioannina (24.VI.1984), Rodopi Mt. (10.VIII.1985). Total: 5 specimens.
Family CYDNIDAE
Subfamily Cydninae
Genus Cydnus Fabricius 1803 (Figure 2)
Figure 2. Map showing collecting sites of species of genera Cydnus, Macroscytus, Ochetostethus and Tritomegas during the present study in Greece.
3.
Cydnus aterrimus (Forster 1771)
Santorini Island [Kamarion (5.V.1982)]. Total: 1 specimen.
Genus Macroscytus Fieber 1860 (Figure 2)
4.
Macroscytus brunneus (Fabricius 1803)
Attiki [Kifissia (2.X.1981, 26.VIII.1985)], Doris [Agios Nikolaos (2.VII.1986)]. Total: 4 specimens.
Subfamily Sehirinae
Genus Ochetostethus Fieber 1860 (Figure 2)
5.
Ochetostethus balcanicus (Wagner 1940)
Doris [Agioi Pantes (26.V.1980), Skaloula (29.IV.1978, 1.VI.1985)], Olympos Mt. [Prionia (21.V.1981), Stavros (20.V.1981)]. Total: 5 specimens.
Genus Tritomegas Amyot et Serville 1843 (Figure 2)
6.
Tritomegas bicolor (Linnaeus 1758)
Epirus [Vryssochori (28.V.1981)], Voeotia [Arachova (17.III.1981)]. Total: 2 specimens.
7.
Tritomegas sexmaculatus (Rambur 1839)
Olympos Mt. [Stavros (20.V.1981)]. Total: 1 specimen.
Family PENTATOMIDAE
Subfamily Asopinae
Genus Arma Hahn 1832 (Figure 4)
8.
Arma insperata (Horváth 1899)
Florina [Vernon Mt-Kalo Nero (21.VIII.1986)]. Total: 1 specimen.
Genus Jalla Hahn 1832 (Figure 4)
9.
Jalla dumosa (Linnaeus 1758)
Doris [Skaloula (1.VI.1985)], Fokis [Elaion (22.VII.1983)], Kozani [Vourinos Mt. 18.VIII.1983)]. Total: 3 specimens.
Genus Picromerus Amyot et Serville 1843 (Figure 4)
10.
Picromerus bidens* (Linnaeus 1758) (Figure 3)
Florina [Vernon Mt-Kalo Nero (30.VII.1982)]. Total: 3 specimens.
Figure 3. Picromerus bidens (Collector S. Drosopoulos, Vernon Mt.-Kalo Nero 30.VII.1982, posited in S. Drosopoulos historical collection. Photo A. Tsagkarakis 12.VIII.2022).
11.
Picromerus conformis (Herrich-Schaeffer 1841)
Doris [Skaloula (27.IX.1983)], Florina [Psarades (18.VIII.1986)]. Total: 2 specimens.
Genus Troilus Stål 1867 (Figure 4)
12.
Troilus luridus (Fabricius 1775)
Olympos Mt. [Stavros (20.V.1981)]. Total: 1 specimen.
Genus Zicrona Amyot et Serville 1843 (Figure 4)
13.
Zicrona caerulea (Linnaeus 1758)
Attiki [Avlon (11.V.1980)], Doris [Monastirakion (18.VIII.1981)]. Total: 2 specimens.
Figure 4. Map showing collecting sites of species of genera Arma, Jalla, Picromerus, Troilus and Zicrona during the present study in Greece.
Subfamily Pentatominae
Genus Acrosternum Fieber 1860 (Figure 6)
14.
Acrosternum heegeri (Fieber 1861)
Doris [Eratini (14.VII.1978)], Zakynthos [Volimes (28.VI.1988)], Athos [Karyes (5.VIII.1986)]. Total: 7 specimens.
15.
Acrosternum millierei (Mulsant et Rey 1866)
Attiki [Marathon (11.VII.1984)], Chios [Chora (5.IV.1979)], Doris [Agios Nikolaos (20.VII.1986)], Kynouria [Astros (31.VIII.1984)]. Total: 9 specimens.
Genus Aelia Fabricius 1803 (Figure 6)
16.
Aelia acuminata (Linnaeus 1758)
Aetoloakarnania [Messolonghion (17.V.1990)], Amphilochia [Anixiatiko (20.VIII.1985)], Arkadia [Menalon Mt. (20.VI.1986)], Attiki [Amaroussion (24.V.1978, 27.IX.1978), Avlon (4.VII.1978, 16.VII.1979), Kifissia (10.I.1978, 8.II.1978), Parnis Mt(1.V.1983)], Chalkidiki [Valti (8.VIII.1985)], Chios [Chora (5.IV.1979, 20.IX.1984)], Doris [Kallion (12.V.1980), Skaloula (24.VII.1977, 28.IV.1978, 20.VI.1978, 12.VII.1978, 17.IX.1978, 25.X.1978, 27.III.1979, 29.IV.1979, 5.X.1979, 2.XII.1979, 16.VII.1987)], Epirus [Vryssochori (26.V.1981, 27.V.1981)], Evia [Oreoi (20.IV.1980)], Evrytania [Megalo Chorio (10.VIII.1986)], Drama [Falakron Mt. (13.VI.1982)], Florina [Kotas (15.VIII.1979, Vernon Mt. (30.VII.1982), Vigla (30.VIII.1983), Florina (15.VIII.1979)], Fokis [Galaxidi (15.IV.1980)], Fthiotis [Anthili (12.VIII.1979), Malessina (3.IV.1979), Oiti Mt. (9.VI.1984, 30.XI.1984, 15.VI.1985)], Grevena [Anoixis (17.VIII.1983, 30.VI.1984)], Ikaria [Gyaliskari (15.VII.1981), Raches (14.VII.1981)], Ioannina [Aristi (13.VIII.1986, 23.VIII.1986), Ioannina (29.V.1981)], Karditsa [Artessiano (1.X.1981)], Konitsa [Aoos Riv. (29.V.1981)], Lakonia [Mystras (30.IV.1985)], Lesvos [Sykamnia (16.VI.1987)], Messinia [Artemissia (29.IV.1985), Dorion (1.V.1985)], Naxos [Apeiranthos (16.VI.1981)], Olympos Mt. (13.VIII.1979), Parnassos Mt. [National Park (13.VII.1985)], Paros [Paroikia (18.VI.1981)], Pella [Vryta (14.VIII.1979)], Pieria [Poroi (12.VIII.1979, 14.VIII.1980)], Pindos Mt. [Miliotades (5.IX.1980, 29.IX.1980, 25.V.1981), Katara (21.VIII.1983)], Rodopi Mt. [Elatia (11.VI.1982, 26.VII.1982, 13.VIII.1985)], Rhodos [Dimilia (31.V.1990)], Strymonas Riv. (1.VI.1982), Thessaloniki [Cedron Hills (26.XII.1979, 31.V.1982, 14.X.1982), Plagiarion (23.V.1981)], Trikala [Orthovounion (30.IX.1981)], Vegoritis Lake (14.VIII.1979), Xanthi [Porto Lagos (9.VI.1982, 23.VIII.1983)], Zakynthos [Keri (29.VI.1988)]. Total: 119 specimens.
17.
Aelia albovittata (Fieber 1868)
Chios [Armolia (20.VI.1987)], Samos [Pythagorion (23.VI.1987)]. Total: 3 specimens.
18.
Aelia germari* (Küster 1852) (Figure 5)
Evros [Metaxades (2.VI.1982)]. Total: 1 specimen.
Figure 5. Aelia germari (Collector S. Drosopoulos, Metaxades, 2.VI.1982, posited in S. Drosopoulos historical collection. Photo A. Tsagkarakis, 12.III.2021).
19.
Aelia klugii (Hahn 1831)
Florina [Vernon Mt. Kalo Nero (30.VII.1982, 12.VI.1982)], Rodopi [Vathyrhemma (12.VI.1982)]. Total: 5 specimens.
20.
Aelia rostrata (Boheman 1852)
Attiki [Avlon (26.V.1978, 11.V.1979), Parnis Mt. (29.V.1985)], Doris [Ghiona (23.VII.1977, 16.VII.1878)], Drama [Potami (12.VI.1982)], Epirus [Vryssochori (28.V.1981)], Evros [Metaxades (2.VI.1982), Evros Riv. (Delta 6.VI.1982)], Florina (15.VIII.1979, Vernon Mt.-Kalo Nero (21.VII.1983)), Fthiotis [Anthili (12.VIII.1980)], Konitsa [Aoos Riv. (30.V.1981)], Kozani [Vourinos Mt. (18.VIII.1985)], Messinia [Artemissia (29.IV.1985), Dorion (1.V.1985)], Oiti Mt. (1.VII.1984, 15.VI.1985)], Olympos Mt. [Stavros (20.V.1981)], Parnon Mt. (18.VI.1986), Prionia (30.V.1982), Rhodos [Plymiri (29.VI.1987)], Rodopi Mt. [Elatia (19.VII.1983)]. Total: 38 specimens.
21.
Aelia virgata (Klug 1841)
Evros [Metaxades (2.VI.1982)], Trikala [Aghiofilo (22.VII.1989)]. Total: 2 specimens.
Genus Antheminia Mulsant et Rey 1866 (Figure 6)
Figure 6. Map showing collecting sites of species of genera Acrosternum and Aelia during the present study in Greece.
22.
Antheminia lunulata (Goeze 1778)
Fthiotis [Kalamakion (12.VII.1979)], Pella [Vryta (14.VIII.1979)], Vegoritis Lake (14.VIII.1979). Total: 3 specimens.
Genus Apodiphus Saunders 1877 (Figure 7)
Figure 7. Map showing collecting sites of species of genera Apines, Apodiphus, Brachynema, Carpocoris and Codophila during the present study in Greece.
23.
Apodiphus amygdali (Germar 1817)
Attiki [Mati (20.VI.1977), Amaroussion (30.VIII.1985)], Doris [Skaloula (5.X.1980)]. Total: 7 specimens.
Genus Brachynema Mulsant et Rey 1852 (Figure 7)
24.
Brachynema cinctum (Fabricius 1775)
Attiki [Schinias (1.VII.1982)]. Total: 1 specimen.
Genus Carpocoris Kolenati 1846 (Figure 7)
25.
Carpocoris (Carpocoris) fuscispinus (Boheman 1851)
Attiki [Kifissia (30.III.1978), Marathon (14.V.1985), Parnis Mt. (13.VII.1984, 12.VII.1985)], Doris [Skaloula (24.VII.1977, 15.X.1979)], Epirus [Vryssochori (27.V.1981)], Evia [Agios Georgios (21.VI.1980), Aedipsos (23.VI.1980)], Kozani [Vourinos Mt. (17.VI.1985, 22.VII.1989)], Parnon Mt. [Kastanitsa (18.VI.1986), Kosmas (14.VII.1982, 11.VI.1985)], Messinia [Kazarma (4.VII.1984)], Olympos Mt. [Prionia (13.VIII.1979)], Paros [Paroikia (18.VI.1981)], Rhodos [Emponas (30.V.1990), Petaloudes (29.V.1990)], Samos [Neochorion (24.VI.1987), Pyrgos (24.VI.1987)], Oiti Mt. (9.VIII.1986), Thessaloniki [Cedron Hills (14.XI.1982)], Vegoritis Lake (14.VIII.1979). Total: 38 specimens.
26.
Carpocoris (Carpocoris) mediterraneus mediterraneus (Tamanini 1958)
Evros Riv. [Delta (7.VI.1982)], Naxos [Moutsouna (15.III.1982)]. Total: 4 specimens.
27.
Carpocoris (Carpocoris) pudicus (Poda 1761)
Attiki [Kifissia (30.III.1978), Parnis Mt. (29.V.1985)], Doris [Skaloula (20.VII.1978, 5.X.1980)], Florina [Vevi (24.VIII.1983)], Rodopi Mt. [Betoula Forest (22.V.1983)], Konitsa [Aoos Riv. (30.V.1981)], Kozani [Vourinos Mt. (17.VI.1985)], Vegoritis Lake (14.VIII.1979). Total: 10 specimens.
Genus Chlorochroa Stål 1872 (Figure 7)
28.
Chlorochroa sp. Stål 1872
Olympos Mt. [Stavros (20.V.1981)]. Total: 1 specimen.
Genus Codophila Mulsant et Rey 1866 (Figure 8)
Figure 8. Map showing collecting sites of species of genera Dolycoris, Dyroderes and Eurydema during the present study in Greece.
29.
Codophila varia (Fabricius 1787)
Aetoloakarnania [Babini (23.VI.1986), Evinos Riv. (22.VI.1986)], Arkadia [Platanos (18.VI.1985), Xeropigado (9.V.1985)], Attiki [Parnis Mt. (13.VII.1984), Schinias (1.VII.1982)], Avlis [Vahty (16.VI.1979)], Doris [Skaloula (18.VII.1978, 20.VII.1978, 17.IX.1978, 15.VIII.1983, 1.VI.1985)], Evia [Agios Georgios (21.VI.1980)], Fokis [Eleon (17.VIII.1979)], Ikaria [Gyaliskari (15.VII.1981)], Ilia [Krestena (16.V.1985)], Kephallinia [Aenos Mt. (23.VI.1988)], Kynouria [Astros (5.VII.1984)], Messinia [Kazarma (4.VII.1984)], Parnassos Mt. (13.VII.1985), Rhodos [Emponas (30.V.1990), Malonas (30.V.1990)], Samos [Psili Ammos (22.VI.1987), Pythagorion (23.VI.1987)], Aoos Riv. (30.V.1981), Giona (23.VII.1982), Oiti Mt. (30.IX.1984), Parnon Mt. (18.VI.1986), Vegoritis Lake (14.VII.1979). Total: 43 specimens.
Genus Dolycoris Mulsant et Rey 1866 (Figure 8)
30.
Dolycoris baccarum (Linnaeus 1758)
Achaia [Rion (9.VI.1980)], Aetoloakarnania [Babini 23.VI.1986)], Attiki [Amaroussion (2.III.1978), Kifissia (30.III.1978), Parnis Mt. (13.VII.1984, 12.VII.1985, 7.VII.1987)], Doris [Skaloula (15.VII.1978, 9.VI.1979, 29.VI.1979, 5.X.1980)], Epirus [Vryssochori (26.V.1981)], Evia [Edipsos (23.VI.1980)], Evros [Metaxades (2.VI.1982)], Florina (15.VIII.1979)], Fokis [Amfissa (19.IX.1978), Galaxidi (15.IV.1980)], Grevena [Anoixis (31.VII.1984)], Ilia [Kyllini (21.VII.1982)], Kephallinia [Aenos Mt. (23.VI.1988)], Konitsa [Aoos Riv. (29.V.1981)], Lakonia [Taygetos Mt. (30.IV.1985)], Messinia [Kazarma (4.VII.1984)], Kozani [Vourinos Mt. (28.V.1982)], Naxos [Moutsouna (16.VI.1981)], Oiti Mt. (30.IX.1984), Olympos Mt. [Stavros (13.VIII.1989, 20.V.1981), Sykaminos (30.V.1982)], Paros [Monastirion (17.VI.1981)], Pindos [Katara (4.IX.1980), Milies 20.VIII.1985)], Rhodos [Nea Afantou (28.VI.1987), Salakos (30.V.1990)], Rodopi Mt. [Elatia (25.VII.1982), Rodopi Mt. (27.VII.1982, 10.VIII.1985)], Samos [Pyrgos (14.VI.1987)], Parnon Mt. (18.VI.1986), Vourinos (28.V.1982)]. Total: 57 specimens.
Genus Dyroderes Spinola 1837 (Figure 8)
31.
Dyroderes umbraculatus (Fabricius 1775)
Doris [Skaloula (28.IV.1981)], Epirus [Vryssochori (26.V.1981)], Messinia [Dorion (1.V.1985)], Pindos [Greveniti (6.IX.1980)]. Total: 9 specimens.
Genus Eurydema Laporte 1833 (Figure 8)
32.
Eurydema (Eurydema) eckerleini (Josifov, 1961)
Syros [near Poseidonia (10.VI.2005)], Attiki [Ymitos Mt. (2.VIII.2005), Alimos (15.VII.2012), Votanikos (20.VII.2012)], Tinos [near Triantaros (29.VII.2016)]. Total: 100 specimens.
33.
Eurydema (Horvatheurydema) fieberi, (Schummel 1837)
Attiki [Kifissia (15.IV.1974)], Doris [Giona (16.VII.1978), Skaloula (3.V.1989)], Parnon Mt. (30.VI.1989), Kozani [Vourinos Mt. (22.VII.1989)]. Total: 5 specimens.
34.
Eurydema (Eurydema) oleracea (Linnaeus 1758)
Creta [Rethymnon-Myloi (8.VIII.1985)], Epirus [Vryssohori (26.V.1981)], Florina [Vernon Mt. (30.VII.1982), Vevi (30.VIII.1983)], Kozani [Vourinos Mt. (27.V.1982, 18.VIII.1983, 28.VI.1984, 18.VIII.1985)], Messinia [Artemissia (29.IV.1985)], Olympos Mt. [Prionia (13.VIII.1979, 20.V.1981, 21.V.1981), Stavros (12.V.1990)], Pindos [Miliotades (29.IX.1980), Panagia-Katara (25.V.1981)], Rodopi Mt. [Krya Vryssi (25.VII.1982), Rodopi Mt. (23.V.1983, 12.VIII.1985)], Oiti Mt. (30.IX.1984). Total: 40 specimens.
35.
Eurydema (Eurydema) ornata (Linnaeus 1758)
Achaia [Bouboukas (21.VI.1986)], Aetoloakarnania [Mytikas-Astakos (23.VI.1986)], Arkadia [Tripolis (19.VI.1986)], Attiki [Amaroussion (27.IX.1978), Avlon (16.X.1979, 7.VIII.1987), Kifissia (9.VIII.1977, 30.III.1978, 11.VII.1980, 25.VIII.1980), Parnis Mt. (13.VII.1984)], Avlis [Vathy (3.VII.1978)], Doris [Agioi Pantes (26.V.1980), Skaloula (13.VII.1977, 13.VIII.1977, 12.VI.1978, 17.IX.1978, 19.VIII.1979, 30.V.1990)], Epirus [Vryssochori (27.V.1981)], Evia [Agios Georgios (21.VI.1980)], Florina [Kotas (15.VIII.1979)], Fthiotis [Malessina (21.III.1979)], Grevena [Anoixis (18.VIII.1983)], Kozani [Vourinos Mt. (22.VII.1979, 18.VIII.1983, 22.VII.1989)], Lesvos [Sykaminea (16.VI.1987)], Parnassos Mt. [National Park (13.VII.1985)], Pieria [Poroi (12.VIII.1979), Varikon (23.VII.1982)], Rhodos [Apolakkia (30.VI.1987), Dimilia (31.V.1990), Petaloudes (29.V.1990)], Samos [Agios Konstantinos (30.VI.1987)], Trikala [Mourgani (22.VII.1989)], Voeotia [Agia Paraskevi (8.VI.1980)], Ioannina (29.V.1981), Oiti Mt. (1.VII.1984, 15.VI.1985), Parnon Mt. (18.VI.1986, 30.VI.1989). Total: 75 specimens.
36.
Eurydema (Horvatheurydema) rugulosa (Dohrn 1861)
Lesvos [Sykaminea (16.VI.1987)], Samos [Chora (25.VI.1987), Pyrgos (24.VI.1987)]. Total: 6 specimens.
37.
Eurydema (Rubrodorsalium) spectabilis (Stichel 1960)
Crete [Agios Vassilios-Heracleon (10.VII.1985)]. Total: 12 specimens.
38.
Eurydema (Rubrodorsalium) ventralis (Kolenati 1846)
Doris [Marathias 1.V.1979)]. Total: 24 specimens.
Genus Eysarcoris Hahn 1834 (Figure 10)
39.
Eysarcoris aeneus* (Scopoli 1763) (Figure 9)
Rodopi Mt. [Sidironero (23.V.1983)]. Total: 1 specimen.
Figure 9. Eysarcoris aeneus (Collector S. Drosopoulos, Sidironero, 23.V.1983, posited in S. Drosopoulos historical collection. Photo A. Tsagkarakis, 12.III.2021).
40.
Eysarcoris ventralis (Westwood, 1837)
Aetoloakarnania [Loutrakion-Vonitsa (22.VIII.1985)], Attiki [Amarousion (12.VII.1977), Avlon 4.VIII.1978), Kifissia (22.VI.1977), Schinias (1.VII.1982)], Crete [Rethymnon-Myloi (8.VII.1985)], Delta Acheloou (21.VII.1980), Delta Aliakmonos (19.VIII.1983), Doris [Kallion (18.VII.1978, 7.X.1979), Monastirakion (29.VII.1982), Skaloula (12.VII.1978)], Fokis [Amfissa (19.IX.1978)], Fthiotis [Anthili (26.VII.1977, 12.VIII.1980), Arkitsa (11.VIII.1980)], Ikaria [Gyaliskari (15.VII.1981)], Ioannina [Aristi (13.VIII.1985), Ioannina (20.VIII.1985)], Karpathos [Arkassa (3.VI.1990)], Korinthia [Kokoni (3.VIII.1977)], Messinia [Artemissia (11.VI.1985)], Olympos Mt. [Stavros (20.V.1981)], Paros [Paroikia (18.VI.1981)], Pindos [Katara (4.IX.1980), Miliotades (29.IX.1980)], Rhodos [Salakos (30.V.1990)], Samos [Agios Konstantinos (27.VI.1987), Psili Ammos (22.VI.1987)], Xanthi [Porto Lagos (9.VI.1982)], Loutra Kaiafa (3.VII.1984). Total: 49 specimens.
Genus Halyomorpha Mayr 1864 (Figure 10)
41.
Halyomorpha halys (Stål 1855)
Attiki [Votanikos (2.II.2015, 10.X.2016, 6.IX.2017), Alimos (20.V.2015), Kifissia (25.V.2015)], Crete [Chania (18.X.2015)], Kastellorizo [around village (15.VIII.2015)]. Total: 25 specimens.
Genus Holcogaster Fieber 1860 (Figure 10)
42.
Holcogaster fibulata (Germar 1831)
Attiki [Marathon (11.VII.1984)], Doris [Agios Nikolaos (20.VII.1986)]. Total: 2 specimens.
Genus Holcostethus Fieber 1860 (Figure 10)
43.
Holcostethus albipes (Fabricius 1781)
Kynouria [Astros (31.VII.1984)]. Total: 1 specimen.
44.
Holcostethus sphacelatus (Fabricius 1794)
Epirus [Vryssochori (26.V.1981)], Konitsa [Aoos Riv. 29.V.1981)], Olympos Mt. [Prionia (21.V.1981)], Rodopi Mt. [Virgin Wood 23.V.1983)]. Total: 7 specimens.
Genus Mustha Amyot et Serville 1843 (Figure 10)
45.
Mustha spinulosa (Lefebvre 1831)
Attiki [Avlon (26.VII.1979), Kifissia (16.VI.1978, 25.VII.1980)], Doris [Skaloula (18.V.1989)], Trikala [Mourgani (19.VI.1985)]. Total: 6 specimens.
Figure 10. Map showing collecting sites of species of genera Eysarcoris, Halyomorpha, Holcogaster, Holcostethus and Mustha during the present study in Greece.
Genus Neostrachia Dallas 1851 (Figure 13)
46.
Neostrachia bisignata (Walker 1867)
Doris [Monastirakion (29.VII.1982)], Kynouria [Astros (24.VIII.1988)], Volos [Amaliapolis (1.IX.1990)]. Total: 3 specimens.
Genus Neottiglossa Kirby 1837 (Figure 13)
47.
Neottiglossa bifida (Costa 1847)
Achaia [Bouboukas (21.VI.1986)], Aetoloakarnania [Messolonghion-Tourlida (20.VII.1990), Mytikas-Astakos (23.VI.1986)], Doris [Kallion (14.VIII.1977), Monastirakion (18.VIII.1981), Skaloula (24.VII.1977, 13.VII.1978, 9.VI.1979, 1.VI.1985)], Messinia [Menina (30.IV.1985), Artemissia 29.VI.1985), Kazarma (4.VII.1984)], Konitsa [Aoos-Monastiri (30.V.1981)], Lakonia [Mystras (30.IV.1985)], Naxos [Apeiranthos (16.VI.1981)], Pieria [Poroi (14.VIII.1980)], Thessaloniki [Plagiarion (23.V.1981)]. Total: 22 specimens.
48.
Neottiglossa flavomarginata (Lucas 1849)
Kozani [Vourinos Mt. (29.VI.1984)]. Total: 6 specimens.
49.
Neottiglossa leporina (Herrich-Schaeffer 1830)
Doris [Krokylion (19.VII.1978), Mavrolithari (8.VIII.1986)], Drama [Potami (12.VI.1982)], Epirus [Vryssochori (26.V.1981)], Florina [Vernon Mt. Kalo Nero (30.VII.1982), Florina (1.X.1983)], Konitsa [Aoos-Monastiri (10.V.1981)], Kozani [Vourinos Mt. (8.VI.1984)], Pieria [Poroi (14.VIII.1980)], Rodopi Mt. [Sidironero (23.V.1983), Vathyrhemma (26.VII.1982)], Oiti Mt. (30.IX.1984, 15.VI.1985)]. Total: 22 specimens.
50.
Neottiglossa lineolata* (Mulsant et Rey 1852) (Figure 11)
Doris [Skaloula (18.VII.1978)], Evia [Agios Georgios (21.VI.1980), Gialtra (23.VI.1980)], Konitsa [Aoos-Monastiri (30.V.1981)], Lesvos [Antissa (17.VI.1987)], Xanthi [Porto Lagos (9.VI.1982)]. Total: 8 specimens.
Figure 11. Neottiglossa lineolata (Collector S. Drosopoulos, Porto Lagos, 9.VI.1982, posited in S. Drosopoulos historical collection. Photo A. Tsagkarakis, 12.III.2021).
51.
Neottiglossa pusilla* (Gallen 1789) (Figure 12)
Rodopi Mt. [Vathyrhemma (26.VII.1982)]. Total: 2 specimens.
Figure 12. Neottiglossa pusilla (Collector S. Drosopoulos, Vathyrhemma, 26.VII.1982, posited in S. Drosopoulos historical collection. Photo A. Tsagkarakis, 12.III.2021).
Genus Nezara Amyot et Serville 1843 (Figure 13)
52.
Nezara viridula (Linnaeus 1758)
Attiki [Chalandri (3.VII.1987), Kifissia (9.VIII.1977, 18.VIII.1978, 2.VII.1979, 29.X.1984, 6.II.1986), Mati (20.VI.1977)]. Crete [Heracleon-Agios Vassilios (10.VII.1985)], Paros [Paroikia (18.IX.1981)]. Total: 13 specimens.
Genus Palomena Mulsant et Rey 1866 (Figure 13)
Figure 13. Map showing collecting sites of species of genera Neostrachia, Neottiglossa, Nezara and Palomena during the present study in Greece.
53.
Palomena prasina (Linnaeus 1761)
Kozani [Vourinos Mt. (28.V.1982)], Oiti Mt. (30.IX.1984), Parnon Mt. [Kastanitsa (19.VII.1982, 15.IX.1983)], Pindos [Miliotades (29.IX.1980)], Rodopi Mt. [Betula Forest (22.V.1983)], Tripolis [Tripolis (19.VI.1986)]. Total: 10 specimens.
Genus Pentatoma Olivier 1789 (Figure 14)
Figure 14. Map showing collecting sites of species of genera Peribalus, Pentatoma, Piezodorus, Pitedia and Rhaphigaster during the present study in Greece.
54.
Pentatoma (Pentatoma) rufipes (Linnaeus 1758)
Florina [Vernon Mt.-Kalo Nero (21.VII.1983, 21.VIII.1986), Vigla (30.VIII.1983)], Rodopi Mt. [Betula Forest (25.VII.1992), Sidironero (27.VII.1992)]. Total: 10 specimens.
Genus Peribalus Mulsant et Rey, 1866 (Figure 14)
55.
Peribalus strictus strictus (Fabricius 1803)
Attiki [Avlon (4.VIII.1978), Kifissia (30.VII.1978, 8.X.1987)], Avlis [Vathy (3.VII.1978)], Crete [Agios Vassilios-Heracleon (25.IV.1975)], Doris [Skaloula (18.VII.1978, 17.IX.1978, 7.VIII.1986)], Florina [Kotas (15.VIII.1979)], Fthiotis [Malessina (21.III.1979)], Parnassos Mt. [National Park (13.VII.1985)], Rhodos [Salakos (30.V.1990)], Samos [Agios Konstantinos (27.VI.1987)]. Total: 16 specimens.
Genus Piezodorus Fieber 1861 (Figure 14)
56.
Piezodorus lituratus (Fabricius 1794)
Attiki [Parnis-Mola (12.VII.1985)], Schinias [(16.V.1984)], Avlis [Vathy 13.IV.1978)], Doris [Eratini (4.III.1979, 8.III.1988), Skaloula (28.IV.1978, 12.VII.1978, 14.VII.1978, 30.X.1982)], Epirus [Vryssochori (26.V.1981)], Evia [Agios Georgios (21.VI.1980)], Evrytania [Megalo Chorio (10.VIII.1986)], Florina [Vernon Mt.-Kalo Nero (30.VII.1982)], Messinia [Menina (30.IV.1985)], Ilia [Killini Mt. (21.VII.1982)], Kerkyra [Tritsi (3.IV.1985)], Oaks [Rodopi (12.VIII.1985)], Parnon Mt., [Kastanitsa (14.VIII.1982, 15.IX.1983)], Ikaria [Raches (14.VII.1981)], Rhodos [Apolakkia (30.VI.1987), Emponas (30.VI.1990), Messanagros (29.VI.1987), Petaloudes (29.V.1990)]. Total: 31 specimens.
Genus Rhaphigaster Laporte 1833 (Figure 14)
57.
Rhaphigaster nebulosa (Poda 1761)
Acheloos Riv. [Delta Acheloou (21.VII.1990)], Attiki [Kifissia (16.II.1978, 18.VIII.1978, 5.VI.1980, 12.IX.1983), Mati (28.VIII.1977)], Avlis [Vathy (5.V.1981)], Crete [Heracleon-Agios Vassilios (10.VII.1985)], Doris [Mavrolithari (8.VIII.1986), Skaloula (2.XI.1979, 27.IX.1983, 26.III.1984, 21.XII.1987)], Evros Riv. [Delta Evrou (7.VI.1982)], Aetoloakarnania [Nafpaktos (4.IX.1979)], Rhodos [Petaloudes (29.V.1990)]. Total: 17 specimens.
Genus Sciocoris Fallen 1829 (Figure 15)
Figure 15. Map showing collecting sites of species of genera Sciocoris, Stagonomus and Staria during the present study in Greece.
58.
Sciocoris (Sciocoris) cursitans cursitans (Fabricius 1794)
Doris [Skaloula (12.IV.1985)], Lesvos [Andissa (17.VI.1987)]. Total: 2 specimens.
59.
Sciocoris (Sciocoris) deltocephalus (Fieber 1861)
Kephallinia [Aenos Mt. (23.VI.1988)], Paros [Paroikia (18.VI.1981)]. Total: 2 specimens.
60.
Sciocoris (Sciocoris) helferii (Fieber 1851)
Aetoloakarnania [Mytikas-Astakos (23.VI.1986)], Arkadia [Xeropigado (9.V.1985)], Attiki [Parnis Mt.-Mola (12.VII.1985)], Ikaria [Gialiskari (15.VII.1981)], Kephallinia [Aenos Mt. (23.VI.1988)], Konitsa [Aoos-Monastiri (30.V.1981)], Kozani [Vourinos Mt. (29.VI.1984)], Aoos Riv. (14.VIII.1986)], Lesvos [Kalloni (18.VI.1987)], Pieria [Varikon (23.VII.1982)]. Total: 11 specimens.
61.
Sciocoris (Aposciocoris) macrocephalus (Fieber 1851)
Arkadia [Xeropigado (9.V.1985)], Attiki [Avlon (26.V.1978, 4.VIII.1978, 4.VII.1979) Marathon (11.VII.1984, 14.V.1985)], Doris [Skaloula (1.VI.1985)], Fokis [Amfissa (2.V.1979)], Kephallinia [K. Katelios (22.VI.1988)], Kerkyra [Liapades (2.IV.1985)]. Total: 16 specimens.
62.
Sciocoris (Neosciocoris) maculatus (Fieber 1851)
Kephallinia [Aenos Mt. (23.VI.1988)], Konitsa [Aoos-Monastiri (29.V.1981, 30.V.1981), Aoos Riv. (14.VIII.1986)], Olympos Mt. [Kryovryssi (30.V.1982)], Samothraki [Loutra (4.VI.1982)], Oiti Mt. (9.VIII.1986). Total: 8 specimens.
63.
Sciocoris (Sciocoris) sulcatus (Fieber 1851)
Aetoloakarnania [Messolongion-Tourlida (20.VII.1990)], Attiki [Avlon (26.V.1978, 11.V.1979), Kifissia (25.VIII.1980), Parnis Mt. (29.V.1985)], Kephallinia [Aenos Mt. (23.VI.1986)], Korinthia [Derveni (15.V.1985)], Kozani [Vourinos Mt. (29.VI.1984)], Kynouria [Kastanitsa (20.V.1982)], Messinia [Dorion (1.V.1985)], Olympos Mt. [Prionia (21.V.1981)], Pindos [Milies (20.VIII.1985)], Rhodos [Kalathos (28.VI.1987), Nea Afantou (28.VI.1987), Petaloudes (29.V.1990)], Rodopi Mt. [Elatia (25.VII.1982)], Voeotia [Tsoukalades (12.V.1988)]. Total: 19 specimens.
Genus Stagonomus Gorski 1852 (Figure 15)
64.
Stagonomus (Stagonomus) amoenus (Brulle 1832)
Crete [Heracleon-Agios Vassilios (25.IV.1985)], Konitsa [Aoos Riv. (29.V.1981), Aoos-Monastirion (30.V.1981)], Paros [Monastirion (17.VI.1981)], Rhodos [Petaloudes (29.V.1990)]. Total: 15 specimens.
65.
Stagonomus (Dalleria) bipunctatus (Linnaeus 1758)
Evia [Agios Georgios (21.VI.1980)], Florina [Vermon Mt.-Kalo Nero (30.VII.1982)], Kozani [Vourinos Mt. (18.VIII.1985)]. Total: 7 specimens.
Genus Staria Dohrn 1860 (Figure 15)
66.
Staria lunata (Hahn 1835)
Arkadia [Xeropigado (9.V.1985)], Attiki [Marathon (11.VII.1984)], Doris [Skaloula (15.VII.1978, 20.VII.1978, 17.IX.1978, 29.IV.1979)], Evros [Metaxades (2.VI.1982)], Fokida [Agia Euthymia (12.V.1988)], Konitsa [Aoos (27.V.1981)], Korinthia [Akrokorinthos (31.VII.1979)], Kozani [Vourinos Mt. (28.VI.1984)] Menalon Mt. (20.VI.1986), Parnassos Mt. [National Park (13.VII.1985), Parnassos Mt. (23.V.1980)], Rodopi Mt. [Vathyrhemma (26.VII.1982)]. Total: 36 specimens.
Subfamily Podopinae
Genus Ancyrosoma Amyot et Serville 1843 (Figure 16)
Figure 16. Map showing collecting sites of species of genera Ancyrosoma, Derula, Graphosoma and Leprosoma during the present study in Greece.
67.
Ancyrosoma leucogrammes (Gmelin 1790)
Aetoloakarnania [Aetolikon (20.VII.1990), Messolonghion-Tourlida (16.V.1990), Kapsorachi-Trichonis (22.VI.1986)], Arkadia [Xeropigado (9.V.1985)], Arta [Louros Riv. (28.IX.1981)], Attiki [Amaroussion (27.IX.1978), Avlon (4.VII.1978, 12.VII.1979, 16.X.1979), Kifissia (14/31978, 11.VII.1980), Marathon (14.V.1985), Parnis Mt. (13.VII.1984), Schinias (1.VII.1982, 20.V.1983)], Avlis [Vathy (16.VI.1979)], Doris [Eratini (11.VI.1979), Ghiona (16.VII.1978), Kallion (18.VII.1978), Karoutes (18.IX.1978), Skaloula (13.VIII.1977, 18.VI.1978, 20.VII.1978, 17.IX.1978, 15.VIII.1983, 1.VI.1985)], Evia [Agios Georgios (21.VI.1980)], Fokis (Agia Euthymia (12.V.1988)], Ioannina (29.V.1981), Karditsa [Artessiano (1.X.1981)], Konitsa [Aoos Riv. (30.V.1981, 17.VI.1981)], Korinthia [Akrokorinthos (31.VII.1979), Derveni (15.V.1985),], Kynouria [Astros (13.VII.1982, 9.IX.1982)], Messinia [Kazarma (4.VII.1984)], Olympos Mt. [Prionia (13.VIII.1979)], Pieria [Poroi (14.VIII.1980)], Platanos [Parnon Mt. (1.VII.1985)], Thessaloniki [Cedron Hills (23.V.1981)]. Total: 57 specimens.
Genus Derula Mulsant et Rey 1856 (Figure 16)
68.
Derula flavoguttata (Mulsant et Rey 1856)
Attiki [Schinias (23.VI.1986)], Doris [Giona (23.VII.1977, 16.VII.1978), Skaloula (18.VI.1978, 1.VI.1985)], Epirus [Vryssochori (26.V.1981)], Evia [Agios Georgios (21.VI.1980), Gialtra (23.VI.1980)], Florina [Vernon Mt.-Kalo Nero (30.VII.1982), Vevi (25.VIII.1983)], Fokis [Agia Euthymia (12.V.1988)], Korinthia [Derveni (15.V.1985)], Pindos [Vouchorina (13.V.1983)], Rhodos [Salakos (30.V.1990)], Samos [Pythagorion (23.VI.1987)], Trikala [Mourgani (18.VI.1985)]. Total: 23 specimens.
Genus Graphosoma Laporte 1833 (Figure 16)
69.
Graphosoma italicum italicum (Müller 1766)
Aetoloakarnania [Aetolikon (20.VII.1990)], Attiki [Kifissia (9.VIII.1977, 10.I.1978), Schinias (23.IV.1986)], Crete [Rethymnon-Myloi (8.VII.1985)], Doris [Skaloula (14.VII.1978, 7.VIII.1986)], Evrytania [Megalo Chorio (10.VIII.1986)], Fokis [Amfissa (8.VI.1980)] Ioannina [Vikos Gorge (13.VIII.1986)], Konitsa [Aoos-Monastiri (3.V.1981)], Korinthia [Derveni (15.V.1985)], Kozani [Vourinos Mt. (28.VI.1984), (18.VIII.1985)], Lakonia [Mystras (30.IV.1986)], Lesvos [Sykaminea (16.VI.1987)], Olympos Mt. [Stavros (13.VIII.1980)], Pieria [Litochoron (13.VIII.1979)], Rhodos [Apolakkia (30.VI.1987)], Voeotia [Aliartos (12.V.1988)], Oiti Mt. (30.IX.1984, 15.VI.1985). Total: 28 specimens.
70.
Graphosoma semipunctatum (Fabricius 1775)
Attiki [Amaroussion (27.IX.1978), Avlon (16.X.1979), Marathon (14.V.1985), Schinias (23.IV.1986)], Crete [Rethymnon-Agioi Apostoloi (30.VI.1993)], Doris [Skaloula (18.VI.1978)], Kephallinia [Myrtos (25.VI.1988)], Lakonia [Mystras (11.VI.1985)], Oidi Mt. (27.VI.1993), Samos [Neochorion (14.VI.1987)]. Total: 23 specimens.
Genus Leprosoma Baerensprung 1859 (Figure 16)
71.
Leprosoma inconspicuum (Baerensprung 1859)
Doris [Kalion (5.V.1975)]. Total: 1 specimen.
Genus Podops Laporte 1833 (Figure 19)
72.
Podops (Opocrates) curvidens (Costa 1843)
Aetoloakarnania [Loutrakion-Vonitsa (22.VIII.1985), Messolongion-Tourlida (20.VII.1990)], Doris [Eratini (11.VI.1979), Kalion (15.VII.1978, 5.III.1979, 7.X.1979), Monastirakion (29.IV.1981), Skaloula (2.XI.1979)], Pieria [Varikon (23.VII.1982)], Pindos [Korydallos (25.V.1981), Miliotates (5.IX.1980)]. Total: 14 specimens.
73.
Podops (Podops) inunctus* (Fabricius 1775) (Figure 17)
Doris [Kalion (15.VII.1978)], Samothraki [Kamariotissa (5.VI.1982)]. Total: 4 specimens.
Figure 17. Podops (Podops) inunctus (Collector S. Drosopoulos, Kamariotissa, 5.VI.1982, posited in S. Drosopoulos historical collection. Photo A. Tsagkarakis, 12.III.2021).
74.
Podops (Opocrates) rectidens Horvath 1883
Achaia [Bouboukas (9.VI.1980)], Aetoloakarnania [Mytikas-Astakos (23.VI.1986)], Arkadia [Kynouria-Astros (20.V.1982, 5.VII.1984)], Attiki [Schinias (1.VII.1982)], Doris [Eratini (11.VI.1979), Kalion (7.X.1979), Monastirakion (9.VI.1980, 25.X.1980)], Ikaria [Gialiskari (15.VII.1981)], Ilia [Pyrgos-Zacharo (3.VII.1984)], Samothraki [Kamariotissa (5.VI.1982)], Zakynthos [L. Keri (29.VI.1988)]. Total: 23 specimens.
Genus Tarisa Amyot et Serville 1843 (Figure 19)
75.
Tarisa pallescens* (Jakovlev 1871) (Figure 18)
Olympos Mt. [Kryovryssi (30.V.1982)]. Total: 1 specimen.
Figure 18. Tarisa pallescens (Collector S. Drosopoulos, Kryovryssi, 30.V.1982, posited in S. Drosopoulos historical collection. Photo A. Tsagkarakis, 12.III.2021).
Genus Tholagmus Stål 1860 (Figure 19)
Figure 19. Map showing collecting sites of species of genera Podops, Tarisa, Tholagmus and Ventocoris during the present study in Greece.
76.
Tholagmus flavolineatus (Fabricius 1798)
Arkadia [Charadros (17.VI.1986)], Attiki [Avlon (7.VI.1978, 4.VII.1979, 31.V.1989)], Avlis [Vathy (16.VI.1979)], Evia [Agios Georgios (21.VI.1980)], Fthiotis [Kalamakion (18.VIII.1979)], Kephallinia [Assos (25.VI.1988), Fiskardo(24.VI.1988)], Aetoloakarnania [Kapsorachi-Trichonis (22.VI.1986)], Menalon Mt. (20.VI.1986), Parnassos Mt. [National Park (13.VII.1985)]. Total: 18 specimens.
Genus Ventocoris Hahn 1843 (Figure 19)
77.
Ventocoris (Selenodera) achivus (Horvath 1889)
Avlis [Vathy (16.VI.1979)]. Total: 2 specimens.
78.
Ventocoris (Ventocoris) rusticus (Fabricius 1781)
Achaia [Kato Klitoria (20.VI.1986)], Attiki [Marathon (14.V.1985)], Avlis [Vathy (23.V.1979, 16.VI.1979)], Evia [Agios Georgios (21.VI.1980)], Fokis [Amphissa (8.VI.1980)], Kephallinia [Assos (25.VI.1988)], Messinia [Kazarma (4.VII.1984)], Santorini Island [Kamarion (5.V.1982)], Voeotia [Agia Paraskevi (23.VII.1980)]. Total: 23 specimens.
Family PLATASPIDAE
Subfamily Plataspinae
Genus Coptosoma Laporte 1833 (Figure 20)
Figure 20. Map showing collecting sites of species of genus Coptosoma during the present study in Greece.
79.
Coptosoma scutellatum (Geoffroy 1785)
Achaia [Bouboukas (21.VI.1986)], Doris [Kallion (15.VII.1978), Krokylion (21.VI.1978, 19.VII.1978), Skaloula (18.VI.1978)], Florina [Vernon Mt.-Kalo Nero (21.VII.1983)], Ioannina [Voutsaras (24.VI.1984)], Kozani [Vourinos Mt. (18.VIII.1985)], Olympos Mt. [Prionia (13.VIII.1980)], Parnassos Mt. [National Park (13.VII.1985)], Pindos [Korydallos (25.V.1981)], Prespa [Bela Voda (13.X.1986)], Rodopi [Vathyrhemma (26.VII.1982)], Aoos Riv. (27.V.1981). Total: 32 specimens.
Family SCUTELLERIDAE
Subfamily Eurygastrinae
Genus Eurygaster Laporte 1833 (Figure 22)
80.
Eurygaster austriaca (Schrank 1778)
Avlis [Vathy (7.VI.1978, 23.V.1979)], Fokis [Galaxidi (15.IV.1980)], Oiti Mt. (1.VII.1984). Total: 4 specimens.
81.
Eurygaster dilaticollis (Dohrn 1860)
Rodopi Mt. (19.VII.1983), Oiti Mt. (1.VII.1984, 15.VI.1985), Vernon Mt. [Kalo Nero (30.VI.1982)]. Total: 6 specimens.
82.
Eurygaster hottentotta* (Fabricius 1775) (Figure 21)
Kynouria [Kastanitsa (20.VII.1982)]. Total: 1 specimen.
Figure 21. Eurygaster hottentotta (Collector S. Drosopoulos, Kastanitsa, 20.VII.1982, posited in S. Drosopoulos historical collection. Photo A. Tsagkarakis, 12.III.2021).
83.
Eurygaster integriceps (Puton 1881)
Arkadia [Xeropigado (9.V.1985)], Attiki [Marathon (14.V.1978)], Avlis [Vathy (13.IV.1978)], Chios [Agios Georgios (6.IV.1979), Armolia (20.VI.1987)], Doris [Skaloula (29.IV.1979, 17.V.1981)], Epirus [Vryssochori (26.V.1981)], Grevena [Agioi Theodoroi (18.VI.1985), Anoixis (31.VII.1984)], Ikaria [Raches (14.VII.1981)], Ioannina (29.V.1981), Kozani [Vourinos Mt. (29.VI.1984)], Kynouria [Astros (20.V.1982)], Naxos [Apeiranthos (16.VI.1981)], Paros [Paroikia (18.VI.1981)], Samos [Agios Konstantinos (27.VI.1987), Pythagorion (23.VI.1987)]. Total: 31 specimens.
84.
Eurygaster maura (Linnaeus 1758)
Doris [Monastirakion (29.VII.1982), Skaloula (15.VII.1978, 28.X.1981)], Epirus [Vryssochori (27.V.1981)], Evia [Agios Georgios (21.VI.1980)], Florina [Megali Prespa (30.VII.1982), Florina (15.VIII.1979)], Kastoria (13.V.1983), Kynouria [Astros (19.V.1982, 5.VII.1984)], Messinia [Kazarma (4.VII.1984)], Pindos [Korydallos (25.V.1981)], Rhodos [Archipolis (31.V.1990), Messanagros (29.IX.1987)], Rodopi Mt. (25.VII.1982, 10.VIII.1985), Xanthi [Porto Lagos (8.V.1982)], Zakynthos [Keri (29.VI.1988)]. Total: 21 specimens.
85.
Eurygaster testudinaria testudinaria (Geoffroy 1785)
Kastoria (13.V.1983), Rodopi Mt. [Silli (23.VIII.1983)]. Total: 3 specimens.
Genus Psacasta Germar 1839 (Figure 22)
86.
Psacasta (Psacasta) exanthematica exanthematica (Scopoli 1763)
Aetoloakarnania [Nafpaktos (22.VII.1978)]. Total: 1 specimen.
Subfamily Odontoscelinae
Genus Irochrotus Amyot et Serville 1843 (Figure 22)
87.
Irochrotus maculiventris (Germar 1839)
Achaia [Bouboukas (21.VI.1986)], Arkadia [Tripolis (19.VI.1986)], Attiki [Avlon (26.V.1978)], Doris [Skaloula (1.VI.1985)], Evia [Agios Georgios (21.VI.1980), Mantoudi (28.X.2007)], Ioannina [Voutsaras (24.VI.1984)]. Total: 8 specimens.
Genus Odontoscelis Laporte 1833 (Figure 22)
88.
Odontoscelis (Odontoscelis) fuliginosa (Linnaeus 1761)
Attiki [Avlon (11.V.1980), Parnis Mt. (29.V.1985)], Karpathos [Messochori (4.VI.1990)], Kephallinia [Aenos Mt. (23.VI.1988)]. Total: 7 specimens.
89.
Odontoscelis (Odontoscelis) lineola (Rambur 1839)
Doris [Skaloula (17.IX.1978)] Arkadia [Charadros (17.VI.1989), Karoutes (18.IX.1978)], Vegoritis Lake (14.VIII.1979). Total: 4 specimens.
Subfamily Odontotarsinae
Genus Odontotarsus Laporte 1833 (Figure 22)
Figure 22. Map showing collecting sites of species of genera Eurygaster, Psacasta, Irochrotus, Odontoscelis and Odontotarsus during the present study in Greece.
90.
Odontotarsus purpureolineatus (Rossi 1790)
Achaia [Bouboukas (21.VI.1986)], Aetoloakarnania [Astakos (14.VI.1984), Mytikas-Astakos (23.VI.1986)], Arkadia [Charadros (17.VI.1986)], Attiki [Avlon (7.VI.1978), Marathon (11.VII.1984, 14.V.1985)], Chios [Armolia (20.VI.1987)], Doris [Skaloula (20.VII.1986)], Epirus [Vryssochori (26.V.1981)], Evros [Metaxades (2.VI.1982)], Grevena [Anoixis (31.VII.1984)], Ilia [Kyllini Mt. (21.VII.1982)], Ioannina [Voutsaras (24.VI.1984)], Konitsa [Aoos Riv.-Monastiri (29.V.1981)], Kozani [Vourinos Mt. (18.VIII.1983), (28.VI.1984)], Lesvos [Petra (17.VI.1987)], Messinia [Artemissia (11.VI.1985), Kazarma (4.VII.1984)], Olympos Mt. [Prionia (21.V.1981)], Pilion Mt. [Chania 16.VIII.1980)], Rhodos [Petaloudes) (29.V.1990), Rodopi Mt. [Livaderon (10.VIII.1985), Sidironero (23.V.1983)], Samos [Neochorion (24.VI.1987), Pythagorion (23.VI.1987)], Trikala [Mourgani (19.VI.1985)], Voeotia [Tsoukalades (12.V.1988)], Parnon Mt. (11.VI.1985). Total: 36 specimens.
91.
Odontotarsus robustus (Jakovlev 1883)
Attiki [Avlon (16.VII.1979), Kifissia (11.VII.1980)], Doris [Skaloula (20.VII.1978, 17.IX.1978, 9.VI.1979, 1.VI.1985)], Pindos [Votonossi (5.IX.1980), Konitsa (22.VI.1986)]. Total: 16 specimens.
92.
Odontotarsus rufescens (Fieber 1861)
Amphilochia [Anoixiatiko (20.VIII.1985)], Attiki [Avlon (20.V.1987)], Evia [Agios Georgios (21.VI.1980)], Ilia [Krestena (16.V.1985)], Kynouria [Astros (13.VII.1982)], Messinia [Kazarma (4.VII.1984)]. Total: 7 specimens.
Table 1. Updated checklist of Pentatomoidea of Greece.

4. Discussion

In Greece, until the present work, very few studies have been conducted on the Pentatomoidea fauna, in which 150 Pentatomoidea species are referred. Except two overall studies performed by Drosopoulos [43] and Ramsey [44], in 1980 and 2019, respectively, only sporadic references are made by numerous researchers, considering the Greek fauna as part of a wider Balkans. In the present study, 8 species are recorded in Greece for the first time, raising the number of the Greek fauna to 158 species. It has to be mentioned that the new records refer to specimens that were collected during 1982–1983 and placed in the historic collection of Sakis Drosopoulos. Our study provides one step toward enriched knowledge of the Pentatomoidea fauna of Greece, which will hopefully trigger more comprehensive taxonomic study of the entire superfamily.

Author Contributions

Conceptualization, A.T. and S.D.; methodology, A.T. and Z.T.; validation, A.T., Z.T., A.C., I.M. and A.K.; investigation, A.T. and Z.T.; data curation, A.C., Z.T.; writing—original draft preparation, A.T., Z.T. and A.C.; writing—review and editing, A.T., Z.T.; visualization, A.T.; supervision, A.T.; project administration, A.T. All authors have read and agreed to the published version of the manuscript.

Funding

This research received no external funding.

Institutional Review Board Statement

Not applicable.

Data Availability Statement

Data is contained within the article.

Conflicts of Interest

The authors declare no conflict of interest.

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