Understory Plant–Soil Carbon Coupling in Agroforestry Systems

A special issue of Plants (ISSN 2223-7747). This special issue belongs to the section "Plant–Soil Interactions".

Deadline for manuscript submissions: 31 December 2026 | Viewed by 279

Editor


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Guest Editor
State Key Laboratory for Development and Utilization of Forest Food Resources, Zhejiang A&F University, Hangzhou 311300, China
Interests: forest soil; ecological stoichiometry; plant–soil carbon

Special Issue Information

Dear Colleagues,

As global agroforestry expands to meet climate mitigation and food security goals, the understory vegetation layer—encompassing medicinal plants, mushroom cultivation beds, and strategic cover crops—has emerged as a critical yet understudied driver of soil carbon dynamics. Unlike traditional monocultures, these multi-strata systems create unique plant–soil interfaces where belowground carbon processes are tightly regulated by the specific functional traits of understory species. These include root exudation chemistry, mycorrhizal association strategies, and the stoichiometric quality of litter.

This Special Issue invites contributions that explore the multi-dimensional coupling between understory vegetation and soil carbon processes across diverse agroforestry systems. We welcome the submission of research investigating how distinct plant functional groups—ranging from nitrogen-fixing legumes and deep-rooted cover crops to medicinal herbs and edible fungi—modulate soil carbon inputs via root exudation, rhizodeposition, and litterfall quality. Studies elucidating the mechanistic pathways of carbon stabilization are particularly encouraged, including physical protection within soil aggregates, chemical recalcitrance of plant-derived compounds, and microbial carbon use efficiency driven by the assembly of fungal–bacterial communities. We also encourage integrative work across spatial scales, from rhizosphere microhotspots to ecosystem-level carbon budgeting, and across climatic gradients, including subtropical, temperate, and tropical agroforestry plantations. Contributions employing isotope tracing, metabolomics, high-throughput sequencing, long-term chromosequence approaches, or models of carbon sequestration in the soil with Century or RhotC, deciphering carbon flow from plant traits to persistent soil pools, are highly valued. By synthesizing perspectives from plant ecophysiology, soil microbiology, and ecosystem ecology, this collection aims to advance evidence-based management strategies that enhance belowground carbon sequestration while sustaining the productive capacity of multi-strata cultivation systems.

Prof. Dr. Jiasen Wu
Guest Editor

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Keywords

  • agroforestry systems
  • understory vegetation
  • rhizosphere processes
  • soil carbon stabilization
  • microbial carbon use efficiency
  • root functional traits
  • mycorrhizal networks

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Published Papers (1 paper)

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Research

31 pages, 9640 KB  
Article
Moss Cover Redirects Soil Organic Carbon from Active Turnover to Mineral-Associated Stabilization in Subalpine Forests
by Jiahui Huang, Xiaoyu Zhang, Yu Tian, Guo Luo, Dajun Xie, Jinxiao Li, Baoli Duan and Shuming Peng
Plants 2026, 15(13), 2098; https://doi.org/10.3390/plants15132098 - 6 Jul 2026
Abstract
Understory mosses modify near-surface soil conditions, but how elevation regulates their influence on active and mineral-associated soil organic carbon (SOC) remains unclear. We compared independently selected moss-covered and non-moss-covered soils across a 3200–3500 m elevational gradient and integrated soil physicochemical measurements, microbial biomass [...] Read more.
Understory mosses modify near-surface soil conditions, but how elevation regulates their influence on active and mineral-associated soil organic carbon (SOC) remains unclear. We compared independently selected moss-covered and non-moss-covered soils across a 3200–3500 m elevational gradient and integrated soil physicochemical measurements, microbial biomass (MB), dissolved organic matter (DOM), microbial necromass carbon (MNC), particulate organic carbon (POC), mineral-associated organic carbon (MAOC), metagenomic profiling, and piecewise structural equation modeling. Moss-covered soils consistently contained higher SOC and MAOC, but lower DOM, MB, and generally lower POC, than non-moss-covered soils. MNC showed an elevation-dependent reversal, with higher values under moss cover at 3200 m but lower values under moss cover at 3300–3500 m. Elevation was not a significant uniform driver of MB, DOM, MNC, POC, or MAOC; instead, its influence was mainly reflected in interactions with surface cover and in elevation-related changes in moss-layer structure, diversity, and hydrothermal conditions. Core carbon-fixation and degradation functions remained broadly stable, whereas specific functional modules shifted within moss-covered soils: acetate and acetyl-CoA metabolism genes (ackA and abfD) were relatively abundant at 3300–3400 m, while the polysaccharide-reprocessing gene SGA1 and oxidative-transformation gene katG increased toward higher elevations, and pmoC/amoC rebounded at 3500 m. Structural equation models linked the microbial functional gene system more strongly to POC, whereas MNC was positively associated with MAOC, and the direct POC-to-MAOC pathway was not significant. These findings indicate that moss cover is associated with contrasting SOC allocation patterns and stronger microbial necromass–MAOC coupling, while elevation modulates these relationships indirectly through changes in moss communities, soil microenvironment, and microbial functional potential. Full article
(This article belongs to the Special Issue Understory Plant–Soil Carbon Coupling in Agroforestry Systems)
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