Quaternary, Volume 7, Issue 4 (December 2024) – 17 articles

Coste San Giacomo (CSG) represents a significant paleontological site to investigate the faunal and environmental changes that occurred in Mediterranean Europe during the Early Pleistocene. In this work, we described for the first time the Carnivoran assemblage. We ascribed the fossil remains to the following taxa: Ursus sp., Homotherium latidens, Canis etruscus, Pliocrocuta perrieri, Martellictis ardea and Vulpes alopecoides. Considering the value of the carnivoran taxa here identified, we discuss their particular biochronological significance, since the CSG site records the last occurrence of P. perrieri and the first occurrences of H. latidens, C. etruscus, M. ardea and V. alopecoides for the Italian Peninsula. These results will allow us to improve the data of the biochronological scheme of the Villafranchian European Land Mammal Age, recognizing the earliest dispersals and latest occurrences across Europe. Full article

The fortified hilltop settlement of Monkodonja, located near Rovinj on the west coast of Istria, Croatia, provides insight into Bronze Age occupation and conflict in the Adriatic region. Established around 2000 BC, as evidenced by a series of C14 dates from human and animal bones, the settlement experienced significant construction phases, particularly in its defensive architecture. Its earliest fortifications, built with limestone blocks using dry-stone wall techniques, date to the 19th century BC, with major expansions in the 16th century BC, where the primary wall was doubled in width and reached over 3 m in thickness. Monkodonja’s architectural complexity, notably the West Gate and Acropolis fortifications, and certain types of artifacts reveal influences from southern regions such the eastern Aegean. However, the settlement appears to have met a violent end around the 15th century BC, suggested by destruction layers, widespread burning, and the presence of weapons such as a lance tip, bronze axe, and slingstones. Monkodonja’s destruction raises questions about broader military conflicts in the Adriatic region during this period. Possible causes could include localized warfare or connections to larger-scale disturbances. Research in Monkodonja is also significant in the context of the debate surrounding the emergence of the so-called Castellieri settlements in Istria at the beginning of the 2nd millennium BC. As early as the beginning of the 20th century, it was proposed that a migration of people to the Istrian peninsula brought this new settlement form and other influences, leading to a significant population increase. The appearance of the Castellieri settlement form coincides with a period marked by documented climatic changes and two major natural disasters in the form of volcanic eruptions. Full article

This article is devoted to the study of the Quaternary aeolian sands of the boreal zone of north Asia. Using the example of the study reference sections of the Selenga Dauria (Western Transbaikalia), it was established that the activation of aeolian processes is determined by the complex interaction of natural and anthropogenic factors. Natural factors include neotectonic movements; wide distribution of alluvial and lacustrine-alluvial deposits; a sharply continental semi-arid climate; and forest-steppe and steppe vegetation. Among the anthropogenic factors, the leading ones are deforestation, plowing of land and construction of new settlements, roads and other line structures. The obtained radiocarbon dating of buried soils and coal from ancient fire pits indicates the activation of aeolian processes during the Holocene. The main sources for aeolian transport (winnowing) are sands located in the areas of river and lake beaches, floodplains and river terraces. Almost all aeolian sands of the boreal zone were formed as a result of short-range wind transport. They form mini-deserts unfixed by vegetation, with active aeolian processes, dunes, barkhans and deflationary basins. Aeolian swells and blowout basins characterize aeolian landscapes weakly fixed by vegetation. It is noted that aeolian deposits of the boreal zone of north Asia, in contrast to similar sands of the subtropical and tropic zones, consist of coarser-grained material. Medium- and fine-grained sands dominate their composition, which is polymineral and well-sorted. In subtropical and tropical deserts, they are predominantly monomineral, fine and fine-grained. At the same time, mainly minerals that are unstable to weathering (feldspars, plagioclases, pyroxenes and amphiboles) represent the mineralogical composition of the studied aeolian sands. Weathering-resistant minerals dominate the sands of classical deserts: quartz, leucoxene, ilmenite, epidote, zircon, garnets, tourmaline, rutile and others. Modern aeolian landscapes are a unique natural formation for the boreal zone of north Asia and can be successfully used for the development of ecotourism. Full article

This research presents an in-depth analysis of large mammal remains first discovered in 1932 in the archaeological area of ancient Rome, central Italy, during the work for the opening of Via dell’Impero (VFI). This work describes the faunal assemblage, its current preservation status, and uses tephrochronology to assess its age. Additionally, it provides paleoecological insights into the evolution of the mammalian fauna in Latium, central Italy, from MIS 13 to MIS 7. Analysis of the fossils updates the identification previously proposed by De Angelis d’Ossat, confirming the presence of Palaeoloxodon antiquus, Cervus elaphus, and Bos primigenius. However, in contrast to the previous author, the hippopotamus remains are assigned to Hippopotamus cf. antiquus, and a second deer is identified as Dama sp.. Furthermore, gnawing marks on the hippopotamus femur suggest the presence of a middle-sized carnivore. Tephrochronological investigation was conducted on pumice retrieved from the VFI fossiliferous layer and ash extracted from sediments adhering to the fossil surfaces. The major element composition of the glass from all pumice/ash samples shows a strong affinity with the Vico β unit, allowing correlation with the Fucino record and constraining the deposition of the VFI fossiliferous level between <406.5 ± 1.3 ka and >405.7 + 1.5/−1.6 ka. Radiometric dating is particularly useful for large mammal faunas of MIS 11-MIS 7, a period lacking significant faunal renewals, as Latium mammalian faunas are often dominated by species (elephants, red deer, aurochs) with broad chronological ranges. Full article

A detailed high-resolution study of climate and vegetation changes in two sedimentary profiles from the Godavari delta in India was conducted to understand extreme climate variability over the last 3000 years. These historical climate records are vital for predicting future climate changes influenced by increased human activity. Pollen records reveal several climatic periods: (1) an initial warm/humid phase (2260–1560 cal yr BP) with mangrove and tropical tree pollen and maximum pollen accumulation rate (PARmax), (2) a gradual shift to drier conditions with reduced mangrove and arboreal pollen, coinciding with to the Dark Ages Cold Period (1580–1070 cal yr BP), (3) a Medieval Warm Period (1090–580 cal yr BP) marked by a stronger summer monsoon, rejuvenation of mangroves and tree pollen, (4) a relatively cool and dry phase with abundant dry deciduous and non-arboreal pollen possibly linked to the Little Ice Age (580–80 cal yr BP), showing a weakened monsoon, and (5) recent warming since 80 cal yr BP. A semi-quantitative aridity, temperature, and moisture index based on pollen concentration variations have also been used to identify centennial-scale climatic cycles in the pollen record. Full article

We present multiproxy records from a 2.25-m-long lake sediment profile from central India, which suggested that between ~22,200 and 18,658 cal yr BP, the Indian Summer Monsoon (ISM) was weak, supporting open vegetation in a cool and dry climate, which is globally correlated with the Last Glacial Maximum (LGM). The grain size data of this phase suggest low-energy conditions, indicating a weak ISM. Environmental magnetic concentration-dependent parameters also confirm this weakened ISM. Between ~18,658 and 7340 cal yr BP, the ISM underwent a notable increase, and open mixed tropical deciduous forests replaced the existing vegetation under a warm and moderately humid climate. Environmental magnetic parameters and the grain size data signal a shift toward higher energy levels, in harmony with the warm and moderately humid climate during this time span. Between ~7340 and 1960 cal yr BP, the ISM intensity further increased, which supported open mixed tropical deciduous forests with a rise in prominent tree species under a warm and a relatively more humid climate, correlated with the global Holocene Climatic Optimum (HCO). The trends in environmental magnetic parameters and grain size data mirror this phase of climatic amelioration. From ~1961 cal yr BP to the present, the ISM has intensified, giving rise to dense mixed tropical deciduous forests under a warm and relatively more humid climate. Environmental magnetic parameters and the grain size data are in tandem with the palynogical findings from this phase of the ISM variability. Full article

On 2015, after the direct study of the most important Late Villafranchian fossil collections of Europe and Western Asia, including Orce (Spain), Pirro Nord and Upper Valdarno (Italy), Appollonia (Greece), Dmanisi (Georgia) and ‘Ubeidiya (Israel), among others, our team proposed the hypothesis that suids disappeared from Europe during the time span between 1.8 and 1.2 Ma. The implications of our conclusions were significant, the arrival of Early Homo into Western Europe, dated to 1.4 Ma at the site of Barranco León in Orce (Spain), preceded the return of pigs into the continent at 1.2 Ma. This hypothesis has been recently challenged because of the finding of an incomplete metatarsal ascribed to Sus sp., with no clear stratigraphic origin, found in the XIX Century Croizet collection of Peyrolles (France), which is housed in the Natural History Museum, London, together with other weak arguments based on the absence of reliable dating for many Early Pleistocene European sites, and other hypothetical records of pigs, with no real fossil support. We answer all these questions and defend that our 2015 hypothesis is correct. Full article

In the Afrotropic biogeographic realm, with its diverse and high-density mammal population, early humans may have been hunting with stone-tipped weapons since ~500,000 years ago. Being able to hunt effectively from a distance has several important adaptive advantages. Yet, until now, African long-range javelin hunting remained unexplored as intermediate between short/medium-range, hand-delivered and long-range, mechanically projected weapons. Insights gained from a new Afrotropic comparative dataset with 950 weapon tips of known use—including several javelin types—provide a contextually appropriate middle-range tool for assessing the probable effective hunting ranges of Middle Stone Age points. We use a novel application of the ballistically relevant tip cross-sectional area (TCSA) statistic to define contact, short-, medium-, long- and maximum-range hunting and discuss the adaptive advantages for each. The approach is applied to suggest developments and variations in the best-fit hunting ranges of 5597 stone points from 62 Middle Stone Age Afrotropic assemblages. By aligning our results with the Marine Isotope Stage (MIS) record we hypothesize that effective long-range (~20–30 m) hunting with stone-tipped weapons was probably not practiced by ≥MIS 8, and that experimentation with long-range javelins—similar to those used by contemporary Ethiopian hunters—over these distances may have started during MIS 6, becoming part of the everyday Afrotropic hunting arsenal by the end of MIS 5. Full article

Collecting is a form of leisure, and even a passion, consisting of collecting, preserving and displaying objects. When we look for its origin in the literature, we are taken back to “the appearance of writing and the fixing of knowledge”, specifically with the Assyrian King Ashurbanipal (7th century BC, Mesopotamia), and his fondness for collecting books, which in his case were in the form of clay tablets. This is not, however, a true reflection, for we have evidence of much earlier collectors. The curiosity and interest in keeping stones or fossils of different colors and shapes, as manuports, is as old as we are. For decades we have had evidence of objects of no utilitarian value in Neanderthal homes. Several European sites have shown that these Neanderthal groups treasured objects that attracted their attention. On some occasions, these objects may have been modified to make a personal ornament and may even have been integrated into subsistence activities such as grinders or hammers. Normally, one or two such specimens are found but, to date, no Neanderthal cave or camp has yielded as many as the N4 level of Prado Vargas Cave. In the N4 Mousterian level of Prado Vargas, 15 specimens of Upper Cretaceous marine fossils belonging to the Gryphaeidae, Pectinidae, Cardiidae, Pholadomyidae, Pleurotomariidae, Tylostomatidae and Diplopodiidae families were found in the context of clay and autochthonous cave sediments. During MIS 3, a group of Neanderthals transported at least fifteen marine fossils, which were collected from various Cretaceous units located in the surrounding area, to the Prado Vargas cave. The fossils, with one exception, show no evidence of having been used as tools; thus, their presence in the cave could be attributed to collecting activities. These activities could have been motivated by numerous tangible and intangible causes, which suggest that collecting activities and the associated abstract thinking were present in Neanderthals before the arrival of modern humans. Full article

Humans are well known to have made paintings and engravings on rock surfaces, both geometric motifs with an unclear representation, and representative motifs that refer to their activities and aspects of their environment. This kind of art is widespread across time and space and has throughout history been subjected to various kinds of approaches. Typically, rock art research focuses on its role in the development of the hominin brain and the capability of abstract thinking, as well as on interpreting representative and non-representative motifs. Ethnography and cognitive research have often stressed that rock art is the result of ritual practises and the expression of a shamanic belief system. However, representative motifs may also shed light on a region’s ecological and human prehistory. Here, we give an overview of the general development of rock art study: we highlight the development of artistic behaviour in humans by discussing aesthetic preferences, and the creation of simple geometric motifs and eventually representative motifs, before describing the theories that developed from the earliest study of rock art. These have largely focused on classification and interpretation of the motifs, and often centred on Palaeolithic material from Europe. We then move on to discuss how ethnography among rock art creating communities often suggests important relationships between specific animals in both the realms of spiritual belief systems and within the local environment. Lastly, we highlight how rock art reflects the local penecontemporaneous environment when it comes to depictions of animals, plants, technologies, humans and their activities. We argue that animal depictions are a useful subject to study on a large scale, as it is the most widespread representative motif, and the most appropriate subject to study when the goal is to draw conclusions on environmental changes. Rock art can fill gaps in the local archaeological record and generate new questions of it, but also offer new insights into the history of local human–animal interaction: animal species depicted and/or referred to in rock art are likely to have been a selection of spiritually important animals and a comparison to known information on human interactions with local species may reveal patterns among which animals are selected for local rock art depictions and which are not. Interregional comparison can in turn shed light on whether humans in general tend to ascribe meaning to the same types of animals. We end the review with suggestions for future study, with a special role for computational methods, which are suitable for the analysis of large databases of visual imagery. Full article

A 7.4 m long sediment core has been retrieved from the central part of Lake Vrana on the island of Cres to reconstruct the paleoenvironmental conditions. Lake Vrana is the deepest freshwater lake in Croatia, located in the karst region of the eastern Adriatic coast. A dated sediment sequence in Lake Vrana of 4.4 m has spanned the past 16.4 kyr, and it featured a dynamic sediment deposition until the beginning of the Holocene, including strong sediment input and supply to the lake by runoff sediments of dolomitic origin from the catchment in the period 16.4–14.4 cal kyr BP. High organic carbon content, which originates from mixed terrestrial and aquatic origins in the periods 14.4–13.3 cal kyr BP and 12.7–11.7 cal kyr BP, indicates fluctuating lake levels in shallow water environments during the Late Glacial to Holocene transition. The Holocene sequence indicates the development of more stable conditions and continuous sediment deposition, characterized by an increasing trend of siliciclastic sediments delivered into the lake during the early Holocene (11.7–10 cal kyr BP) and dominantly from 8 to 4.4 cal kyr BP, indicating enhanced input and erosion, which coincides with the humid and pluvial period recorded in the central Mediterranean region. It is followed by sediments with high organic carbon content between 4.4 and 1.6 cal kyr BP, which points to higher lake productivity. Calcite sedimentation prevailed between 1.6 to 0.4 cal kyr BP, indicating stable deeper-lake conditions. Predominantly, siliciclastic sediments from 0.4 to 0.1 cal kyr BP pointed to erosion during the Little Ice Age (LIA), with enhanced precipitation and sediment discharge from the catchment. The re-establishment of calcite sedimentation has been observed over the last 100 years. Full article

Quaternary stream sediments and beach black sand in north-western Saudi Arabia (namely Wadi Thalbah, Wadi Haramil and Wadi Al Miyah) are characterized by the enrichment of heavy minerals. Concentrates of the heavy minerals in two size fractions (63–125 μm and 125–250 μm) are considered as potential sources of “strategic” accessory minerals. A combination of mineralogical, geochemical and spectroscopic data of opaque and non-opaque minerals is utilized as clues for provenance. ThO₂ (up to 17.46 wt%) is correlated with UO₂ (up to 7.18 wt%), indicating a possible uranothorite solid solution in zircon. Hafnoan zircon (3.6–5.75 wt% HfO₂) is a provenance indicator that indicates a granitic source, mostly highly fractionated granite. In addition, monazite characterizes the same felsic provenance with rare-earth element oxides (La, Ce, Nd and Sm amounting) up to 67.88 wt%. These contents of radionuclides and rare-earth elements assigned the investigated zircon and monazite as “strategic” minerals. In the bulk black sand, V₂O₅ (up to 0.36 wt%) and ZrO₂ (0.57 wt%) are correlated with percentages of magnetite and zircon. Skeletal or star-shaped Ti-magnetite is derived from the basaltic flows. Mn-bearing ilmenite, with up to 5.5 wt% MnO, is derived from the metasediments. The Fourier-transform infrared transmittance (FTIR) spectra indicate lattice vibrational modes of non-opaque silicate heavy minerals, e.g., amphiboles. In addition, the FTIR spectra show O-H vibrational stretching that is related to magnetite and Fe-oxyhydroxides, particularly in the magnetic fraction. Raman data indicate a Verwey transition in the spectrum of magnetite, which is partially replaced by possible ferrite/wüstite during the measurements. The Raman shifts at 223 cm⁻¹ and 460 cm⁻¹ indicate O-Ti-O symmetric stretching vibration and asymmetric stretching vibration of Fe-O bonding in the FeO₆ octahedra, respectively. The ultraviolet-visible-near infrared (UV-Vis-NIR) spectra confirm the dominance of ferric iron (Fe³⁺) as well as some Si⁴⁺ transitions of magnetite (226 and 280 nm) in the opaque-rich fractions. Non-opaque heavy silicates such as hornblende and ferrohornblende are responsible for the 192 nm intensity band. Full article

Two zoogenic deposits from the central part of the Negev Desert (Israel) were investigated by stable isotopes (carbon ¹³C/¹²C and nitrogen ¹⁵N/¹⁴N) and pollen analyses. The merger of these data and results of radiocarbon dating of Atzmaut and Ramon I deposits enabled us to reconstruct climate and vegetation changes in the Negev Desert over the past 8500 years. Decrease of the δ¹³C value in plant remains is a sensitive indicator of paleoclimatic conditions in the region. The decline of the δ¹³C value over the past 8500 years almost fully coincides with an increase of the total pollen concentration, the proportion of Poaceae pollen in the profile of zoogenic deposits and wetter periods. Thus, four humid periods are identified, from the middle of the 4th millennium to the end of 3rd millennium BC, the 1st half of the 2nd millennium BC, from the 2nd half of the 1st millennium BC to the 1st half of the 1st millennium AD and the middle of the 2nd millennium AD. The δ¹⁵N value of plant remains is a less sensitive indicator of climate dynamics and represents the most significant change of precipitation in the region by the end of the 3rd millennium BC. Full article

We present the reconstruction of palaeoenvironmental conditions in the Gdansk, Bornholm, and Arkona Basins of the Baltic Sea over the last millennium. A multiproxy study (including geochemical, XRF, grain size, AMS, and micropalaeontological analyses) of five short sediment cores was performed. The relative age of the sediments was determined based on the Pb distribution along the sediment sequences, as radiocarbon dating has resulted in an excessively old age. The retrieved cores cover two comparable warm periods, the Medieval Climate Anomaly and the Modern Warm Period, for which the increase in surface water productivity was reconstructed. Notably, the production of diatoms was higher during the colder periods (the Dark Ages and Little Ice Age), but this was also the case within the Modern Warm Period. In the Gdansk Basin, the initial salinity increase during the Littorina transgression started after 7.7 cal. a BP. The increased inflow activity was reconstructed during the Medieval Climate Anomaly, even in the Gdansk Basin, despite, in general, very low foraminiferal amounts and diversity. The strongly positive North Atlantic Oscillation Index during this period led to the prevalence of westerly winds over the Baltic region and stronger saltwater intrusions. In the recent sediments, the reconstructed inflow frequency demonstrates a variability against the reduction trend, and a general decline compared to the Medieval Climate Anomaly is seen. Full article

The Villafranchian stage in the mammal fauna evolution in Eurasia (ca. 3.6/3.4 Ma—ca. 1.2 Ma) is associated with the beginning of the formation of the modern appearance of the mammal megafauna of today’s Palaearctic. The cooling and the aridification starting with the beginning of the Early Pleistocene gradually eliminated the quasi-tropical appearance of the Late Neogene landscapes and fauna of Europe. The time from the Mid-Piacenzian (ca. 3.3–3.0 Ma) to the end of the Early Pleistocene was a time of particularly intense dispersal of species, of faunal exchange between Eurasia and Africa, and of the entry of new mammals into Europe from the East. That is why the correlation of the biochronology of the Villafranchian fauna between Eastern and Western Europe is of particular interest. Accumulated data make possible a more precise correlation of these faunas today. A correlation of selected Eastern European localities with established faunal units and MNQ zones is made in the present work. Usually, the dispersal from Asia or from E. Europe to W. Europe is instantaneous from a geological point of view, but in a number of cases, reaching W. Europe happens later, or some species known to be from Eastern Europe do not reach Western Europe. The main driving forces of the faunal dispersals, which are the key bioevents in the faunal formation, are climate changes, which in turn, affect the environment. We can summarize the following more significant Villafranchian bioevents in Europe: the End Pliocene (Early Villafranchian: MNQ16) turnover related to the first appearance of a number of taxa, for example, felids, canids, proboscideans, and ungulates; the Quaternary beginning turnover. Correlated with this are the beginning of the Middle Villafranchian, which should be placed at about 2.6 Ma; the Coste San Giacomo faunal unit turnover (Senèze and Slivnitsa localities should be included here, and the FU itself, at the very beginning of the late Villafranchian (=MNQ18a)); the Pachycrocuta event at the very beginning of the Olivola FU; and the events related to the Late Villafranchian/Epivillafranchian bounfary. Full article

The known Plio-Pleistocene mammalian record, mainly represented by small mammals, and its biotic and geological context in the vast region of Eastern Turkey and Transcaucasus provides a sound base for regional biochronology. Recently obtained faunal associations and the main evolutionary lineages found in the region support direct correlations to the European (ELMA/MN/MQ) and the Eastern European (faunal complexes/MQR-MNR) biochronological systems. Important data on palynology, aquatic and terrestrial mollusks, and magnetostratigraphy integrate the reviewed material into a robust local biochronology. The range of standard biochrons of Early Pliocene through late Early Pleistocene and the regional Anatolian zones M-P are reliably detected. The Early Pleistocene time range (zone P) is refined based on rhizodont lagurines Borsodia and Euro-Asian larger voles Mimomys ex gr. pliocaenicus. The successive zone R for Early Pleistocene faunas with early rootless Microtini is proposed. Full article

Recent investigations in the upper Lower–Middle Pleistocene deposits of the Megalopolis Basin (Greece) led to the discovery of several sites/findspots with abundant faunal material. Here, we provide an updated overview including new results on the micro- and macro-mammal fauna. Important new discoveries comprise partial hippopotamus skeletons from Marathousa 1 and the new Lower Pleistocene site Choremi 6, as well as a second partial elephant skeleton from Marathousa 1, including a complete tusk and the rarely found stylohyoideum. Based on the first results from the newly collected micromammals, we discuss age constraints of the sites, and we provide biostratigraphic/biochronologic remarks on key mammal taxa for the Middle Pleistocene of Greece and southeastern Europe. The presence of mammals highly dependent on freshwater for their survival, together with temperate-adapted ones in several stratigraphic layers of the basin, including those correlated with glacial stages, when conditions were colder and/or drier, indicate the capacity of the basin to retain perennial freshwater bodies under milder climatic conditions, even during the harsher glacial periods of the European Middle Pleistocene, and further support its refugial status. Yet, the smaller dimensions of the Megalopolis hippopotamuses may represent a response to the changing environmental conditions of the epoch, not optimal for hippopotamuses. Overall, the Megalopolis Basin comprises a unique fossil record for southeastern Europe and provides valuable insights into the Middle Pleistocene terrestrial ecosystems of Europe, and hominin adaptations in particular. Full article