1
BioProcess Engineering Research Center, KAIST, 291 Daehak-ro, Yuseong-gu, Daejeon 305-701, Korea
2
Department of Chemical & Biological Engineering, Chungnam National University, 99 Daehangno, Yuseong-gu, Daejeon 305-764, Korea
3
Center for Nanobio Integration & Convergence Engineering, National Nanofab Center, 291 Daehak-ro, Yuseong-gu, Daejeon 305-806, Korea
4
Department of Chemical & Biomolecular Engineering (BK21 Program), Department of Bio & Brain Engineering, Department of Biological Sciences, Bioinformatics Research Center, Center for Systems & Synthetic Biotechnology, and Institute for the BioCentury, KAIST, 291 Daehak-ro, Yuseong-gu, Daejeon 305-701, Korea
5
Department of Microbiology & Research Institute for Medical Science, Chungnam National University, 6 Moonhwa-dong, Jung-gu, Daejeon 301-747, Korea
6
Biotechnology Research Division, National Fisheries Research & Development Institute (NFRDI), 408-1 Sirang-ri, Gijang, Busan 619-705, Korea
7
Department of Chemistry, Chung-Ang University, 84 Heukseok-ro, Dongjak-gu, Seoul 156-756, Korea
†
Theses authors contributed equally to this work.