Search Results (15)

Crickets use the pulse pattern of the species-specific calling song as a primary cue for mate recognition. Here we combined transcriptome profiling of brain regions with network-based analyses in Gryllus bimaculatus exposed to silence or pulse trains known to elicit strong or weak phonotactic attraction. Acoustic stimulation triggered specific transcriptional changes in the brain, with the anterior protocerebrum showing the most pronounced and selective responses to the calling song pattern, characterized by enrichment in neuromodulatory and neurotransmitter-related pathways. Weighted gene co-expression analysis identified a specific cluster of highly co-expressed genes in the anterior brain (termed the calling song-responsive module) that responded selectively only to the calling song stimulus. Genetic network topology analysis revealed six highly connected key hub genes within the calling song-responsive module—GbOrb2, Gbgl, Gbpum, GbDnm, GbCadN, and GbNCadN. These genes showed extensive interactions with many other genes in the network, suggesting their central regulatory role in response to calling song in female crickets. These findings support the anterior brain as a central integrator of cricket auditory mate recognition cues and point to a core molecular network that likely underpins this behavior. Full article

This work investigates how different language modeling techniques affect the performance of an end-to-end automatic speech recognition (ASR) system for the Amazigh language. A (CNN-BiLSTM-CTC) model enhanced with an attention mechanism was used as the baseline. During decoding, two external language models were integrated using shallow fusion: a trigram N-gram model built with KenLM and a recurrent neural network language model (RNN-LM) trained on the same Tifdigit corpus. Four decoding methods were compared: greedy decoding; beam search; beam search with an N-gram language model; and beam search with a compact recurrent neural network language model. Experimental results on the Tifdigit dataset reveal a clear trade-off: the N-gram language model produces the best results compared to RNN-LM, with a phonetic error rate (PER) of 0.0268, representing a relative improvement of 4.0% over the greedy baseline model, and translates into an accuracy of 97.32%. This suggests that N-gram models can outperform neural approaches when reliable, limited data and lexical resources are available. The improved N-gram approach notably outperformed both simple beam search and the RNN neural language model. This improvement is due to higher-order context modeling, its optimized interpolation weights, and its adaptive lexical weighting tailored to the phonotactic structure of the Amazigh language. Full article

Phonemic processing is largely influenced by how stimulus-specific characteristics of a language are computed, but person-level variables represent important moderators as well. The current study investigates how such characteristics, in parallel, affect receptive-level phonemic processing across the preschool age, and whether these effects are comparable across different languages. Using a child-friendly ‘odd-man-out’ discrimination task, we examined 239 monolingual German- and French-speaking preschoolers, aged three to five. Results revealed that phonotactic probability-based syllable frequency, nonword length, and mismatching nonword positioning effects explained independent variance components of phonemic processing. Age significantly affected how memory-related, but not linguistically relevant, stimulus characteristics were utilized for phonemic processing. Additionally, cross-language differences in rhythmic structure between German and French influenced which nonword segments received more attention focus. These findings provide novel insights into critical determinants of phonemic processing in preschoolers and highlight the need for further research to explore these effects over time and within varying language backgrounds. Full article

Insect phonotaxis is influenced by various external and internal factors related to the environment and to the inner states of an organism. Previously, we found that flight, a natural behavior common among insects, significantly enhances the subsequent phonotactic response of female crickets (Gryllus bimaculatus) via serotonergic mechanisms. It is known that phonotaxis is also present in male crickets. However, the natural factors influencing phonotaxis in males and the neurochemical mechanisms behind these effects remained unknown. Our goal was to clarify these points and to compare phonotaxis of freely moving male and female crickets using a recently developed experimental paradigm. First, we found that freely moving male and female crickets demonstrate similar phonotactic response, quantitatively evaluated based on the time that animals spent near the source of calling song, their average distance to it and the number of approaches to it. We also found that the experience of previous flight increased phonotaxis in male crickets as it does in females. Also, as in females, this effect was associated with serotonergic neurotransmission. Thus, the manifestation of phonotactic behavior in freely moving male and female crickets, as well as the behavioral and monoaminergic modulation of phonotaxis in both sexes, appear to be quite similar in our experimental paradigm. Full article

Effective language processing relies on pattern detection. Spanish monolinguals predict verb tense through stress–suffix associations: a stressed first syllable signals present tense, while an unstressed first syllable signals past tense. Low-proficiency second language (L2) Spanish learners struggle to detect these associations, and we investigated whether they benefit from game-based training. We examined the effects of four variables on their ability to detect stress–suffix associations: three linguistic variables—verbs’ lexical stress (oxytones/paroxytones), first-syllable structure (consonant–vowel, CV/consonant–vowel–consonant, CVC), and phonotactic probability—and one learner variable—working memory (WM) span. Beginner English learners of Spanish played a digital game focused on stress–suffix associations for 10 days and completed a Spanish proficiency test (Lextale-Esp), a Spanish background and use questionnaire, and a Corsi WM task. The results revealed moderate gains in the acquisition of stress–suffix associations. Accuracy gains were observed for CV verbs and oxytones, and overall reaction times (RTs) decreased with gameplay. Higher-WM learners were more accurate and slower than lower-WM learners in all verb-type conditions. Our findings suggest that prosody influences word activation and that digital gaming can help learners attend to L2 inflectional morphology. Full article

Pseudowords are essential in (psycho)linguistic research, offering a way to study language without meaning interference. Various methods for creating pseudowords exist, but each has its limitations. Traditional approaches modify existing words, risking unintended recognition. Modern algorithmic methods use high-frequency n-grams or syllable deconstruction but often require specialized expertise. Currently, no automatic process for pseudoword generation is designed explicitly for Greek, which is our primary focus. Therefore, we developed SyBig-r-Morph, a novel application that constructs pseudowords using syllables as the main building block, replicating Greek phonotactic patterns. SyBig-r-Morph draws input from word lists and databases that include syllabification, word length, part of speech, and frequency information. It categorizes syllables by position to ensure phonotactic consistency with user-selected morphosyntactic categories and can optionally assign stress to generated words. Additionally, the tool uses multiple lexicons to eliminate phonologically invalid combinations. Its modular architecture allows easy adaptation to other languages. To further evaluate its output, we conducted a manual assessment using a tool that verifies phonotactic well-formedness based on phonological parameters derived from a corpus. Most SyBig-r-Morph words passed the stricter phonotactic criteria, confirming the tool’s sound design and linguistic adequacy. Full article

Arabic phonotactics significantly differ from English phonotactics in that they usually follow a framework that forbids the presence of consonant clusters in syllabic onsets. This study examines the relationship between Arabic-speaking EFL learners’ spelling accuracy and pronunciation, concentrating on the difficulties caused by consonant clusters in initial and final positions. A mixed-methods approach was employed, involving audio recordings to assess pronunciation accuracy and a spelling test to evaluate spelling performance. According to the results, words with initial consonant clusters (clusters at the beginning of a syllable, or onset) are harder to pronounce and spell than words with final consonant clusters (clusters that follow the vowel). Initial consonant clusters are considered to demand a quick transition in consonants at the start of a syllable, requiring more exact articulation. On the other hand, final clusters allow for a more progressive and straightforward articulation by involving a consonant transition following the initial vowel sound. Additionally, epenthesis may be used as a method to break up consonant clusters in L1 phonological interference, which is consistent with Arabic phonotactic patterns. Full article

Recent work found a correspondence between consonant clustering probability in monosyllabic lexemes and the three vowel types, short and long monophthong and diphthong, in German dialects. Furthermore, that correspondence was found to be bound to a North–South divide. This paper explores the preferences in consonant clustering of particular vowels by analyzing the PhonD2-Corpus, a large database of phonotactic and morphological information. The clustering probability of the diphthongs is positively correlated with frequency while the other vowels showed particular preferences that are not positively correlated with frequency. However, all of them are determined by a threefold pattern: short monophthongs prefer coda clusters, diphthongs onset clusters and long monophthong are balanced. Furthermore, it was found that this threefold pattern seems to have evolved from an originally twofold pattern (short monophthong prefers coda clusters and long monophthong and diphthong prefer onset clusters) in Middle High and Low German. This result is then further considered under the aspect of the compensation of the syllable weight and moraicity. Furthermore, some interesting parallels with the syllable vs. word-language typology framework are noted. Full article

We have recently shown that experience of flight remarkably enhanced subsequent terrestrial phonotaxis in females in response to the male calling song. Here, we elucidated the possible roles of octopamine and serotonin in the enhancing effect of flying on phonotactic behavior. Octopamine is known to be released into the hemolymph during flight in insects; however, the octopamine receptor antagonist epinastine did not abolish the effects of flight in our study. On the contrary, the drug significantly potentiated the influence of flying on phonotactic behavior. The octopamine receptor agonist chlordimeform, at a concentration of 2 mM, which was previously found to activate aggression in crickets, dramatically reduced the phonotactic response. However, at a 10-times-lower dose, chlordimeform produced a light but significant decrease in the time that females took to reach the source of the calling song. A similar effect was produced by octopamine itself, which hardly passes the blood–brain barrier in insects. The effect of flight was completely abolished in female crickets treated with alpha-methyl tryptophan (AMTP). AMPT suppresses the synthesis of serotonin, decreasing its content in the nervous systems of insects, including crickets. An activation of the serotonin synthesis with 5-hydroxytryptophan mimicked the effect of flight by increasing the number of visits to and the time spent in the zone near the source of the calling song. The 5-HT content in the third thoracic ganglion was significantly higher in flyers compared to the control group. In contrast, no changes in the octopamine level were observed in the third thoracic ganglion, which is known to play a crucial role in decision-making involved in intraspecific interactions. Therefore, the results suggest that although octopamine is known to be released into the hemolymph during flight, it is likely to inhibit rather than activate the central mechanisms related to phonotaxis. The weak facilitating effect of a low dose of chlordimeform can be attributed to the activation of peripheral octopaminergic receptors. Our results suggest that the serotoninergic system may contribute to the facilitation of female phonotactic behavior by flying. We suggest that both flying and serotonin enhance sexual motivation in females and, by these means, impact their behavioral response to the male calling song. Full article

The study investigated word recognition during neural activation in monolinguals and bilinguals. We specifically examined word retrieval and blood-oxygenation changes in the prefrontal cortex during a code-mixed word recognition task. Participants completed a gating task incorporating monolingual sentences and Spanish-English code-mixed sentences while using functional near-infrared spectroscopy (fNIRS) to measure blood-oxygenation changes. Word recognition contained four phonotactic conditions: (1) voiceless initial consonants, (2) voiced initial consonants, (3) CV-tense words, and (4) CV-lax words. Bilingual speakers had word-recognition capabilities similar to monolingual speakers even when identifying English words. Word recognition outcomes suggested that prefrontal cortex functioning is similar for early age of acquisition (AOA) bilinguals and monolinguals when identifying words in both code-mixed and monolingual sentences. Monolingual speakers experienced difficulty with English-voiced consonant sounds; while bilingual speakers experienced difficulties with English-lax vowels. Results suggest that localization of speech perception may be similar for both monolingual and bilingual populations, yet levels of activation differed. Our findings suggest that this parity is due to early age of acquisition (AoA) bilinguals finding a balance of language capabilities (i.e., native-like proficiency) and that in some instances the bilingual speakers processed language in the same areas dedicated to first language processing. Full article

Ideophones are believed to exhibit distinct phonotactic patterns compared to regular language, in their expressiveness. Vowel harmony can be observed in ideophones in Modern Korean. However, over time, Korean has gradually lost its regular vowel harmony process, due to the influx of foreign words, especially from Chinese, and historical sound changes like the vowel shift of /ɔ/ to different vowel types. Previous studies have mainly focused on the vowel patterns of ideophones without necessarily comparing the degree of vowel harmony between ideophones and other lexical strata. This lack of comparison makes it challenging to assess the level of corruption in vowel harmony specifically within ideophones, relative to other components of the lexicon. To address this gap, this paper examines vowel patterns extracted from the online dictionary of Korean, developed by the National Institute of the Korean Language (NIKL) with contributions from anonymous users and specialists. The analysis specifically explores vowel patterns across lexical items with varying syllable lengths, focusing on the lexical stratum, adverbial parts of speech, and the semantic meaning of the adverbials. This examination aims to assess the regularity of vowel sequencing and determine the extent of purity in vowel harmony patterns. The quantitative analysis of the compiled dictionary provides valuable insights into the degree of irregular phonotactics and its relationship to sound symbolism in Modern Korean. Full article

Mate choice is the attempt of an individual to gain higher reproductive fitness by preferring to mate with some individuals and not with others. We studied the role of mate choice in the mating system of the neotropical toad Rhinella arenarum by assessing male reproductive tactics for mate acquisition and the contribution of female choice for pair formation. In a shallow pond in central Argentina, we estimated male mating success and the corresponding reproductive tactics by focal observation. The variation of phenotypic and genotypic traits (size and shape, longevity, vocalization features, heterozygosity) was related to the observed mating success in 110 males. The phonotactic response of 21 reproductive females to conspecific advertisement call features was tested in arena experiments. Mating success was limited to 32 males, pair formation was size-assortative. The dominant reproductive tactics were advertising from call positions near suitable breeding sites and pre-mating fights with intruding males, whereas the interception of amplectant pairs and the displacement of mated males were never observed. Female phonotaxis was directed to conspecific choruses but complex and simple call structures were not distinguished. We conclude that the mating system is a lek combining pre-mating fights among males and female choice of slightly smaller males. Fights interfere with female choice, undermining size-assortative mating. This is a unique system in the R. marina species group, in which interception behavior dominates reproduction. Full article

This paper focuses on prosodic adjunction at the Prosodic Word level in a polysynthetic language. I argue that recursion at a depth of more than two levels can only be generated by a theory which requires exact correspondence between certain syntactic phrases and Prosodic Words. Such a theory is similar to Phonological Phrase correspondence in Match Theory, suggesting there is an underlying shared property between correspondence at the Prosodic Word and Phonological Phrase levels. In addition, this theory must include a constraint which prohibits recursive prosodic constituents in order to generate the attested typology of clitics across languages. The empirical focus is the prosodic structure of the verbal complex in Blackfoot (Algonquian; ISO 639-3: bla). Using phonotactic evidence I argue that the vP phase corresponds to a Prosodic Word, and that each prefix to the stem is a Prosodic Word adjunct. I then compare several theories of the syntax-prosody interface, including versions of Alignment Theory, Wrap Theory, and Match Theory. A subset of schematic candidates with one or two prefixes to a stem are used to determine which theories generate the attested typology of clitics as well as a multiply recursive Prosodic Word structure. Full article

Functional load (FL) quantifies the contributions by phonological contrasts to distinctions made across the lexicon. Previous research has linked particularly low values of FL to sound change. Here, we broaden the scope of enquiry into FL to its evolution at higher values also. We apply phylogenetic methods to examine the diachronic evolution of FL across 90 languages of the Pama–Nyungan (PN) family of Australia. We find a high degree of phylogenetic signal in FL, indicating that FL values covary closely with genealogical structure across the family. Though phylogenetic signals have been reported for phonological structures, such as phonotactics, their detection in measures of phonological function is novel. We also find a significant, negative correlation between the FL of vowel length and of the following consonant—that is, a time-depth historical trade-off dynamic, which we relate to known allophony in modern PN languages and compensatory sound changes in their past. The findings reveal a historical dynamic, similar to transphonologization, which we characterize as a flow of contrastiveness between subsystems of the phonology. Recurring across a language family that spans a whole continent and many millennia of time depth, our findings provide one of the most compelling examples yet of Sapir’s ‘drift’ hypothesis of non-accidental parallel development in historically related languages. Full article

In Linguistics and Psycholinguistics, phonotactics refers to the constraints on individual sounds in a given language that restrict how those sounds can be ordered to form words in that language. Previous empirical work in Psycholinguistics demonstrated that phonotactic knowledge influenced how quickly and accurately listeners retrieved words from that part of memory known as the mental lexicon. In the present study, we used three computer simulations to explore how three different cognitive network architectures could account for the previously observed effects of phonotactics on processing. The results of Simulation 1 showed that some—but not all—effects of phonotactics could be accounted for in a network where nodes represent words and edges connect words that are phonologically related to each other. In Simulation 2, a different network architecture was used to again account for some—but not all—effects of phonotactics and phonological neighborhood density. A bipartite network was used in Simulation 3 to account for many of the previously observed effects of phonotactic knowledge on spoken word recognition. The value of using computer simulations to explore different network architectures is discussed. Full article