Abstract
We first investigate the dynamical behavior of an active Brownian particle influenced by a viscoelastic memory effect characterized by a power-law kernel, under the effects of thermal and active noises. We then analyze the dynamics of an active Brownian particle confined in a harmonic trap in the presence of the same noise sources. To derive the Fokker–Planck equation for the joint probability density of the active particle, we obtain analytical solutions for the joint probability density and its moments using double Fourier transforms in the limits , , and . As a result, the mean squared displacement of an active Brownian particle driven by thermal noise exhibits a super-diffusive scaling of in the short-time regime (). In contrast, for a particle in a harmonic trap driven by active noise, the mean squared velocity scales linearly with when . Moreover, the higher-order moments of an active Brownian particle in a harmonic trap with thermal noise scale with in the long-time limit () and for , consistent with our analytical results.
1. Introduction
Recently, the nonequilibrium motion of active particles has been extensively studied in various models, including the run-and-tumble model [1,2], active Brownian particles [3,4], active Langevin particles [5], active Ornstein–Uhlenbeck particles [6,7,8], self-propelled Janus colloids [9,10], and biological microswimmers [11]. Active diffusion, investigated through theoretical analyses, computer simulations, and experiments on active viscoelastic systems, has revealed both similarities and differences among these systems. Representative examples include the transport of passive tracers in active baths and living cells [12], chromosomal dynamics [13], lateral diffusion of membrane proteins [14], and tracer diffusion in dense colloidal suspensions [15]. Active particles have also been explored in polymeric environments such as actomyosin and endoplasmic reticulum networks [16], microtubule assemblies, and macromolecules bound to polymer strands, including DNA and chromosomes.
Over the past two decades, anomalous diffusion dynamics [17,18] have been widely discussed and extended in natural and complex scientific systems. In the typical form of anomalous diffusion, characterized by a scaling exponent , the system exhibits sub-diffusive behavior for , indicated by a sub-linear growth of the mean squared displacement over time, and super-diffusive behavior for , characterized by super-linear growth of the mean squared displacement. Sub-diffusion has been observed for endogenous submicron tracers in biological systems, such as living cells [19], artificially crowded biological environments [20], protein motion in supercomputing simulations [21], and dilute or protein-crowded lipid bilayer membranes. Super-diffusion has been reported in several cellular systems [22]. For instance, a restoring force applied by an optical tweezer in a biological cell [23,24] enabled the fractional Ornstein–Uhlenbeck process [25]. Active processes in the underdamped limit exhibit ballistic motion, while complex underdamped dynamics can lead to hyper-diffusion in the regime [26,27].
Furthermore, theories based on the active fractional Langevin equation have been developed to quantitatively describe transport phenomena in various active viscoelastic systems. These studies also introduced weak ergodicity breaking, a novel phenomenon not previously reported in other systems [28,29]. For Gaussian processes, all statistical properties can be inferred from the mean and covariance functions [30,31,32]. Indirect approaches to assessing the ergodic properties of Gaussian processes have been compared with the behaviors of the mean squared displacement and the time-averaged MSD [33,34].
Fractional Brownian motion is a type of Gaussian process characterized by a two-time autocovariance function [35], with the mean squared displacement given by for . The corresponding probability density function is . For , increments are positively correlated, leading to super-diffusive motion, whereas for , increments are negatively correlated, resulting in sub-diffusive behavior. Standard Brownian motion with independent increments corresponds to normal diffusion for . Fractional Gaussian noise is defined as where is a small, finite time interval. The autocovariance function for an active particle driven by fractional Gaussian noise is then which asymptotically satisfies for . The variance is given by , and in the ballistic limit (), the autocovariance reduces to .
Until now, most of these problems have been addressed numerically using perturbation methods, special functions, or approximations. This study builds on the classical formulations of Heinrichs [30], Athanassoulis et al. [31], and Mamis and Farazmand [32], but extends the framework in several significant ways. By considering thermal noise, active noise, viscoelastic memory effects, and optical trapping forces, we provide novel analytical and numerical comparisons that reveal new scaling relationships, correlation coefficients, entropy connections, and stability properties not covered in previous studies.
Recently, it has become more straightforward to describe the motion of active particles using equations of motion when they experience varying forces within a viscous medium. However, obtaining analytical solutions for the probability densities of displacement, velocity, orientation, and other dynamical quantities remains challenging. Previous studies have investigated the fractional generalized Langevin equation for a passive particle subjected to thermal equilibrium noise and active noise with exponentially decaying correlations. In these viscoelastic systems, the particle experiences two distinct types of noise: thermal noise, which satisfies the fluctuation–dissipation theorem, and active noise, modeled as an active Ornstein–Uhlenbeck process. In this work, we derive the Fokker–Planck equation for the joint probability density and obtain its solution using double Fourier transforms in three distinct time regimes. The organization of this paper is as follows. In Section 2, we derive the Fokker–Planck equation from the fractional generalized Langevin equation. In Section 3, using double Fourier transforms, we obtain approximate solutions for the joint probability density of an active Brownian particle subject to harmonic and viscous forces in three regimes of correlation times and . Section 4 presents numerical calculations of the non-Gaussian parameter, correlation coefficients, and entropy. Finally, in Section 5, we summarize the key findings and provide concluding remarks.
2. Thermal and Active Fractional Generalized Langevin Equations
As a nonequilibrium dynamic model, our first model derives the Fokker–Planck equation for the two variables of displacement and velocity by introducing the viscoelastic memory effect, as well as thermal and active noises, in the fractional Langevin equations, thereby obtaining the joint probability density. The second model introduces the fractional Langevin equations that account for the viscoelastic memory effect, an optical trapped force, and thermal and active noises to obtain a new joint probability density for displacement and velocity.
In this section, we introduce a class of nonequilibrium dynamic models referred to as the fractional Langevin equation which incorporates a viscoelastic memory effect [26] with a power-law kernel .
The active fractional Langevin equation, a class of nonequilibrium dynamic models, is presented; its unique viscoelastic memory effect is characterized by a power-law decay of correlations over time. We derive the Fokker–Planck equation for the joint probability density from the fractional generalized Langevin equation with thermal equilibrium noise. The fractional generalized Langevin equations in our model are expressed as
where and denote the thermal equilibrium noise and the active noise (with exponentially decaying correlation), respectively. We introduce these two types of noise as
where the persistent Hurst exponent h ranges as and the correlation times are and . The thermal equilibrium noise is a fractional Gaussian noise coupled to the memory kernel via the fluctuation–dissipation theorem. The joint probability density for the displacement and the velocity is defined by
for . By taking time derivatives in the joint probability density and inserting Equations (1) and (2) into Equation (5), we can write the time derivative of the joint probability equation for and as
where , , , and . The memory effect is expressed as for . Assuming that the particle is initially at rest at , the forms of the joint probability density [30,31,32] can be written as
where . The parameters are
It is apparent that Equations (8) and (9) are the Fokker–Planck equations, as mentioned in the Introduction. Later, the time evolutions of the joint probability densities and , Equations (8) and (9), will be derived in Appendix A.
Some derivations related to the correlated Gaussian force are given in Ref. [30]. We define the double Fourier transform of the joint probability density as
The Fourier transforms of the Fokker–Planck equation, Equations (8) and (9), are expressed as
2.1. and with Thermal Noise
2.1.1. and with Thermal Noise in
In this subsection, we derive the solutions of the probability densities and in the short-time domain . To find the special solutions for and by the variable separation from Equation (12), the two equations for the displacement and the velocity are written as
where denotes the separation constant. Taking in the steady state, we get as
To find the solution of the probability density for from , we include terms up to order and write
Assuming arbitrary functions of variable , the probability density becomes . Therefore, we have
By a similar method, from Equation (16) to Equation (19), for , we also get the Fourier transform of the probability density for as
By calculating Equations (19) and (20), the Fourier transform of the joint probability density is
By taking the inverse Fourier transform, we obtain
The mean squared displacement for and the mean squared displacement for are
2.1.2. and with Thermal Noise in
We now consider the long-time domain . From Equation (14), an approximate equation for the Fourier-transformed probability density is
The Fourier transform of the probability density from Equation (25) is calculated as
The steady probability density for , from , is
As the Fourier transform of probability density in the short-time domain is given by , is derived as
where . Applying Equation (15) for to a similar method from Equation (25) to Equation (28) of derived, we also get the Fourier transforms of the probability density for velocity as
By calculating Equations (28) and (29), we have
By using the inverse Fourier transform, the probability densities and are, respectively, presented by
Finally, the mean squared values and for the probability densities and are, respectively, given by
2.1.3. and with Thermal Noise for
In this subsection, we find the probability densities and in the time domain . The approximate equation from Equation (14) for is written as
In the steady state, we can calculate from Equation (34) as
We find the Fourier transform of the probability density as
Using a similar procedure of , we get the Fourier transform of the probability density as
We can calculate by calculating Equations (36) and (37) as
Using the inverse Fourier transform, the probability densities and are, respectively, presented by
Thus, the mean squared displacement and the mean squared velocity are
2.2. and with Active Noise
2.2.1. and with Active Noise in the Short-Time Domain
In this subsection, we settle the solutions of the probability densities and in the short-time domain . In order to find the special solutions for , with the variable separation from Equation (13), the two equations for the displacement and the velocity are given by
where denotes the separation constant. As we take in the steady state, we get as
To get the solution of the probability density for from , we calculate the Fourier transform of the probability density after including terms up to order as
Taking the solutions as arbitrary functions of variable , the arbitrary function becomes . As a result, we find that
Using a similar method from Equation (44) to Equation (49) for , we also get the Fourier transform of the probability density for the velocity as
Therefore, by calculating Equations (49) and (50), we get the Fourier transform of the joint probability density as
Using the inverse Fourier transform, we get
The corresponding mean squared displacement and the mean squared displacement for and are, respectively, given by
2.2.2. and with Active Noise in the Long-Time Domain
We find the probability densities and in the long-time domain . We write an approximate equation from Equation (42) as
The Fourier transform of the probability density from Equation (55) is calculated as
We find that the steady probability density for , from , is derived as
As the Fourier transform of the probability density in the short-time domain is given by , is derived as
where and is equal to Equation (44). Applying Equation (43) for to a similar method from Equation (55) to Equation (58) of derived, we also get the Fourier transforms of the probability density for velocity as
By calculating Equations (58) and (59), we have
Using the inverse Fourier transform, the probability densities and are, respectively, presented by
The mean squared values and for the probability densities and are
2.2.3. and with Active Noise for
In this subsection, we find and for the time domain . The approximate equation from Equation (42) for is written as
In the steady state, we can calculate from Equation (64) as
where , for . We find the Fourier transform of the probability density as
Using a similar method to that of derived, we get the Fourier transform of the probability density as
We can calculate by calculating Equations (66) and (67) as
Using the inverse Fourier transform, the probability densities and are, respectively, presented by
The mean squared displacement and the mean squared velocity are, respectively, given by
3. Thermal and Active Fractional Fokker–Planck Equations
We consider a nonequilibrium dynamic model referred to as the active fractional Langevin equation, having a viscoelastic memory effect with a power-law kernel [25], with an optical trapped force and thermal and active noises. The two altered fractional generalized Langevin equations in our model are expressed in terms of
where thermal equilibrium noise and active noise denote and , respectively. We introduce two noises, i.e., thermal noise and active noise, as
where for . The thermal energy is and the correlation times are and . Thermal equilibrium noise is fractional Gaussian noise coupled to the memory kernel via the fluctuation dissipation theorem, while active noise is responsible for both the self-propelled motion of an active particle and nonequilibrium motion from an active heat reservoir.
We next derive the Fokker–Planck equation from the active fractional generalized Langevin equation. First, the joint probability density for displacement and velocity is defined by
where . Taking time derivatives in the joint probability density and inserting Equations (71) and (72) into Equation (75), we can write the time derivative of the joint probability equation for as follows:
where , , and the persistent Hurst exponent h ranges to . In Equations (76) and (77), we have . We assume from the joint probability density that the particle is initially at rest at time . Then, the joint probability densities [34,35,36] satisfy the Fokker–Planck equations derived as
where . The parameters are
Equations (78) and (79) are called the Fokker–Planck equations, as mentioned in the Introduction. Some derivations in relation to the correlated Gaussian force are given in Ref. [30]. We define the double Fourier transform of the joint probability density by the equation
where . The Fourier transforms of the Fokker–Planck equations, Equations (78) and (79), are, respectively, expressed in terms of
3.1. and with Thermal Noise
3.1.1. and with Thermal Noise in the Short-Time Domain
In this subsection, we obtain the solutions of the probability densities and in the short-time domain . To find the special solutions for by the variable separation from Equation (82), the two equations for the displacement and the velocity are given by
where denotes the separation constant.
In the steady state, assuming , we obtain
We assume that . To obtain the probability density for from , we calculate the Fourier transform of the probability density after including terms up to order :
Considering the solutions as arbitrary functions of variable , the arbitrary function becomes . Thus,
Using a similar method from Equation (86) to Equation (89) for , we also get the Fourier transform of the probability density for the velocity as
Hence, by calculating Equations (89) and (90), we get the Fourier transform of the joint probability density as
Using the inverse Fourier transform, we get
The mean squared displacement for and the mean squared displacement for are
3.1.2. and with Thermal Noise in the Long-Time Domain
Now, we derive the probability densities and in the long-time domain (). An approximate equation from Equation (84) can be written as
The Fourier transform of the probability density from Equation (95) is obtained as
We find the Fourier transform of the steady probability density for , defined by , as
Since the Fourier transform of probability density in the short-time domain is given by , we can write as
where . Applying Equation (85) for in the same manner as in Equations (95)–(98) of derived, we also obtain the Fourier transforms of the probability density for velocity as
By calculating Equations (98) and (99), we have
By performing the inverse Fourier transform, the probability densities and are, respectively, calculated as
The mean squared values and for the probability densities and are, respectively,
3.1.3. and with Thermal Noise for
In this subsection, we derive and for the case . The approximate equation from Equation (84) for is written as
In the steady state, can be obtained from Equation (104) as
The Fourier transform of the probability density is then given by
Using a similar procedure to that for , the Fourier transform of the probability density becomes
We can get by calculating Equations (106) and (107) as
Using the inverse Fourier transform, the probability densities and are, respectively, given by
The mean squared displacement and the mean squared velocity are
3.2. and with Active Noise
3.2.1. and with Active Noise in the Short-Time Domain
In this subsection, we obtain the solutions of the probability densities and in the short-time domain . To find the particular solutions for and by separating variables from Equation (83), the two equations for displacement and velocity are given by
where E denotes the separation constant. By setting in the steady state, we get as
To find the solution of the probability density for from , we calculate the Fourier transform of the probability density after including terms up to order as
Taking the solutions as arbitrary functions of variable , the arbitrary function becomes . As a result, we find that
Using a similar method as Equations (114)–(119), we also get the Fourier transform of the probability density for the velocity as
Therefore, by calculating Equations (119) and (120), we get the Fourier transform of the joint probability density as
Using the inverse Fourier transform, we get
The mean squared displacement and the mean squared displacement for and are thus given by
3.2.2. and with Active Noise in the Long-Time Domain
We now find the probability densities and in the long-time domain . An approximate equation from Equation (112) can be written as
The Fourier transform of the probability density from Equation (125) is
We find the steady-state probability density for from ,
As the Fourier transform of the probability density in the short-time domain is given by , is derived as
where and is equal to Equation (114). Applying Equation (113) for to a similar method from Equation (125) to Equation (128) for derived, we also get the Fourier transforms of the probability density for velocity as
By calculating Equations (128) and (129), we have
Using the inverse Fourier transform, the probability densities and are, respectively, presented by
Consequently, the mean squared values and for the probability densities and are
3.2.3. and with Active Noise for
In this subsection, we derive and in the time domain . The approximate equation from Equation (112) for is written as
In the steady state, we can calculate from Equation (134) as
We find the Fourier transform of the probability density as
Using a similar procedure to that for , we get the Fourier transform of the probability density as
We can calculate by calculating Equations (136) and (137) as
Using the inverse Fourier transform, the probability densities and are, respectively, presented by
The mean squared displacement and the mean squared velocity are, respectively, given by
4. Statistical Quantities
In this section, we calculate statistical quantities, including the non-Gaussian parameter for displacement and velocity, the correlation coefficient, the entropy, the combined entropy, and the moments from the moment equations. The moment equations for an active particle with a viscoelastic memory effect, derived from Equations (8) and (9), are expressed as
where . For an active particle in an optical trap, the moment equations from Equations (78) and (79) are
Next, the entropy, the non-Gaussian parameter, and the correlation coefficient are calculated numerically. The entropies and are calculated as
The combined entropy is defined by
Entropy provides a numerical measure of the uncertainty or information content associated with a probability distribution; as the probability of a particular value increases and that of other values decreases, the entropy decreases.
The non-Gaussian parameters for displacement and velocity are, respectively, given by
The non-Gaussian parameter quantifies the degree of heavy tails in the probability distribution of a real-valued variable, providing insight into the deviation from Gaussian behavior. The correlation coefficient is defined as
The correlation coefficient describes the strength and direction of the linear relationship between two variables, and . Here, we assume that an active particle is initially at and at for . The parameters and denote the root-mean-squared displacement and the root-mean-squared velocity of the joint probability density, respectively.
Table 1 and Table 2, respectively, summarize the values of the entropy, non-Gaussian parameter, and correlation coefficient for the joint probability density with thermal equilibrium noise and active noise in the three-time domains. Table 3 and Table 4 show the non-Gaussian parameter, the correlation coefficient, the moment, the entropy, and the combined entropy for the joint probability density for an active Brownian particle with a harmonic trap, , and in the three-time domains.
Table 1.
Values of the non-Gaussian parameter, the correlation coefficient, the moment, the entropy, and the combined entropy for the joint probability density with in the three-time domains.
Table 2.
Values of the non-Gaussian parameter, the correlation coefficient, the moment, the entropy, and the combined entropy for the joint probability density with in the three-time domains.
Table 3.
Values of the non-Gaussian parameter, the correlation coefficient, the moment, the entropy, and the combined entropy for the joint probability density with optical trap and in the three-time domains.
Table 4.
Values of the non-Gaussian parameter, the correlation coefficient, the moment, the entropy, and the combined entropy for the joint probability density with an optical trap and in the three-time domains.
5. Conclusions
In conclusion, we studied the nonequilibrium effects of an active Brownian particle in a viscoelastic medium, subjected to thermal noise and active noise , using the fractional generalized Langevin equation. Thermal noise satisfies the fluctuation–dissipation theorem, whereas active noise is modeled as an active Ornstein–Uhlenbeck process. We derived the Fokker–Planck equation for the joint probability density and obtained its solution via double Fourier transforms across three-time regimes.
The main work and conclusions are as follows:
- (1)
- In the fractional generalized Langevin equation with a viscoelastic memory effect and thermal noise for (where is the Hurst exponent), the mean squared displacement scales with in the limits of and and for , exhibiting super-diffusive behavior, while the mean squared velocity scales with . In particular, for , the mean squared displacement scales with in the limit of , while the mean squared velocity behaves as , consistent with simulated results [33,34]. The mean squared displacement for the fractional generalized Langevin equation scales with in the short-time limit , consistent with the simulation result reported in Ref. [26].
- (2)
- As we solve the fractional Fokker–Planck equation with a harmonic force and thermal noise, the mean squared displacement behaves as in and . The mean squared velocity for exhibits normal diffusion from the fractional generalized Langevin equation with a harmonic force and active noise.
- (3)
- From statistical quantities, the entropy for the joint probability density with as is the same value as that for the joint probability density with in and .
- (4)
- The combined entropy for the joint probability density with is larger than that for the joint probability density with in each time domain. In Table 3, the combined entropy of displacement and velocity follows a scaling behavior of in the time regime , where the scaling exponent lies in the range . Since the combined entropy maintains functional dependence, , in the other regimes ( and ), comparison among these regimes indicates that the system in the region reaches equilibrium more rapidly. This result suggests that the long-time dynamics are dominated by a faster relaxation toward equilibrium despite the identical entropy scaling form.
- (5)
- For , the displacement non-Gaussian parameter for the joint probability density with is the same value proportional to time as that for the joint probability density with in . The velocity non-Gaussian parameter for the joint probability density with is also the same value proportional to time as that for the joint probability density with for in . In Table 1, Table 2, Table 3 and Table 4, as the non-Gaussian parameter increases, the distribution exhibits heavier tails and stronger deviations from Gaussian behavior. Such behavior typically appears in the time regime for , and , . In contrast, for and at , the non-Gaussian parameters of , and , correspond to flatter central regions and shorter tails, indicating light-tailed distributions.
- (6)
- Moments for an active Brownian particle in a harmonic trap with thermal noise in the limits of and for scale with , consistent with our calculation result.
The formal approach of our study shares similarities with analytical frameworks, but extends them in several important aspects. We particularly addressed the roles of active noise, the non-Gaussian parameter, and entropy aspects that have not been sufficiently analyzed in previous studies. Moreover, we hope that the present results provide new physical insights into the mechanisms of transition kinetics, fractional diffusion, and the suppression of rare events. Future work combining theoretical [36,37,38,39], computational [40,41,42,43], and experimental approaches can clarify how inertial effects, finite correlation times, and non-Gaussian statistics influence transport and stability in active matter. The methodology developed here offers a versatile framework applicable to generalized stochastic systems, anomalous diffusion, and self-propelled particle dynamics in complex environments, such as shear flows [44,45]. Interdisciplinary extensions of this work are expected to deepen our understanding of noise-driven processes in fractional active and nonequilibrium systems [46,47,48,49].
Author Contributions
K.K.: Writing, conceptualization, review, editing, and methodology. Y.J.K.: review, editing, investigation, table analysis, and funding acquisition. S.K.S. and S.K.: Review, editing, investigation, and table analysis. All authors have read and agreed to the published version of the manuscript.
Funding
Y.J.K. was funded; This research was funded by Wonkwang University in 2024.
Data Availability Statement
Data will be made available on request.
Conflicts of Interest
Author Sung Kyu Seo was employed by the company Haena Ltd., Author Kyungsik Kim was employed by the company DigiQuay Ltd. The remaining authors declare that the research was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest.
References
- Patteson, A.E.; Gopinath, A.; Goulian, M.; Arratia, P.E. Running and tumbling with E. coli in polymeric solutions. Sci. Rep. 2015, 5, 15761. [Google Scholar] [CrossRef] [Scilit]
- Bechinger, C.; Di Leonardo, R.; Löwen, H.; Reichhardt, C.; Volpe, G.; Volpe, G. Active particles in complex and crowded environments. Rev. Mod. Phys. 2016, 88, 045006. [Google Scholar] [CrossRef] [Scilit]
- Pinto-Goldberg, M.; Soto, R. Hydrodynamic Equations for Active Brownian Particles in the High Persistence Regime. arXiv 2025, arXiv:2506.17509. [Google Scholar] [CrossRef] [Scilit]
- Romanczuk, P.; Bar, M.; Ebeling, W.; Lindner, B.; Schimansky-Geier, L. Active Brownian particles: From individual to collective stochastic dynamics. Eur. Phys. J. Spec. Top. 2012, 202, 1. [Google Scholar] [CrossRef] [Scilit]
- Shaebani, M.R.; Wysocki, A.; Winkler, R.G.; Gompper, G.; Rieger, H. Computational models for active matter. Nat. Rev. Phys. 2020, 2, 181. [Google Scholar] [CrossRef] [Scilit]
- Fodor, E.; Nardini, C.; Cates, M.E.; Tailleur, J.; Visco, P.; van Wijland, F. How far from equilibrium is active matter. Phys. Rev. Lett. 2016, 117, 038103. [Google Scholar] [CrossRef] [Scilit]
- Bonilla, L.L. Active Ornstein-Uhlenbeck particles. Phys. Rev. E 2019, 100, 022601. [Google Scholar] [CrossRef] [Scilit]
- David Martin, D.; O’Byrne, J.; Cates, M.E.; Fodor, E.; Nardini, C.; Tailleur, J.; van Wijland, F. Statistical mechanics of active Ornstein-Uhlenbeck particles. Phys. Rev. E 2021, 103, 032607. [Google Scholar] [CrossRef] [Scilit]
- Nishiguchi, D. Deciphering Long-Range Order in Active Matter: Insights from Swimming Bacteria in Quasi-2D and Electrokinetic Janus Particles. J. Phys. Soc. Jpn. 2023, 92, 121007. [Google Scholar] [CrossRef] [Scilit]
- Rao, D.V.; Reddy, N.; Fransaer, J.; Clasen, C. Self-propulsion of bent bimetallic Janus rods. J. Phys. D 2018, 52, 014002. [Google Scholar]
- Yasuda, K.; Hosaka, Y.; Komura, S. Generalized Three-Sphere Microswimmers. J. Phys. Soc. Jpn. 2023, 92, 121008. [Google Scholar] [CrossRef] [Scilit]
- Gal, N.; Weihs, D. Experimental evidence of strong anomalous diffusion in living cells. Phys. Rev. E 2010, 81, 020903. [Google Scholar] [CrossRef] [Scilit]
- Bronstein, I.; Israel, Y.; Kepten, E.; Mai, S.; Shav-Tal, Y.; Barkai, E.; Garini, Y. Transient Anomalous Diffusion of Telomeres in the Nucleus of Mammalian Cells. Phys. Rev. Lett. 2009, 103, 018102. [Google Scholar] [CrossRef] [Scilit]
- Jeon, J.-H.; Tejedor, V.; Burov, S.; Barkai, E.; Selhuber-Unkel, C.; Berg-Sørensen, K.; Oddershede, L.; Metzler, R. In Vivo Anomalous Diffusion and Weak Ergodicity Breaking of Lipid Granules. Phys. Rev. Lett. 2011, 106, 048103. [Google Scholar] [CrossRef] [Scilit]
- Narinder, N.; Gomez-Solano, J.R.; Bechinger, C. Active particles in geometrically confined viscoelastic fluids. New J. Phys. 2019, 21, 093058. [Google Scholar] [CrossRef] [Scilit]
- Lin, C.; Zhang, Y.; Sparkes, I.; Ashwin, P. Sructure and Dynamics of ER: Minimal Networks and Biophysical Constraints. Biophys. J. 2014, 107, 763. [Google Scholar] [CrossRef] [Scilit]
- Klages, R.; Radons, G.; Sokolov, I.M. Anomalous Transport; Wiley-VCH Verlag: Berlin, Germany, 2008. [Google Scholar]
- Jeon, J.-H.; Monne, H.M.S.; Javanainen, M.; Metzler, R. Lateral motion of phospholipids and cholesterols in a lipid bilayer: Anomalous diffusion and its origins. Phys. Rev. Lett. 2012, 109, 188103. [Google Scholar] [CrossRef] [Scilit]
- Newman, T.A.C.; Beltran, B.; McGehee, J.M.; Elnatan, D.; Cahoon, C.K.; Paddy, M.R.; Chu, D.B.; Spakowitz, A.J.; Burgess, S.M. Diffusion and distal linkages govern interchromosomal dynamics during meiotic prophase. Proc. Natl. Acad. Sci. USA 2022, 119, e2115883119. [Google Scholar] [CrossRef] [Scilit]
- Jeon, J.-H.; Leijnse, N.; Oddershede, L.B.; Metzler, R. Anomalous diffusion and power-law relaxation in wormlike micellar solution. New J. Phys. 2013, 15, 045011. [Google Scholar] [CrossRef] [Scilit]
- Lizana, L.; Ambjornsson, T.; Lomholt, M.A. Tracer particles in two-dimensional elastic networks diffuse logarithmically slow. J. Phys. A 2016, 50, 034001. [Google Scholar] [CrossRef] [Scilit]
- Song, M.S.; Moon, H.C.; Jeon, J.-H.; Park, H.Y. Neuronal messenger ribonucleoprotein transport follows an aging Lévy walk. Nat. Comm. 2018, 9, 344. [Google Scholar] [CrossRef] [Scilit]
- Reverey, J.F.; Jeon, J.-H.; Leippe, M.; Metzler, R.; Selhuber-Unkel, C. Superdiffusion dominates intracellular particle motion in the supercrowded space of pathogenic Acanthamoeba castellanii. Sci. Rep. 2015, 5, 11690. [Google Scholar] [CrossRef] [Scilit]
- Meroz, Y.; Sokolov, I.M. A toolbox for determining subdiffusive mechanisms. Phys. Rep. 2015, 573, 1. [Google Scholar] [CrossRef] [Scilit]
- Cheridito, P.; Kawaguchi, H.; Maejima, M. Fractional Ornstein–Uhlenbeck Processes. Electron. J. Probab. 2003, 8, 1. [Google Scholar] [CrossRef] [Scilit]
- Joo, S.; Jeon, J.H. Viscoelastic active diffusion governed by nonequilibrium fractional Langevin equations: Underdamped dynamics and ergodicity breaking. Chaos Solitons Fractals 2023, 177, 114288. [Google Scholar] [CrossRef] [Scilit]
- Hida, T.; Hitsuda, M. Gaussian Processes. Metrika 1997, 46, 85. [Google Scholar]
- Deng, W.; Barkai, E. Ergodic properties of fractional Brownian–Langevin motion. Phys. Rev. E 2009, 79, 011112. [Google Scholar]
- Slezak, J.; Metzler, R.; Magdziarz, M. Codifference can detect ergodicity breaking and non-Gaussianity. New J. Phys. 2019, 21, 053008. [Google Scholar] [CrossRef] [Scilit]
- Heinrichs, J. Probability distributions for second-order processes driven by Gaussian noise. Phys. Rev. E 1993, 47, 3007. [Google Scholar] [CrossRef] [Scilit]
- Athanassoulis, G.A.; Nikolaos, P.; Nikoletatos-Kekatos, N.P.; Mamis, K.I. A systematic path to non-Markovian dynamics II: Probabilistic response of nonlinear multidimensional systems to Gaussian colored noise excitation. arXiv 2024, arXiv:2405.10236. [Google Scholar] [CrossRef] [Scilit]
- Mamis, K.I.; Farazmand, M. Mitigation of rare events in multistable systems driven by correlated noise. Phys. Rev. E 2021, 104, 034201. [Google Scholar] [CrossRef] [Scilit]
- Volpe, G.; Volpe, G. Simulation of a Brownian particle in an optical trap. Am. J. Phys. 2013, 81, 224. [Google Scholar] [CrossRef] [Scilit]
- Volpe, G.; Volpe, G.; Petrov, D. Brownian motion in a nonhomogeneous force field and photonic force microscope. Phys. Rev. E 2007, 76, 061118. [Google Scholar] [CrossRef] [Scilit]
- Nguyen, G.H.P.; Wittmann, R.; Löwen, H. Active Ornstein-Uhlenbeck model for self-propelled particles with inertia. J. Phys. Condens. Matter 2021, 34, 035101. [Google Scholar] [CrossRef] [Scilit]
- Jung, J.W.; Seo, S.K.; Kim, K. Joint probability densities of an active particle coupled to two heat reservoirs. Physical A 2025, 668, 130483. [Google Scholar] [CrossRef] [Scilit]
- Jung, J.W.; Seo, S.K.; Kim, K. Dynamical behavior of passive particles with harmonic, viscous, and correlated Gaussian forces. Phys. Lett. A 2025, 546, 130512. [Google Scholar] [CrossRef] [Scilit]
- Kang, Y.J.; Jung, J.W.; Seo, S.K.; Kim, K. On the Stochastic Motion Induced by Magnetic Fields in Random Environments. Entropy 2025, 27, 330. [Google Scholar] [CrossRef] [Scilit]
- Caprini, L.; Marini Bettolo Marconi, U. Inertial self-propelled particles. J. Chem. Phys. 2021, 154, 024902. [Google Scholar] [CrossRef] [Scilit]
- Sandoval, M. Pressure and diffusion of active matter with inertia. Phys. Rev. E 2020, 101, 012606. [Google Scholar] [CrossRef] [Scilit]
- Li, Q.; Zheng, X.; Wang, H.; Yang, Z.; Guo, X. An Analysis and Global Identification of Smoothless Variable Order of a Fractional Stochastic Differential Equation. Fractal Fract. 2023, 7, 850. [Google Scholar] [CrossRef] [Scilit]
- Alberti, T.; Carbone, V. Lagrangian Particle Dispersion in a Poor Man’s Magnetohydrodynamic Turbulence Model. Fractal Fract. 2023, 7, 662. [Google Scholar] [CrossRef] [Scilit]
- Taloni, A. Diffusion of an Active Particle Bound to a Generalized Elastic Model: Fractional Langevin Equation. Fractal Fract. 2024, 8, 76. [Google Scholar] [CrossRef] [Scilit]
- Muhuri, S.; Rao, M.; Ramaswamy, S. Shear-flow–induced isotropic-to-nematic transition in a suspension of active filaments. Europhys. Lett. 2007, 78, 48002. [Google Scholar] [CrossRef] [Scilit]
- Hagen, B.; Wittkowski, R.; Löwen, H. Brownian dynamics of a self-propelled particle in shear flow. Phys. Rev. E 2011, 84, 031105. [Google Scholar] [CrossRef] [Scilit]
- Lutz, E. Fractional Langevin equation. Phys. Rev. E 2001, 64, 051106. [Google Scholar] [CrossRef] [Scilit]
- Wei, Q.; Wang, W.; Tang, Y.; Metzler, R.; Chechkin, A. Fractional Langevin equation far from equilibrium: Riemann-Liouville fractional Brownian motion, spurious nonergodicity, and aging. Phys. Rev. E 2025, 111, 014128. [Google Scholar] [CrossRef] [Scilit]
- Jolakoski, P.; Basnarkov, L.; Kocarev, L.; Popovska-Mitrovikj, A.; Bakeva, V.; Sandev, T. Response to an external field of a generalized Langevin equation with stochastic resetting of the memory kernel. Phys. Rev. E 2025, 112, 024120. [Google Scholar] [CrossRef] [Scilit]
- Goychuk, A. Anomalous diffusion and directed coalescence of condensates out of equilibrium. arXiv 2025, arXiv:2506.07753. [Google Scholar]
Disclaimer/Publisher’s Note: The statements, opinions and data contained in all publications are solely those of the individual author(s) and contributor(s) and not of MDPI and/or the editor(s). MDPI and/or the editor(s) disclaim responsibility for any injury to people or property resulting from any ideas, methods, instructions or products referred to in the content. |
© 2025 by the authors. Licensee MDPI, Basel, Switzerland. This article is an open access article distributed under the terms and conditions of the Creative Commons Attribution (CC BY) license (https://creativecommons.org/licenses/by/4.0/).