Search Results (69)

Commercial forest plantations (CFPs) provide timber and ecosystem services, particularly carbon (C) sequestration, but the performance of native tropical hardwoods in pure versus mixed systems is still poorly understood. We evaluated growth, productivity, biomass, and C storage in 17-year-old plantations of Tabebuia rosea, T. donnell-smithii, and Swietenia humilis in western Mexico. Four plantation systems were assessed: pure T. rosea (PPT1), pure T. donnell-smithii (PPT2), mixed T. rosea + T. donnell-smithii (MPT1T2), and mixed T. donnell-smithii + S. humilis (MPT2S). Tree structure (DBH, height, basal area, volume), litter layer, and soils (0–15 cm) were measured. Thirty trees per species were destructively sampled to develop species-specific biometric models. Model performance was evaluated with adjusted R², RMSE, and residual analysis. PPT1 was the most productive system (39.8 m³ ha⁻¹; 55 Mg C ha⁻¹), while PPT2 had the lowest values (20.5 m³ ha⁻¹; 45.1 Mg C ha⁻¹). MPT1T2 increased basal area (+29.8% vs. PPT1) and litter layer C (3.3 Mg C ha⁻¹; +190% vs. PPT2) but did not surpass PPT1 in standing volume. Soil C was highest in PPT1 (36.5 Mg C ha⁻¹). Biometric models achieved high accuracy (R² = 0.91–0.99), confirming DBH as a reliable predictor of biomass and C. We conclude that pure T. rosea maximizes short-term productivity and soil C, whereas mixed systems diversify C allocation by enhancing litter layer pools. These findings highlight the complementary roles of pure and mixed CFPs and provide reliable models for C accounting in tropical hardwood plantations. Full article

Norway spruce (Picea abies (L.) H. Karst.) is a primary forest-forming species in the European part of Russia, both in terms of its distribution and economic importance. A number of studies indicate that one of the reasons for the disturbance of spruce forests is linked to rising temperatures, particularly the detrimental effects of extreme droughts. The aim of our research is to identify changes in the structural and functional organization of mature spruce forests at the center of the East European Plain. The study was conducted in intact spruce forests using resurveyed vegetation relevés within the Smolensk–Moscow Upland, with relevés repeated after 40 years (in 1985 and 2025). Changes in structural and functional parameters of spruce communities were analyzed. The results showed that significant disturbances of the tree layer led to changes in the vegetation of subordinate layers, as well as the successional dynamics of spruce forests. It was found that following the collapse of old-growth spruce stands, two types of secondary succession developed: (1) with the renewal of spruce and (2) with active development of shrubs (hazel and rowan) and undergrowth of broadleaved species. It was also demonstrated that the typological diversity of the studied communities changed over 40 years not only due to the loss of the tree layer and the formation of new “non-forest” types but also because several mixed spruce-broadleaved communities transitioned into broadleaved ones, and pine–spruce communities of boreal origin shifted to nemoral types. An analysis of the complete species composition of spruce forests based on Ellenberg’s scales scoring revealed changes in habitat conditions over the 40-year period. A noticeable trend was an increase in the proportion of thermophilic and alkaliphilic species, indicating a shift toward a nemoral vegetation spectrum. It is expected that under the current forest management regime, the next 40 to 60 years will see a decline in the proportion of spruce within mixed stands, potentially culminating in the complete collapse of monospecific spruce forests in the center of the East European Plain. Full article

Global climate change is driving profound transformations in forest ecosystems, particularly in monsoon-influenced regions of the Pacific coast of Asia, such as the Sikhote-Alin Mountains. Long-lived conifer species, notably Korean pine (Pinus koraiensis Siebold & Zucc.), play a central ecological role in mixed broadleaf–coniferous forests of the region. We examined how the radial growth response of Korean pine to climatic variability changes with tree age and ontogenetic stage, using 191 increment cores from trees ranging from early virginile to old generative stages. We employed two approaches: (i) a stage-based analysis, constructing tree-ring chronologies for each ontogenetic stage; (ii) an individual-tree analysis, applying correlation and regression directly to individual ring-width series. Climate–growth relationships were assessed using monthly temperature, precipitation, and drought indices (PDSI, SPEI). For the stage-based approach, radial growth was positively correlated with the mean August temperature of the previous year (up to r = 0.61), minimum November temperature (up to r = 0.50), and summer drought indices (up to r = 0.57). Age-related trends in climate sensitivity, assessed from regression models under both approaches, were significant for 9 of the 18 monthly climate variables examined. For stage-specific chronologies, simple regressions across six ontogenetic stages described up to 98% of the variance, whereas cambial-age-based relationships were much weaker (R² = 0.03–0.14). These findings highlight the importance of accounting for ontogenetic structure in dendroclimatic analyses and climate reconstructions. Such insights are critical for understanding long-term forest dynamics and informing climate adaptation strategies in Korean pine-dominated ecosystems. Full article

The grey-headed woodpecker (Picus canus) (GHW) is one of the least-studied European woodpeckers, listed in Annex I of the Birds Directive. We examined the key environmental characteristics that determine the possibility of GHW occurrence in vast forests in northeast Poland. Woodpeckers were inventoried in spring on 54 study plots (4 km²) covering 20% of the forest area. Active territories were detected and mapped using the playback experiment of territorial voices and drumming. The generalized linear model GLM, random forest RF, and Boosting were used for modeling. GLM was used to indicate the most critical factors affecting the abundance of GHW. The number of territories in a single study plot ranged from 0 to 3; the most frequent were areas without woodpeckers. The probability of the nesting of the GHW was increasing at plots with watercourses, a bigger share of mixed forest area, and a proportion of stands over 120 years old. The calculation for all 400 quadrats allowed us to estimate the population size at approximately 180–200 breeding pairs. The overall density of GHW in the study area was assessed at 0.13/km², while at the optimal quadrats, it increased to about 0.75/km². Preference for watercourses was linked to alders growing along water banks. Near the water, there are often small meadows where the GHW can prey on ants. In turn, old-growth forests above 120 years old increased the probability of the presence of the GHW. There are more dead and dying trees in older forests, which are the ones the GHW chooses to excavate cavities. To effectively protect the habitats of the GHW, it is necessary to maintain a larger area of stands over 120 years old, mainly on wet sites. Full article

Old-growth forests are rare in Europe, yet they play a critical role in biodiversity and carbon storage. This study examines the structural dynamics of the Janj old-growth forest in the Dinaric Alps using repeated field measurements from 2011 and 2021 at 39 systematically arranged 12 m radius plots. All trees (DBH ≥ 7.5 cm), regeneration (10 cm height to 7.5 cm DBH), and coarse woody debris (CWD) were assessed. Results revealed that total basal area declined by 3.5 m² ha⁻¹ over the decade, primarily driven by significant reductions in stem density for silver fir (p = 0.001) and Norway spruce (p = 0.001). In contrast, European beech maintained a stable basal area throughout the study period. Moreover, silver fir exhibited a significant increase in mean diameter (p = 0.032) and a pronounced rise in regeneration individuals (t = 3.257, p = 0.002). These findings underscore a gradual compositional shift towards European beech dominance, with conifers facing higher mortality in larger diameter classes. The substantial volume of CWD (463 m³ ha⁻¹) highlights advanced decay dynamics consistent with mature forest conditions. This study emphasizes the value of repeated measurements to capture subtle yet important successional changes in primeval forests, which is essential for conservation planning and sustainable forest management. Full article

Variation patterns in plant functional traits and their interrelationships play a crucial role in understanding species coexistence mechanisms and ecological differentiation within local plant communities. However, the dynamic patterns of plant functional traits across different forest successional stages remain insufficiently understood. Here, we investigated the woody species composition of subtropical evergreen–deciduous broadleaved mixed forest across 75 plots, representing three successional stages (20-year-old secondary forest, 35-year-old secondary forest, and old-growth forest (>80 years)), in Xingdoushan and Mulinzi National Nature Reserves, Hubei Province, Central China. We measured four functional traits of woody plants: leaf area (LA), specific leaf area (SLA), leaf dry matter content (LDMC), and wood density (WD). For each different age plant community, we calculated (1) species abundance-weighted mean community trait values, and (2) species-level mean trait values. We applied trait gradient analysis to partition and assess correlations of four functional traits across communities of different successional stages, separating within-community (α components) and between-community (β components) variation. To quantify the extent to which environmental constraints influence trait expression, we used the ecological constraint index (C_i). The results revealed significant variation in the four functional traits across communities at different successional stages. Community-level mean LA and SLA decreased significantly with age, WD increased significantly with age, and there was no significant relationship between LDMC and age. The α trait components consistently varied more widely than β components at different successional stages, indicating that biological competition dominates the assembly of local forest communities across various successional stages. Correlations between the four functional traits were dynamically adjusted with the study scale (community-level and species-level) and forest age. The ecological constraints on the four functional traits varied significantly across forest successional stages, with SLA being subject to the strongest constraints. Our findings reveal that biotic competition predominantly shapes community assembly during the succession of subtropical evergreen–deciduous broadleaved mixed forests, while stronger ecological filtering in old-growth stands underscores their role in maintaining ecosystem stability. These insights support more effective conservation and restoration strategies. Full article

Seasonal droughts induced by climate change pose a significant threat to the normal growth patterns of forests in the subtropical regions of southern China. Therefore, it is crucial to explore the response of tree water use patterns to seasonal drought to maintain tree physiological activities. However, it remains unknown whether changes in dry and wet seasons have an impact on the water use patterns of trees of different ages. In this study, a two-year experiment was conducted in Eucalyptus urophylla × Eucalyptus grandis (hereinafter referred to as Eucalyptus) plantations at three ages (4, 7, and 17 years). Specifically, the water use patterns of Eucalyptus in dry and wet seasons were calculated using hydrogen stable isotopes (including the isotopes in xylem water and 0–150 cm soil layers) coupled with MixSIAR. The results showed that there were notable variations in the proportions of water absorption from different soil layers by Eucalyptus during dry and wet seasons. During the dry season (April 2024), 4-year-old and 7-year-old Eucalyptus primarily utilized water from the 40–90 cm soil layer, while 17-year-old Eucalyptus mainly relied on deep soil water at depths of 60–150 cm, with a utilization ratio of 50.9%. During the wet season (August 2023), the depth of water uptake by Eucalyptus of different ages significantly shifted towards shallow layers, and the trees primarily utilized surface soil water from the 0–60 cm layer, with utilization ratios of 59.9%, 64.8%, and 61.6% for 4-year-old, 7-year-old, and 17-year-old Eucalyptus, respectively. The water sources of Eucalyptus during dry and wet seasons were variable, which allowed Eucalyptus to cope with seasonal drought stress. The differences in the water uptake strategies of Eucalyptus between dry and wet seasons can be attributed to their long-term adaptation to the environment. Our research revealed the differences in the water utilization of Eucalyptus with various ages between dry and wet seasons in subtropical China, providing new insights for a better understanding of the adaptive mechanisms of subtropical forests in response to alterations in water conditions caused by climate change. Full article

Forest Orchidaceae are important for European temperate forests, yet their distribution and abundance have so far interested limited research. In three pure or mixed silver fir stands in the Foreste Casentinesi National Park (NP) (Northern Apennines, Italy) we analysed how structural traits in mature and old-growth forests affected orchid communities in terms of abundance of the main genera, trophic strategy and rarity in the NP. We established three 20 × 60 m plots to quantify the structure of living and dead tree community, including a set of old-growth attributes connected to large trees, deadwood, and established regeneration. In each plot, we measured the abundance of all orchid species and explored their behaviour according to the trophic strategy (autotrophy/mixotrophy, obligate mycoheterotrophy), rarity within the NP, and threatened status according to the IUCN Red List. We used multivariate ordination and classification techniques to assess plot similarities according to forest structure and Orchid Community and identify the main structural factors related to orchid features. The main structural factors were used as predictors of community traits. Forest composition (i.e., the dominance/abundance of silver fir) affected the presence of the main orchid genera: Epipactis were abundant in silver fir-dominated forests, Cephalanthera in mixed beech and fir forests. Interestingly, Cephalanthera could become limited even in beech-dominated conditions if fir regeneration was abundant and established. Old-growth attributes like the density of deadwood and large tree volume were important determinants of the presence of rare and mycoheterotrophic species. Our results provided a first quantitative description of forest reference conditions to be used in the protection and restoration of threatened and rare orchid species. Full article

Tsuga forrestii Downie, a vulnerable species endemic to China, is confined to small, fragmented habitats in the northwestern Yunnan and southwestern Sichuan Provinces, China. Understanding the current status of its communities and populations is crucial for protecting existing natural forest resources. We surveyed 33 plots in Yunnan and Sichuan, where T. forrestii is the primary dominant species. We analyzed their community characteristics including vertical stratification, species diversity, and population structure, and classified them into four distinct forest types: (1) Tsuga forrestii evergreen coniferous forest; (2) Tsuga forrestii-Lithocarpus variolosus evergreen coniferous and broad-leaved mixed forest; (3) Tsuga forrestii-Quercus guyayifolia evergreen coniferous and broad-leaved mixed forest; (4) Tsuga forrestii-Abies forrestii evergreen coniferous forest. These forests exhibited a multilayered vertical structure, and T. forrestii as a dominant species appeared in the arborous and shrub layers, and the associated taxa were mainly species of Abies, Lithocarpus, Quercus, Castanopsis, and Rhododendron. No significant differences in species richness or diversity indices or phylogenetic relatedness metrics among the forest types were found. The maximum age of the remaining wild specimens was 344 years, with individuals under 20 years or over 170 years old being rare. The average growth rate of T. forrestii, based on ring width, fluctuated over time, and the range of mean values was 0.92 to 3.31 mm/year. Established seedlings/saplings were rare and mainly found in unstable microhabitats. These findings highlight the poor regeneration and a decline in its populations. Improving its regeneration status is crucial to maintaining its population status. Full article

Forest development forms the foundation for the advancement of sustainable forest management that integrates the knowledge of natural and anthropogenic processes with ecological and biological insights. This study aims to emphasize the role of assisted natural regeneration and balanced forest development phases in fostering closer-to-nature management approaches, contributing to resilient forest ecosystems capable of self-regulation and biodiversity support in the face of anthropogenic and climatic challenges. This study focuses on forest development in Lithuania based on five National Forest Inventories (NFIs) from 2002 to 2022. We examine the tree volume structure of the growing stock by stand type and forest type series from the point of view of stand age and forest development phases. This is performed by applying the standardized methodologies of the Lithuanian National Forest Inventory. Our analysis focuses on broader patterns derived from the selected NFI data rather than stand-level details. Our findings demonstrate that long-term observation of dynamic National Forest Inventories can aid in the development of closer-to-nature forest management methods for different forest type series. In order to implement the European Union’s strategy and policy for closer-to-nature forest management, we call for the use of “assisted succession” methods in commercial forests, promoting the formation of mixed-species forest stands with multi-cohort age profiles, including old-growth all-aged forest patches of >121 years. Full article

Background: Pinus pinea L. (stone pine) produces pine nuts of high value. Its cultivation is carried out in forests and plantations, with intensive management techniques being studied to stimulate diameter growth, which is positively related to cone production. Aims: To evaluate the effect of fertilization in a 30-year-old plantation and to understand if adult trees respond to nutritional management. Methods: A trial with completely randomized block design was established with two treatments (fertilization/control) and three repetitions. The plantation, with a density of 204 trees/ha, is located in central Chile, on a sandy-loam soil with neutral pH, medium organic matter content, and a fertility condition that limits tree development. Fertilization considered the repeated application of macro (N, P, K, S, Mg) and micronutrients (B, Fe, and Zn). Periodic measurements of height, stem and crown diameter, and cone production were made up to age 36. Cone production was evaluated using mixed generalized linear models and growth variables using ANOVA (analysis of variance). Results: Significant effects of fertilization on DBH annual growth (35% higher than the control, p < 0.001) and in cone production (3 times higher, p < 0.0001) were found. Conclusions: Fertilization is a useful practice to improve the growth and cone productivity of the species. Full article

Seasonally dry mature and old-growth (MOG) forests in the western USA face increasing threats from catastrophic wildfire and drought due to historical fire exclusion and climate change. The Emerald Point forest at Lake Tahoe in the Sierra Nevada of California, one of the last remaining old-growth stands at lake level, is at high risk due to elevated fuels and tree densities. The stand supports huge trees and the highest tree diversity in the Lake Tahoe Basin and protects important raptor habitat. In this study, we simulate forest response to vegetation management and wildfire to assess the impacts of four fuel-reduction scenarios on fire behavior and stand resilience at Emerald Point. Results: Our results demonstrate that restorative forest management can greatly improve an MOG forest’s resistance to catastrophic fire. Thinning to the natural range of variation for density, basal area, and fuel loads, followed by a prescribed burn, was most effective at reducing large-tree mortality, maintaining basal area, and retaining live tree carbon post-wildfire, while reducing secondary impacts. Conclusions: Our findings highlight the value of proactive management in protecting old-growth forests in seasonally dry regions from severe fire events, while also enhancing their ecological integrity and biodiversity. Full article

Increased tree species diversity can promote forest production by reducing intra-specific competition and promoting an efficient unitization of resources. However, questions remain on whether and how mixed stands affect the dynamics of intra–annual xylem formation in trees, especially in subtropical forests. In this study, we randomly selected 18 trees from a monoculture of 63-year-old Masson pine (Pinus massoniana) growing in pure stands and mixed them with 39-year-old Castanopsis hystrix in Pinxiang, southern China. A total of 828 microcores were collected biweekly throughout the growing season from 2022 to 2023 to monitor the intra-annual xylem formation. Cell production started in early March and ended in late December and lasted about 281 to 284 days. Xylem phenology was similar between mixed and pure stands. During both seasons, the Masson pine in mixed stands showed higher xylem production and growth rates than those in pure stands. The Masson pine in mixed stands produced 45–51 cells in 2022 (growth rate of 0.22 cells day⁻¹) and 35–41 cells in 2023 (0.17 cells day⁻¹). Growth rate, and not growth seasons, determined the superior xylem growth in the mixed stands. Our study shows that after 39 years of management, Masson pine and C. hystrix unevenly aged mixed stands have a significant positive mixing effect on Masson pine xylem cell production, which demonstrates that monitoring intra-annual xylem growth dynamics can be an important tool to evaluate the effect of species composition and reveal the mechanisms to promote tree growth behind the mixing effect. Full article

Old-growth forests are essential for maintaining biodiversity, as they are formed by the complexity of diverse forest structures, such as broad variations in tree height and diameter (DBH) and conditions of living and dead trees, leading to various ecological niches. However, many efforts of old-growth forest mapping from LiDAR have targeted only one specific forest structure (e.g., stand height, basal area, or stand density) by deriving information through a large number of LiDAR metrics. This study introduces a novel approach for identifying old-growth forests by optimizing a set of selected LiDAR standards and structural metrics. These metrics effectively capture the arrangement of multiple forest structures, such as canopy heterogeneity, multilayer canopy profile, and canopy openness. To determine the important LiDAR standard and structural metrics in identifying old-growth forests, multicollinearity analysis using the variance inflation factor (VIF) approach was applied to identify and remove metrics with high collinearity, followed by the random forest algorithm to rank which LiDAR standard and structural metrics are important in old-growth forest classification. The results demonstrate that the LiDAR structural metrics (i.e., advanced LiDAR metrics related to multiple canopy structures) are more important and effective in distinguishing old- and second-growth forests than LiDAR standard metrics (i.e., height- and density-based LiDAR metrics) using the European definition of a 150-year stand age threshold for old-growth forests. These structural metrics were then used as predictors for the final classification of old-growth forests, yielding an overall accuracy of 78%, with a true skill statistic (TSS) of 0.58 for the test dataset. This study demonstrates that using a few structural LiDAR metrics provides more information than a high number of standard LiDAR metrics, particularly for identifying old-growth forests in mixed temperate forests. The findings can aid forest and national park managers in developing a practical and efficient old-growth forest identification and monitoring method using LiDAR. Full article

Vegetation cover in urban and peri-urban areas is threatened by urban sprawl, through habitat fragmentation, loss of green space, biodiversity reduction, and the urban heat island effect intensifying. The intrusion of cities into natural landscapes reduces vital ecosystem services provided by vegetation. Hence, sustainable and integrated urban planning practices are required. Our study aims to investigate the dynamics of the urban and peri-urban fabric by exploring the relationship between the green fabric distribution and recent trends in urban expansion, focusing specifically on the peri-urban areas of Iași Municipality, Romania. We designed a mixed-method approach combining a multivariate analysis of four critical indicators (vegetation cover, built-up space, land surface temperature, and population density), emerging hot-spots, and space-time cubes in a GIS environment to achieve our research aims. Our results demonstrate that uncontrolled urban expansion has manifested in diverse patterns, impacting territories next to road transport networks and with construction-suitable topography. Concurrently, the development of green spaces prevails in forests and unexpected locations such as brownfields, railway corridors, and old industrial zones, through the growth of urban greenery. This approach provides a comprehensive understanding of how urban sprawl impacts the environment and how different land types are prone to this transformation, creating a path towards sustainability, resilience, and equitable development. Full article