Search Results (62)

The Caucasus region, characterized by its complex topography and diverse climatic regimes, exhibits pronounced spatial variability in temperature and precipitation patterns. This study investigates the seasonal behavior of air temperature, precipitation, vertical temperature gradients, and inversion phenomena across distinct landscape types using observational data from 63 meteorological stations for 1950–2022. Temperature trends were analyzed using linear regression, while vertical lapse rates and inversion layers were assessed based on seasonal temperature–elevation relationships. Precipitation regimes were evaluated through Mann-Kendall trend tests and Sen’s slope estimators. Results reveal that temperature regimes are strongly modulated by landscape type and elevation, with higher thermal variability in montane and subalpine zones. Seasonal temperature inversions are most frequent in spring and winter, especially in western lowlands and enclosed valleys. Precipitation patterns vary markedly across landscapes: humid lowlands show autumn–winter maxima, while arid and semi-arid zones peak in spring or late autumn. Some landscapes exhibit secondary maxima and minima, influenced by Mediterranean cyclones and regional atmospheric stability. Statistically significant trends include increasing cool-season precipitation in humid regions and decreasing spring rainfall in arid areas. These findings highlight the critical role of topography and landscape structure in shaping regional climate patterns and provide a foundation for improved climate modeling, ecological planning, and adaptation strategies in the Caucasus. Full article

The Caucasian Sea was among the vast tropical water masses that existed on Earth in the Mesozoic. The knowledge of Kimmeridgian–Hauterivian deposits from the central Western Caucasus can facilitate the understanding of the Caucasian paleogeography at the Jurassic–Cretaceous transition. Taking into account the scale of the study area and its geological complexity, a generalized synthesis of the published information seems to be an appropriate option to propose a tentative paleogeographical model. Some original field and laboratory studies, including the examination of the composition of Hauterivian alluvial sandstones, contribute to this model. Kimmeridgian–Hauterivian deposits crop out in the northern, western, and southern domains of the study area, but older rocks are exposed in its central and eastern parts. The Caucasian Sea covered the study area in the early Kimmeridgian, but a large land appeared in the late Kimmeridgian and existed until the end of the Hauterivian despite certain shoreline shifts. The land was eroded deeply, with exposure of pre-Upper Jurassic rocks, including Precambrian–Paleozoic crystalline complexes, and the sedimentary material was delivered to an alluvial plain on its periphery. The registered sea–land interplay was controlled tectonically. Full article

The smooth snake (Coronella austriaca) has a wide but fragmented distribution across the Western Palearctic, with limited records in Kazakhstan. This study aims to provide an updated distribution map and to explore the phylogenetic affinities of C. austriaca in Kazakhstan. The species had not been documented for over 60 years until its recent rediscovery in the region. Field surveys conducted between 2019 and 2024 in the West Kazakhstan and Aktobe regions have yielded novel records, including the southernmost observation in the Mugodzhar mountain range. Mitochondrial DNA analysis confirmed that the Kazakh populations belong to the Eastern lineage, sharing haplotypes with specimens from the northwestern Caucasus and Crimea. Habitat assessment revealed that the species’ distribution is restricted to open habitats of petrophytic and calciphyte steppe communities on chalks and rocky steppes. Of particular interest is that 70% of the observed individuals exhibited patternless coloration, suggesting the potential for regional morphological variation. These findings offer the first evidence for the phylogenetic affiliation of the smooth snake in Kazakhstan and reflect its rarity in the country, highlighting the need for local conservation efforts, including habitat protection and population monitoring. Full article

Climate-induced hydrological risks and deteriorating infrastructure present major challenges for small river basins in border regions, particularly in non-EU countries with limited institutional capacity and funding. These issues are especially acute in post-socialist contexts, where outdated hydrotechnical systems no longer meet current environmental and safety standards. This study investigates the vulnerabilities of the Uzh River basin in Uzhhorod, Ukraine—a non-EU border city with strong ecological and institutional ties to neighboring EU regions—and proposes an adaptive river management model tailored to such environments. An integrated assessment of flood protection systems, sediment transport, drainage performance, and governance gaps was conducted to inform the proposed framework, which combines structural and ecosystem-based interventions with a focus on transboundary water governance. Unlike many existing approaches that lack mechanisms for localized implementation and cross-border coordination, this model offers a transferable, evidence-based methodology for enhancing flood resilience and hydrological sustainability in similar urban areas. The insights are relevant to border cities across Eastern Europe, the Western Balkans, and the South Caucasus, contributing to both engineering practice and regional policy by aligning hydrotechnical solutions with cooperative climate adaptation strategies. Full article

This study aimed to reveal intraspecific variations in two Dryomys species distributed in Türkiye, based on mitochondrial DNA cytochrome b gene sequences, and to discuss the factors driving these variations in the context of phylogeography and genetic species concepts. As a result of Maximum Likelihood, Bayesian Inference, and Network analyses, which included haplogroups or lineages from Italy, Russia, the Caucasus, and Iran identified in previous studies, along with Turkish haplotypes, three major clades (MC1, MC2, and MC3) were identified within Dryomys nitedula. These clades began to diverge evolutionarily in the middle of the Late Miocene (8.82 million years ago) and exhibit significant genetic differences from one another. The Turkish haplotypes were divided into five distinct lineages (N1–N5), each within five subclades (SC1–SC5), which were nested within these MCs. These lineages, their geographical distributions, and the subspecies defined in previous studies that correspond to these lineages are as follows: N1 from the Thrace region (Dryomys nitedula wingei), N2 from the Black Sea region (potentially a new subspecies), N3 from western and central Anatolia (Dryomys nitedula phrygius), N4 from northeastern Anatolia (Dryomys nitedula tichomirowi), and N5 from eastern Anatolia (Dryomys nitedula pictus). The N2 lineage, distributed in areas close to the coastal side of the Eastern Black Sea region and with a range close to both N3 (D. n. phrygius) and N4 (D. n. tichomirowi), exhibited high genetic differentiation from these two lineages and was a candidate to be treated as a new subspecies of Dryomys nitedula in Türkiye. The N5 lineage, which includes haplotypes from the distribution areas of the populations initially classified as Dryomys pictus and later as Dryomys nitedula pictus in previous studies, was found to be more closely related to Dryomys nitedula kurdistanicus from the Zagros Mountains than to D. n. pictus from the central regions of Iran. Combining the results of this study with previous research, it is clear that the D. nitedula lineages in Türkiye, along with haplogroups or subspecies in neighboring regions diverged between the middle Late Miocene and Middle Pleistocene. This divergence is believed to have been driven by climatic cycles and geomorphological processes that shaped the topography of their distribution range. The high genetic diversity observed in the lineages of Anatolia suggests that the region may have served as a glacial refuge for D. nitedula. Similarly to the processes and factors shaping the evolution of D. nitedula, Dryomys laniger was found to have diverged into two lineages, western (L1) and eastern (L2 or Dryomys anatolicus), within its distribution range during the Late Pliocene (2.94 Mya). To make a more accurate taxonomic assessment of D. laniger, a larger number of samples is needed, and the distribution limits should be more clearly defined. Full article

By exploring seed samples from vineyards and natural habitats across the Eurasian range of Vitis vinifera, our analysis revealed substantial morphological variation within populations. Through the analysis of domestication index values, probabilities, and entropy, we assessed seed diversity. Samples with high domestication probability values—predominantly from vineyards—exhibited low heterogeneity and entropy, with similar patterns observed in natural habitats, suggesting the presence of feral vines. In parallel, seeds with low domestication index values, found mainly in natural habitats, also displayed low entropy and are likely associated with Vitis sylvestris or other wild Vitaceae species. Intermediate domestication values pointed to hybrid swarms, highlighting the crucial role of hybridization in the development of modern grapevine cultivars. The study identified mixed populations across the Iberian Peninsula, Italy, and the South Caucasus, which act as significant gene reservoirs. A domestication gradient is evident, with higher domestication rates in the South Caucasus compared to Western Europe and East Asia. The results demonstrate the significance of these mixed populations as repositories of genetic diversity, underscoring their conservation value, particularly considering the negative impact of habitat alterations, especially in riparian forests due to major public works. Full article

Common walnut (Juglans regia L.) is a valuable nut crop widely distributed within the temperate climate zone. It has several secondary centers of origin associated with refugia formed during the LGM (Last Glacial Maximum) period, one of which was presumably located in the Caucasus region. Currently, walnut populations of the Caucasus are studied fragmentarily, especially in the northeast. Our study is aimed at investigating the genetic diversity, population structure, and possible walnut introduction processes in the Caspian region of the Northeast Caucasus (Dagestan). Genotyping using 11 SSR markers was performed for 158 samples from local walnut populations. Also, to identify the introduction facts, 42 varieties of different geographical origin were added to research. The SSR markers genetic diversity in the analysis of population varied from four to 20 alleles per locus, with an average value of 9.82. Analysis of the genetic structure revealed significant genetic differences between the subpopulations of the Mountain and Flatland Dagestan. Presumably, the structure of the gene pool could have been influenced by terrain factors and ethnocultural characteristics. Subpopulations of Flatland Dagestan have an intermediate position between the western cultivars and the gene pool of Mountain Dagestan, which subpopulations showed significant genetic isolation. In the studied samples set of population selections, the established introduction facts were rare. We have suggested that a significant part of the walnut gene pool from Mountain Dagestan has an autochthonous origin from plants that survived the LGM in the Caucasian refugia. Our results are the basis for further studies of the local gene pool and verification of hypothesis about the presence of an indigenous germplasm preserved in this region from the LGM period. Full article

The Eurasian otter (Lutra lutra) is a widespread semiaquatic carnivorous mammal in Eurasia. The nominate subspecies (L. l. lutra) occupies vast areas between Western Europe and the Russian Far East, but its phylogeography and genetic diversity are still unclear across Northern Eurasia. Another subspecies, L. l. meridionalis, located in the Caucasus mountains, is morphologically almost identical to L. l. lutra but needs genetic revision. We compared the genetic diversity of Eurasian otters from Russia and Armenia using a mtDNA fragment (820 bp) and 20 autosomal microsatellite loci (N = 117). A total of 32 haplotypes were observed with 17 novel haplotypes. The MtDNA median-joining network was mostly star-shaped with a branch of haplotypes from Far Eastern Russian otters. Both mtDNA analysis and Bayesian clustering of microsatellite data indicated that Far Eastern otters are more genetically differentiated than European and Siberian otters (Φst = 0.565 and 0.467; Rst = 0.306 and 0.256), as well as Caucasian otters (L. l. meridionalis) from Russia and Armenia (Φst = 0.515, Rst = 0.253). Haplotype and nucleotide diversities of Far Eastern otters are also the highest between sample groups (H = 0.882, π = 0.003) and, of Caucasian otters, the lowest (H = 0.464, π = 0.001). Our results suggest Caucasian otters are more similar to the otters from European Russia than to the other groups (but with lower genetic diversity) and lack the genetic variability typical to different subspecies. On the contrary, otters from the Russian Far East are more genetically differentiated, have higher genetic diversity than otters from Europe, and likely belong to another genetic lineage. Full article

Territorial inventories of geosites remain on the international agenda, and they can help in acquiring information for solving pure geological research tasks. New field investigations in the northern part of Mountainous Adygeya (geoheritage-rich territory in the western Greater Caucasus) permitted us to extend our knowledge of its geosites. Five geosites were described qualitatively and assessed semi-quantitatively, namely the Polkovnitskaya River Valley (ammonite-bearing concretions of Aptian glauconitic sandstones), the Little Khadzhokh River Valley (Aptian glauconitic sandstones with fossils and trace fossils), the Shakhan and Middle Khadzhokh River Valley (Upper Jurassic variegated siliciclastics, Hauterivian fluvial and deltaic sandstones, mixed ancient and modern clastic material), the Big Khadzhoh Waterfall (splendid waterfall and exposures of locally folded Oxfordian–Kimmeridgian limestones), and the Verblyud Mountain (small, camel-shaped cuesta’s remnant). They were ranked nationally (one geosite), regionally (three geosites), and locally (one geosite). Close examination of the considered geosites permitted to register pure geological peculiarities (changes in the dip direction between sedimentary packages), which indicate the tectonic activity across the Jurassic–Cretaceous transition. Additionally, complex accessibility patterns were established in some geosites, and these patterns should be addressed in further refinements of the semi-quantitative approaches to geosite assessment. Full article

Granitoids are known to crop out on ancient seashores, but the related geological evidence remains limited. The information from the Western Caucasus sheds light on the stratigraphical distribution of coarse siliciclastic beds associated with late Carboniferous granitoids of the Dakh, Rafabgo, and Sakhray crystalline massifs. For the purposes of this study, the available information was reviewed and verified against the modern stratigraphical scales. It is established that the considered coarse sisliciclastic beds occur at five stratigraphical levels of the Triassic–Jurassic succession. A rocky seashore with granitoid exposures existed for a short time around the Sakhray and probably Rufabgo massifs at the very beginning of the Triassic. The Dakh Massif possessed such a shore twice (at least), i.e., in the Norian–Rhaetian and the Early Toarcian. However, it cannot be excluded that rocky shores persisted there for >50 Ma. Generally, the Western Caucasus provides an example of granitoid exposures on Mesozoic seashores and adds knowledge of the global distribution of rocky shores in the Triassic and Jurassic periods. Full article

Geomorphosite inventory remains an urgent task for many geographical domains where it can facilitate the exploration of geoheritage resources. The present study reports a new and sufficient portion of information from Mountainous Adygeya in southwestern Russia, which is a geoheritage-rich area in the western part of the Greater Caucasus Mountains. The Cossack, Monk, and Three Teeth mountains were analyzed qualitatively and semi-quantitatively. Their morphology and geological settings were described, their geoheritage properties were scored by several criteria, and their cultural meanings were considered. It was established that the total heritage values of these geomorphosites are comparable. The challenging accessibility of these landforms is offset by the availability of sites for distant observation. Each proposed geomorphosite has its own cultural meaning reflected by its name. The proposed geomorphosites are constituents of the geoheritage resources of Mountainous Adygeya, and they can be exploited via local product branding and georoute functioning. Full article

The grapevine was one of the earliest domesticated fruit crops and has been cultivated since ancient times. It is considered one of the most important fruit crops worldwide for wine and table grape production. The current grape varieties are the outcome of prolonged selection initiated during the domestication process of their wild relative. Recent genetic studies have shed light on the origins of the modern domestic grapevine in western Europe, suggesting that its origin stems from the introgression between eastern domestic grapes and western wild grapes. However, the origin of ancient grapevines remains largely unexplored. In this study, we conducted an extensive analysis of 2228 well-preserved waterlogged archaeological grape pips from two sites in Sardinia (Italy), dated to the Late Bronze Age (ca. 1300–1100 BC) and the Iron Age (4th and 3rd centuries BC). Using morphometrics and linear discriminant analyses, we compared the archaeological grape pips with modern reference collections to differentiate between wild and domestic grape types and to investigate similarities with 330 modern cultivars. Grape pips from the Late Bronze Age displayed a high percentage of similarity with domesticated grapevines, with a small percentage assigned to wild ones, while the majority of grape pips from the Iron Age were classified as domestic. Discriminant analyses revealed that both white and red grape varieties were cultivated during the Late Bronze and Iron Ages, suggesting a high level of diversification in grape cultivation. Furthermore, a high percentage of archaeological grape pips from both periods showed strong similarities with modern cultivars from the Caucasus and Balkans. This suggests that the great diversity of grapevines present in Sardinia could result from interbreeding between western Asian cultivars and local grapevines that began in the Late Bronze Age. Additionally, a substantial proportion of archaeological grape pips exhibited similar morphometric characteristics to two important Mediterranean grape cultivars: “Muscat à petits grains blancs” and “Garnacha”. Full article

Located at the crossroads of two continents and at the southeastern edge of the Mediterranean Basin, Anatolia was one of the most important Pleistocene glacial refugia in the Western Palaearctic. As part of the Irano-Anatolian, Caucasus and Mediterranean Basin biodiversity hotspots, this region is also home to a rich avian community including nearly 400 breeding species. Nevertheless, research addressing the genetic structure and diversity of local bird populations is limited, and information on glacial refugia in this region is still scant, especially when compared to other large Mediterranean peninsulas, namely the Balkan, Italian and Iberian ones. In this study, we contribute to filling this gap by addressing the biogeographic pattern of four common resident songbirds—the Eurasian blue tit (Cyanistes caeruleus), the great tit (Parus major), the Eurasian chaffinch (Fringilla coelebs) and the Eurasian blackbird (Turdus merula)—and one endemic species—the Krüper’s nuthatch (Sitta krueperi)—by amplifying two mitochondrial DNA genes in individuals from Anatolia (n = 329) and comparing their sequences to those of conspecifics from the rest of their distribution range across the western Palaearctic (n = 357) deposited in public databases. The overall genetic structure of these species is consistent with a scenario of isolation for multiple populations in different refugia across Anatolia and subsequent secondary contact in the wake of ice retreat, which makes this region a hotspot of genetic diversity for both widespread and endemic avian species. Full article

In instances of severe infestations, Nepticulidae larvae can inflict damage on cultivated plants. Previously, it was assumed that the Prunus-feeding Nepticulidae have continuous distribution from Europe to the neighboring Caucasus. During recent fieldwork in the Caucasus, leaf mines were found on plum trees that initially resembled those of Stigmella plagicolella (Stainton) in Europe. However, upon rearing the adults, significant differences emerged, leading to the hypothesis that a different Prunus-feeding species exists in the Caucasus; this challenges previous records in Western Asia. This paper presents the outcomes of our morphological, molecular, and statistical investigations, unveiling S. colchica sp. nov., a previously unknown potential plum-tree pest. Distinguished by male genitalia characteristics, the new species differs from S. plagicolella. The inter- and intraspecific divergences between S. colchica sp. nov. and S. plagicolella range from 3.5% to 6.02%. Moreover, the utilized delimitation algorithms reliably clustered two species separately, as does our mitotype network. A statistical analysis also shows a discernible trend between the leaf mines of S. colchica sp. nov. and S. plagicolella. This unexpected discovery not only documents a new potential pest, enhancing our understanding of the Caucasian fauna, but also contributes to the broader biological inventory. Full article

In our study, using the analysis of DNA barcodes and morphology (wing color, male genitalia, and female sphragis shape), we show that the group of species close to P. mnemosyne comprises the western and eastern phylogenetic lineages. The eastern lineage includes P. stubbendorfii, P. glacialis, and P. hoenei. The western lineage includes three morphologically similar species: P. mnemosyne (Western Eurasia), P. turatii (southwestern Europe), and P. nubilosus stat. nov. (Turkmenistan and NE Iran), as well as the morphologically differentiated P. ariadne (Altai). The latter species differs from the rest of the group in the presence of red spots on the wings. Parnassius mnemosyne s.s. is represented by four differentiated mitochondrial clusters that show clear association with specific geographic regions. We propose to interpret them as subspecies: P. mnemosyne mnemosyne (Central and Eastern Europe, N Caucasus, N Turkey), P. mnemosyne adolphi (the Middle East), P. mnemosyne falsa (Tian Shan), and P. mnemosyne gigantea (Gissar-Alai in Central Asia). We demonstrate that in P. ariadne, the red spots on the wing evolved as a reversion to the ancestral wing pattern. This reversion is observed in Altai, where the distribution areas of the western lineage, represented by P. ariadne, and the eastern lineage, represented by P. stubbendorfii, overlap. These two species hybridize in Altai, and we hypothesize that the color change in P. ariadne is the result of reinforcement of prezygotic isolation in the contact zone. The lectotype of Parnassius mnemosyne var. nubilosus Christoph, 1873, is designated. Full article