Abstract
The study considers a nonlinear multi-parameter reaction–diffusion system of two Lotka–Volterra-type equations with several delays. It treats both cases of different diffusion coefficients and identical diffusion coefficients. The study describes a few different techniques to solve the system of interest, including (i) reduction to a single second-order linear ODE without delay, (ii) reduction to a system of three second-order ODEs without delay, (iii) reduction to a system of three first-order ODEs with delay, (iv) reduction to a system of two second-order ODEs without delay and a linear Schrödinger-type PDE, and (v) reduction to a system of two first-order ODEs with delay and a linear heat-type PDE. The study presents many new exact solutions to a Lotka–Volterra-type reaction–diffusion system with several arbitrary delay times, including over 50 solutions in terms of elementary functions. All of these are generalized or incomplete separable solutions that involve several free parameters (constants of integration). A special case is studied where a solution contains infinitely many free parameters. Along with that, some new exact solutions are obtained for a simpler nonlinear reaction–diffusion system of PDEs without delays that represents a special case of the original multi-parameter delay system. Several generalizations to systems with variable coefficients, systems with more complex nonlinearities, and hyperbolic type systems with delay are discussed. The solutions obtained can be used to model delay processes in biology, ecology, biochemistry and medicine and test approximate analytical and numerical methods for reaction–diffusion and other nonlinear PDEs with delays.
Keywords:
reaction–diffusion equations with delay; Lotka–Volterra-type systems of equations; nonlinear systems of PDEs; reductions; exact solutions; generalized separable solutions; solutions in terms of elementary functions MSC:
35K57; 35R10; 35C05
Contents
1. Introduction
1.1. Preliminary Remarks. The Lotka–Volterra-Type Diffusion System with Delays
1.2. The System of Delay PDEs in Question. Exact Solutions and Generalized Separable Solutions
1.3. Simplest Solutions (Stationary Points)
2. Reductions and Exact Solutions of the System of Delay PDEs with Different Diffusion Coefficients (a1 ≠ a2)
2.1. Reduction of the System with Three Delays to a Single Second-Order Linear ODE without Delay
2.2. Reduction of the System of PDEs with Three Delays to a System of Two Second-Order ODEs without Delays
2.3. Exact Solutions in Terms of Elementary Functions to the Reduced System of Two Second-Order ODEs
2.4. Reduction of a System of PDEs with Three Delays to a System of Three Second-Order ODEs without Delays
2.5. Exact Solutions in Terms of Elementary Functions to the Reduced System of Three Second-Order ODEs
2.6. Reduction of the System of PDEs with a Single Delay to a Unsteady System of Two First-Order Delay ODEs
2.7. The First Integral and Exact Solutions of the Reduced System of Two First-Order Delay ODEs
3. Reductions and Exact Solutions of the System of Delay PDEs with Identical Diffusion Coefficients (a1 = a2)
3.1. Reduction of the System of PDEs with Three Delays to a System of Two Second-Order ODEs without Delay and One Linear PDE
3.2. Exact Solutions to the Reduced System Consisting of Two ODEs and One PDE
3.3. Reduction of the System of PDEs with a Single Delay to a System of Two First-Order Delay ODEs and One Linear PDE
4. Related Systems of PDEs with Several Space Variables and Delays. Other Generalizations
4.1. Reductions and Exact Solutions of a Multidimensional System with Different Diffusion Coefficients and Delays
4.2. Reductions and Exact solutions of a Multidimensional System with Identical Diffusion Coefficients and Delays
4.3. Further Generalizations and Modifications
5. Brief Conclusions
1. Introduction
1.1. Preliminary Remarks The Lotka–Volterra-Type Diffusion System with Delays
Linear and nonlinear differential equations, ordinary and partial, with delay are not uncommon for mathematical modeling of phenomena and processes in various areas of theoretical physics, mechanics, control theory, biology, biophysics, biochemistry, medicine, ecology, economics and technical applications.
Below are a few factors that lead to the need to introduce delay into mathematical models described by differential equations. In biology and biomechanics, the delay is due to the limited transmission rate of nerve and muscle reactions in living tissues. In medicine, in studying the spread of infectious diseases, the delay time is determined by the incubation period (the time from the moment of infection to the first signs when the disease begins to manifest itself). In the dynamics of populations, a delay arises because individuals participate in reproduction only after reaching a certain age. In control theory, delays are usually associated with the finite speed of signal propagation and the limited speed of technological processes.
The presence of delay in mathematical models and differential equations is a complicating factor, which, as a rule, leads to a narrowing of the stability region of the solutions obtained. The study and solution of ordinary differential equations (ODEs) with a delay are comparable in complexity to the study and solution of partial differential equations (PDE) without delays. Importantly, unlike simpler PDEs without delay, which often have self-similar solutions, PDEs with delay do not admit self-similar solutions.
The present article deals with a Lotka–Volterra-type reaction–diffusion system with two components and four constant delays described by two coupled PDEs with quadratic nonlinearities:
where and are the unknown functions; , ; , ; and are delay times; and are diffusion-type coefficients; and , , , , , and are some constants.
Systems of the form (1) are used in the mathematical modeling of various processes in biology, ecology, biochemistry, medicine, etc. For example, in population theory, the delays and characterize the mean reproductive age of species. At the same time, and are responsible for the time required for changes in the population size of one species to cause changes in the other. All the delays are non-negative and can be zero in some models. The terms with nonzero coefficients and make the model different from a single equation, while the coefficients are responsible for the interaction between individuals of the two populations. In the case of cooperation, when one species persists in the absence of the other or when species mutually increase each other’s growth rate, the coefficients and are positive. In the case of competition, an increase in one population leads to a decrease in the other (for example, an increase in the number of predators leads to a decrease in the prey population), and the coefficients and are negative. Lotka–Volterra cooperative models with delays, described by PDEs (1), were treated in [1,2]; for competitive models, see [3,4,5].
The special case of the system of delay PDEs (1) with , , and describes a Belousov–Zhabotinsky oscillating chemical reaction, where u and v are the bromic acid and bromide ion concentrations (see [6]).
The system of delay PDEs (1) is a generalization of simpler equations and systems outlined below.
. In the simplest case of no diffusion or delays, with and , the system of PDEs (1) degenerates into a system of ODEs [7,8], which is known as the classical Lotka–Volterra system.
. With no diffusion, when , system (1) degenerates into a simpler system of delay ODEs. For qualitative features and the dynamics of such delay systems, see, for example, [9,10,11,12,13,14].
. If , the diffusion system of PDEs (1) simplifies significantly when there are no delays, . Numerous studies are devoted to qualitative analysis of such systems (e.g., see [15,16,17,18,19] and references in [18,19]).
The system of PDEs (1) without delays admits simple traveling wave solutions
with the functions and described by a system of ODEs. The studies [20,21,22] obtained some exact solutions to this system, of the form (2), which are expressible in terms of elementary functions.
The studies [23,24,25] considered nonclassical symmetries and exact solutions to system (1) and related Lotka–Volterra-type systems in the absence of delays (see also [26,27]).
Remark 1.
For exact solutions to various single nonlinear reaction–diffusion-type equations without delays, see, for example, [26,28,29,30,31,32,33,34,35,36,37,38,39,40,41,42,43,44]. The books [26,32,37,40,43,44] describe solution methods for such equations.
. In general, the presence of delays in system (1) complicates its analysis dramatically, since the majority of analytical methods that are effective in constructing exact solutions to nonlinear PDEs without delay are either inapplicable for solving delay PDEs at all or extremely limited for this purpose.
The system of delay PDEs (1) admits simple traveling wave solutions (2) (e.g., see [1,5,45]). At present, there is only one publication [46] where two other exact solutions to system (1) were obtained and some reductions of related nonlinear reaction–diffusion systems of PDEs were described.
Remark 2.
For exact solutions to single nonlinear reaction–diffusion-type delay equations, see [47,48,49,50,51,52,53] (see also [54], where time-fractional reaction–diffusion PDEs with delay were considered). The studies [55,56] describe some exact solutions to nonlinear Klein–Gordon-type PDEs with delay.
1.2. The System of Delay PDEs in Question Exact Solutions and Generalized Separable Solutions
. In general, the multi-parameter nonlinear Lotka–Volterra-type diffusion system of delay PDEs (1) contains 12 free parameters, four of which can be removed by scaling the dependent and independent variables. We will consider a special case of system (1) while imposing one constraint, , on the free parameters. With this in mind, we set , , , and to obtain
where , (, ); , , , , , , , and are free parameters; and are some constants to be determined later. The present study does not deal with the degenerate cases of or , when system (3) becomes semi-coupled, meaning that one equation of the system is isolated, while the other one is linear in the unknown.
. A solution is called exact if it satisfies the equation in question exactly (i.e., when substituted into the equation, the solution turns it into an identity). At the same time, any approximations or simplifications of the equation are not allowed, and no a priori assumptions are used.
The term ‘exact solution’ for nonlinear PDEs with delay is normally used in cases where the solution is expressed:
- (i)
- In terms of elementary functions;
- (ii)
- In terms of elementary and special functions and indefinite integrals;
- (iii)
- In terms of solutions to ODEs without or with delay or systems of such equations.
Combinations of solutions from items (i)–(iii) are allowed. In cases (i) and (ii), an exact solution can be represented in an explicit, implicit or parametric form.
Exact solutions play an essential role as standard reference results that can be used to verify the consistency and estimate errors of various numerical, asymptotic, and approximate analytical methods for solving nonlinear PDEs with delay. These solutions can also serve as a basis for perfecting and testing computer algebra software packages such as Maple and Mathematica.
The simplest and most preferred exact solutions for testing are solutions in terms of elementary functions (see Item (i)). Currently, the vast majority of the obtained exact solutions of such equations are more complex and relate to Item (iii).
. We will be looking for exact generalized separable solutions to the system of delay PDEs (3) in the form
or in an alternative form that can be obtained from (4) by replacing the functions and with and , respectively. In some cases, we will look for more complicated solutions involving functions with two arguments. Special attention will be given to the construction of exact solutions expressible in terms of elementary functions.
1.3. Simplest Solutions (Stationary Points)
Constants (stationary points) are the simplest solutions to system (3):
These constants are determined by the formulas
2. Reductions and Exact Solutions of the System of Delay PDEs with Different Diffusion Coefficients ()
2.1. Reduction of the System with Three Delays to a Single Second-Order Linear ODE without Delay
The four delay times in system (3) are assumed to be related by the single constraint
This suggests that any three of these delays can be set arbitrarily. Notably, relation (7) holds, for example, in the following three important special cases:
We seek a generalized separable solution to system (3) in the form
where and are the stationary points (5) of system (3). We assume that the function satisfies the second-order linear ODE
The functions (8) are selected so as to ensure that under condition (7), the following simple relations hold:
Substituting (8) into (3) and taking into account relations (10) and Equation (9) followed by a simple rearrangement of terms, we arrive at the linear algebraic system
It serves to determine the parameters and .
If , the solution of system (11) is expressed as
In case (d), the parameters degenerate, , resulting in solutions of little interest.
The general solution to the linear ODE (9) with reads
where and are arbitrary constants.
Formulas (8), (13) and (14) and the first three stationary points (6) determine six nondegenerate solutions, three for and each, to the nonlinear system (3) with and four constant delays satisfying condition (7). The study [46] presents exact solutions to system (3) corresponding to the stationary point (b) in (6). These are written out below.
Solution for :
where and are arbitrary constants.
Solution for :
2.2. Reduction of the System of PDEs with Three Delays to a System of Two Second-Order ODEs without Delays
Generalized separable solutions exponential in t. We seek a generalized separable solution to system (3) in the form
with the functions , , and and parameter to be determined in the subsequent analysis.
The functions (17) are selected so as to ensure that under condition (7), the following relations hold:
Substituting (17) into (3) and taking into account (18), we obtain
with the notation used for short.
Relations (19) can be satisfied by setting
The first two equations in (20) form a closed system of ODEs for determining the functions and , while the other two equations make up an overdetermined system for the single function . On requiring that the last two equations in (20) coincide, we find the parameter and other constants:
On substituting (21) into (20), we arrive at the following system of stationary second-order ODEs:
where .
Statement 1.
Let , , be a solution of system (22). Then, the functions
where A and B are arbitrary constants () are also a solution of the system.
System (22) simplifies significantly if either or is set to zero, in which case the system will only consist of two equations.
Properties and some transformations of the system of ODEs (22) with . The autonomous systems of ODEs (22) with can be rewritten in the form
where , , and . We assume that (the special case was discussed above).
Below are three linear transformations of the system of ODEs (24) and (25), which will be required for the construction of exact solutions.
. The linear transformation
takes system (24) and (25) to the same system with other determining parameters
. The linear transformation
also takes system (24) and (25) to the same system with other parameters
We will use to refer to systems (24) and (25). Then, transformations and and their composition connect this system with three other systems of the same form, which can schematically be depicted as
These transformations allow one to multiply exact solutions to (24) and (25). In addition, the following statement holds true.
Statement 2.
Exact solutions to the system of ODEs (24) and (25). The general solution to the autonomous ODE (24) can be represented in the implicit form
where and are arbitrary constants. In general, the integral on the right-hand side of (34) is not expressible in terms of elementary functions.
Solutions to ODE (24) are expressible in terms of the Weierstrass elliptic function , which is defined explicitly via the elliptic integral [57]:
where and are free parameters called invariants. The first ODE in (33) has a solution (see [57,58]), where is an arbitrary constant. By virtue of transformation (32) and the first property in (23), Equation (24) with admits the solutions
where and are arbitrary constants.
Notably, once is known, the general solution to the linear homogeneous Equation (25) can be written as [58]:
where is any nontrivial particular solution to Equation (25).
If , Equation (25) with coincides with Equation (24). Therefore, the function is a particular solution to Equation (25) in this case. On substituting into (36), we obtain the general solution to Equation (25) with ; the representation of the function in terms of the Weierstrass elliptic function (35) can also be used.
2.3. Exact Solutions in Terms of Elementary Functions to the Reduced System of Two Second-Order ODEs
Consider a few special cases where solutions to the system of ODEs (24) and (25) are expressible in terms of elementary functions.
. For and , the function in (34) can be represented in explicit form as
Up to renaming, it coincides with a special case of an ODE discussed in the book [58] (see Equation (132) with and on page 535).
For , the general solution of Equation (38) is
where and are arbitrary constants.
For , the general solution of Equation (38) can be represented as
where and are arbitrary constants, and are Bessel functions of the first and second kind, and and are modified Bessel functions of the first and second kind. These can be expressed in terms of elementary functions as [58]:
. For and , the integral in (34) can be expressed in terms of elementary functions, and the function can be written explicitly as
where is an arbitrary constant.
For , the general solutions of Equation (25) corresponding to solutions (40) are
where
and and are arbitrary constants.
Substituting into Equation (25) and in view of the first solution (40), we find the following allowed values of the parameters:
As a result, taking into account (36), we obtain four general solutions for different :
where .
In view of (36), we obtain four general solutions for different :
where .
. For , , the radicand in (34) can be factorized to . In this case, the integral can be expressed in terms of elementary functions, and the function can be represented in explicit form as
The respective general solutions of Equation (25) are
where .
The respective general solutions of Equation (25) are
where .
. Apart from the above exact solutions for , Equation (24) admits the solutions
The following replacements should be made in the respective solutions for : , cos, tan, with , sin, cot, and , cosh, tanh, with , sinh, coth, , respectively.
In particular, besides the first solution in (40) with and the first solution in (41) with , there are solutions
Moreover, apart from the first solution in (43) with and the first solution in (44) with , there are solutions
Table 2 summarizes the exact solutions in terms of elementary functions to the system of ODEs (24) and (25) for various values of the parameters b and . The space variable x can everywhere be replaced with , where is an arbitrary constant.
The exact solutions to the original Lotka–Volterra system of delay PDEs (3) for and and under condition (7) are determined by substituting the functions and from Table 2, where , , and , into Formula (17) with and .
Remark 3.
Remark 4.
By setting in system (22) and denoting , , , and , one can obtain exact solutions for the pair of functions and analogous to those described above for the pair of functions and for .
2.4. Reduction of a System of PDEs with Three Delays to a System of Three Second-Order ODEs without Delays
System (3) with four delays that satisfy the single relation (7) also admits generalized separable solutions linear in time t:
An analysis similar to that above results in the following parameters of Equation (3):
where is an arbitrary constant. In this case, the functions , , and are described by the stationary system of ODEs
Let us look at the special case . By adding up the first two equations of (49) multiplied by and , we obtain an isolated ODE for . We rewrite this equation, together with the third and first ODEs of (49) and the algebraic relation for , as the mixed algebraic-differential system of equations
which contains three second-order ODEs.
2.5. Exact Solutions in Terms of Elementary Functions to the Reduced System of Three Second-Order ODEs
First, let us describe two simple classes of exact solutions to system (50) with .
. Exact solution to system (50) with :
where , …, are arbitrary constants.
. Exact solution to system (50) with :
. The first two ODEs of system (50) coincide, up to renaming, with system (24) and (25) at and , whose exact solutions were described above. For known , the second equation in (50) for is a second-order linear homogeneous ODE, which has a particular solution . Hence, the general solution of this equation is given by Formula (36). Once is found, the third equation in (50) is a second-order linear nonhomogeneous ODE for , with the particular solution of the homogeneous ODE already known. Considering the aforesaid and using relevant formulas from [58], one can find the general solution to the ODE for . As a result, we obtain the solution to system (50) in the form
where , …, are arbitrary constants, and is a particular solution to the third equation of system (50), which is defined as
Here, the function is the Wronskian determinant. Simple computations show that .
Exact solutions to system (50) can be obtained by substituting the functions and from Table 3 into formulas (51) and (52).
. For arbitrary , the function in solution (51) can be taken to be the Weierstrass elliptic function:
where is an arbitrary constant.
2.6. Reduction of the System of PDEs with a Single Delay to a Unsteady System of Two First-Order Delay ODEs
Let the four delays in system (3) be identical:
We look for generalized separable solutions to system (3) in the form
with the functions , , , and to be determined in the subsequent analysis.
The functions (54) are selected to ensure that the composite arguments of are only dependent on t. We impose the following additional condition (a linear differential constraint) on the function :
where the constants and are to be found in the subsequent analysis. Notably, without loss of generality, one can set in (55) in the nondegenerate case of , since the translation of by a constant only leads, by virtue of representation (54), to new definitions of and .
Substituting (54) into (3) and taking into account (55), we get
where the notations , , and are used for short.
Relation (56) can be satisfied if we set
The system of ODEs (57) is overdetermined because it consists of four equations for three functions , , and . On requiring that the last two equations of system (57) coincide, we find the parameter and other constants:
For , it follows from (58) that , and hence, we can set in Equations (55) and (57). The general solution of Equation (55) with is given by Formula (14), while the first two equations of (57) are independent of and make up a closed nonlinear system of first-order delay ODEs for and :
where and . Integrating the last equation in Equation (57) and using relations (58), we find that the function is expressed in terms of and as
where A is an arbitrary constant.
It is noteworthy that the system of delay ODEs (59) generally admits one-component solutions , and , . In the absence of delay, these solutions can be expressed in terms of elementary functions.
2.7. The First Integral and Exact Solutions of the Reduced System of Two First-Order Delay ODEs
On eliminating the combination from the system of delay ODEs (59), we get
Integrating yields the first integral
where is an arbitrary constant, which may depend on . Substituting (61) into the first equation of system (59), we arrive at one first-order delay ODE
Example 1.
ODE (62) for the Lotka–Volterra system of PDEs without delay (3), where is a Bernoulli equation. The integration of it gives, in view of (61), an exact solution to system (59):
where is an arbitrary constant. The respective function is defined by Formula (60) with . It can be expressed in terms of elementary functions if, for example,
Formulas (14), (54), (58) and (64) describe a new solution to the Lotka–Volterra system of PDEs without delay (3).
Similarly, if and , we get
The Cauchy-type problem for the delay ODE (62) subject to the initial condition (66) can be solved using the method of steps [59,60]. To this end, we split time t into segments of length τ and denote
where and Then, on integrating Equation (62) from to t on the interval [, we get
The function on the left-hand side of Formula (68) is sought on the interval , whereas on the right-hand side is defined on the preceding interval . The computations are carried out successively starting from , when the right-hand side is a known function defined on the initial interval (66). This step results in . Then, one sets and finds the function using (68) via the already known function . The procedure continues further in a similar fashion.
For , in order to determine , , and , one has to solve the nonlinear system consisting of the last three equations in (57) with and .
3. Reductions and Exact Solutions of the System of Delay PDEs with Identical Diffusion Coefficients ()
3.1. Reduction of the System of PDEs with Three Delays to a System of Two Second-Order ODEs without Delay and One Linear PDE
On setting , , and , we look for solutions to system (3) in the form
where is a free parameter, while , , and are functions to be determined in the subsequent analysis.
Solutions of the form (69), where is a function of two independent variables and is an arbitrary parameter, generalize substantially solutions (17), in which is independent of t, and is expressed in terms of the system constants. We will refer to such solutions as incomplete separable solutions.
We assume the function to satisfy the additional periodicity condition
One can verify that relations (18) remain valid under conditions (7) and (70). Considering the above and substituting (69) into (3), we obtain a closed system for and consisting of two second-order ODEs without delay:
The function satisfies the nonstationary Schrödinger equation
and the periodic condition (70). Importantly, the coefficients of Equation (73) depend on the space variable x alone.
We multiply ODEs (71) and (72) by and , respectively, and add up to obtain an ODE for . We write this equation in conjunction with ODE (71) and PDE (70) as the following mixed algebraic-differential system of equations:
Systems (74)–(77) can be solved sequentially from one equation to another. We start from the isolated stationary ODE (74). Divided by a, it coincides, up to obvious renaming, with Equation (24). PDE (77) is subjected to the additional conditions (7) and (70), which hold true, for example, in the stationary case and the nonstationary case of and .
3.2. Exact Solutions to the Reduced System Consisting of Two ODEs and One PDE
. More complicated stationary solutions to system (74)–(77) with and can be constructed in a few steps as shown below (see Table 4).
- (i)
- The isolated subsystem consisting of two ODEs (74) and (75) for and coincides, up to notation, with the system of ODEs (24) and (25) with . It follows that exact solutions to Equations (74) and (75) can be obtained using Table 2 and formulas with , in which the functions and determining parameters must be renamed as follows: , , , , and .
- (ii)
- The function is determined by substituting the functions and obtained in step (i) into (76), which results in
- (iii)
. If and , some nonstationary exact solutions to systems (74)–(77) with are defined by Formulas (78) and (79) for and with any expression of the function specified below:
where C and are arbitrary constants. Some other nonstationary exact solutions to systems (74)–(77) with can be obtained using Formulas (78) and (79) and the expression , where is any solution of the standard linear heat equation .
. If and , there are nonstationary exact solutions to systems (74)–(77) with that are defined by Formula (81) for and and any expression of from (83) with .
. For , one can seek exact solutions to PDE (73) that satisfy the periodicity condition (70) and relation (7) in the form
where corresponds to a stationary solution. Substituting (84) into (71) yields the following linear stationary system of ODEs for and :
Since PDE (73) is linear in , an arbitrary linear combination of exact solutions (84),
where are arbitrary constants, is also an exact solution to this equation.
Example 2.
For any of the four simplest solutions (5), the general solution of Equation (73) satisfying the periodicity condition (70) can be represented as
where
The constants , , , and , otherwise arbitrary, must ensure the convergence of the series (86) and derivatives and ; the convergence is certainly ensured if, for example, we set for all , where M is an arbitrary positive integer.
Let us highlight two special cases:
3.3. Reduction of the System of PDEs with a Single Delay to a System of Two First-Order Delay ODEs and One Linear PDE
We look for incomplete separable solutions to system (3) with , , and and a common delay time (53) in the form
with the functions , , and to be determined in the subsequent analysis.
On substituting (87) into the system of PDEs (3), we arrive at a nonlinear system of delay ODEs for and ,
and a linear parabolic PDE with variable coefficients for :
The system of delay ODEs (88) coincides with system (59) at . Hence, it can be integrated as described in Section 2.7, where some exact solutions can also be found. With the substitution
Equation (89) can be reduced to the standard linear heat equation
whose exact solutions can be found, for example, in [61].
4. Related Systems of PDEs with Several Space Variables and Delays Other Generalizations
4.1. Reductions and Exact Solutions of a Multidimensional System with Different Diffusion Coefficients and Delays
Now, let us look at the following Lotka–Volterra-type nonlinear diffusion system of PDEs with n space variables:
where is the Laplace operator and are Cartesian coordinates, while the other notations are the same as in system (3). As above, we assume that relation (7) holds.
System (90) admits reductions to simpler equations, which are similar to the reductions of system (3). We will illustrate this with specific examples and, in addition, present some exact solutions to (90) as well as solutions expressible in terms of solutions to linear PDEs.
. There is an exact solution to system (90) with that generalizes solutions (15) and (16):
where , and is a function satisfying the Helmholtz equation
In the three-dimensional case, with , the linear PDE (92) admits, for example, the following exact solutions [61]:
where , , , , and are arbitrary constants. Any instance of the function sinh in the last three solutions can be replaced with cosh.
. Suppose that
Then, system (90) admits a radially symmetric exact solution of the form
where . The functions and are described by the nonautonomous system of ODEs
where , , and . Systems (96) and (97) are a generalization of systems (24) and (25).
Equation (96) with admits the exact solution
Substituting (98) into (97) and multiplying by , we arrive at the equation
which coincides, up to notation, with a special case of an ODE discussed in the book [58] (see Equation (132) with on page 535). It follows that the general solution of ODE (99) is given by
where and are arbitrary constants, and are the Bessel functions of the first and second kind, and and are the modified Bessel functions of the first and second kind. In the three-dimensional case, which corresponds to and , Formula (100) can be rewritten in terms of elementary functions.
4.2. Reductions and Exact solutions of a Multidimensional System with Identical Diffusion Coefficients and Delays
We assume that
System (90) with coefficients (101) admits exact solutions of the form
where is a free parameter, while , , and are described by the mixed algebraic-differential system of equations
with additionally adopted to satisfy the periodicity condition . System (103) generalizes systems (74)–(77).
Presented below are two simple stationary solutions to system (103) with .
. Solution with expressible in terms of solutions to two independent Helmholtz equations:
where is expressed via by the formula .
In the three-dimensional case, a few exact solutions to the linear PDEs (104) can be obtained using Formula (93) in which one must set and , respectively. For other solutions to the linear PDEs (104), see [61].
. Solution with expressible in terms of solutions to the independent Laplace and Helmholtz equations
where is expressed via by the formula . For solutions to the linear PDEs (105), see [61].
4.3. Further Generalizations and Modifications
. The nonlinear system of delay PDEs (3) can be generalized to the anisotropic case (when the diffusion coefficients depend on the spatial coordinate):
where is a given function.
As previously, the constants (5) and (6) are the simplest exact solutions of system (106). If condition (7) is satisfied, then system (106) admits generalized separable solutions of the form (8), where the function is a solution to the second-order linear ODE
Exact solutions of the linear ODE (107) for various can be found in the handbook [58]. In particular, for and , the general solutions of ODE (107) can be expressed in terms of Bessel functions or modified Bessel functions.
Under conditions (7) and (21), system (106) admits more complicated generalized separable solutions of the form (17).
. A further generalization of the system of delay PDEs (3) is
Here, is an arbitrary function of two arguments, while is an arbitrary linear differential operator with respect to the spatial variable of the form
where are some given functions and m is a positive integer.
Under conditions (7) and (21), systems (108) and (109) admit a generalized separable solution of the form (17), where the functions , , and satisfy the system of ODEs
where .
If the function is explicitly independent of x, so that , then system (110) admits particular solutions of the form
5. Brief Conclusions
We have described several reductions of a Lotka–Volterra-type nonlinear multi-parameter reaction–diffusion system with several delay times to simpler systems of ODEs with or without delay, as well as reductions to mixed systems consisting of ODEs and a single linear PDE without delay.
We have found many new exact solutions to the reaction–diffusion delay system in question, including over 50 solutions expressible in terms of elementary functions. Moreover, setting the delay times in these solutions to zero can result in many new exact solutions in terms of elementary functions to a simpler nonlinear Lotka–Volterra-type system without delays.
The exact solutions presented in this paper all involve several free constants of integration. Such solutions may be suitable for testing approximate analytical and numerical methods for solving reaction–diffusion equations as well as other nonlinear PDEs with delays.
Author Contributions
Conceptualization and methodology, A.D.P.; formal analysis and investigation, A.D.P. and A.I.Z.; construction of solutions, A.D.P. and A.I.Z.; checking solutions, A.I.Z.; writing—original draft preparation, A.D.P. and A.I.Z.; writing—review and editing, A.D.P.; supervision, A.D.P. All authors have read and agreed to the published version of the manuscript.
Funding
The study was supported by the Ministry of Education and Science of the Russian Federation, Russia within the framework of the State Assignment (Reg. No. AAAA-A20-120011690135-5 and No. AAAA-A20-120092190052-9).
Institutional Review Board Statement
Not applicable.
Informed Consent Statement
Not applicable.
Data Availability Statement
Not applicable.
Conflicts of Interest
The authors declare no conflict of interest.
References
- Huang, J.; Zou, X. Traveling wavefronts in diffusive and cooperative Lotka–Volterra system with delays. J. Math. Anal. Appl. 2002, 271, 455–466. [Google Scholar] [CrossRef]
- Li, W.-T.; Yan, X.-P.; Zhang, C.-H. Stability and Hopf bifurcation for a delayed cooperation diffusion system with Dirichlet boundary conditions. Chaos Solitons Fractals 2008, 38, 227–237. [Google Scholar] [CrossRef]
- Faria, T. Stability and bifurcation for a delayed predator–prey model and the effect of diffusion. J. Math. Anal. Appl. 2001, 254, 433–463. [Google Scholar] [CrossRef]
- Pao, C. Global asymptotic stability of Lotka–Volterra competition systems with diffusion and time delays. Nonlinear Anal. Real World Appl. 2004, 5, 91–104. [Google Scholar] [CrossRef]
- Lv, G.; Wang, M. Traveling wave front in diffusive and competitive Lotka–Volterra system with delays. Nonlinear Anal. Real World Appl. 2010, 11, 1323–1329. [Google Scholar] [CrossRef]
- Wu, J.; Zou, X. Traveling wave fronts of reaction–diffusion systems with delay. J. Dyn. Differ. Equ. 2001, 13, 651–687. [Google Scholar] [CrossRef]
- Lotka, A.J. Undamped oscillations derived from the law of mass action. J. Am. Chem. Soc. 1920, 42, 1595–1599. [Google Scholar] [CrossRef]
- Volterra, V. Variazionie fluttuazioni del numero d’individui in specie animali conviventi. Mem. Acad. Lincei 1926, 2, 31–113. [Google Scholar]
- May, R.M. Time delay versus stability in population models with two and three trophic levels. Ecology 1973, 4, 315–325. [Google Scholar] [CrossRef]
- Gopalsamy, K. Stability and Oscillations in Delay Differential Equations of Population Dynamics; Kluwer: Dordrecht, The Netherlands, 1992. [Google Scholar]
- Kuang, Y. Delay Differential Equations with Applications in Population Dynamics; Academic Press: Boston, MA, USA, 1993. [Google Scholar]
- Song, Y.; Wei, J. Local Hopf bifurcation and global periodic solutions in a delayed predator-prey system. J. Math. Anal. Appl. 2005, 301, 1–21. [Google Scholar] [CrossRef]
- Nakaoka, S.; Saito, Y.; Takeuchi, Y. Stability, delay, and chaotic behavior in a Lotka–Volterra predator-prey system. Math. Biosci. Eng. 2006, 3, 173–187. [Google Scholar] [CrossRef] [PubMed]
- Xu, C.J.; Tang, X.H.; Liao, M.X.; He, X.F. Bifurcation analysis in a delayed Lokta–Volterra predator-prey model with two delays. Nonlinear Dyn. 2011, 66, 169–183. [Google Scholar] [CrossRef]
- Conway, E.D.; Smoller, J.A. Diffusion and the predator-prey interaction. SIAM J. Appl. Math. 1977, 33, 673–686. [Google Scholar] [CrossRef]
- Jorné, J.; Carmi, S. Liapunov stability of the diffusive Lotka–Volterra equations. Math. Biosci. 1977, 37, 51–61. [Google Scholar] [CrossRef]
- Hastings, A. Global stability in Lotka–Volterra systems with diffusion. J. Math. Biol. 1978, 6, 163–168. [Google Scholar] [CrossRef]
- Alhasanat, A.; Ou, C. Minimal-speed selection of traveling waves to the Lotka–Volterra competition model. J. Differ. Equ. 2019, 266, 7357–7378. [Google Scholar] [CrossRef]
- Lam, K.Y.; Salako, R.B.; Wu, Q. Entire solutions of diffusive Lotka–Volterra system. J. Differ. Equ. 2020, 269, 10758–10791. [Google Scholar] [CrossRef]
- Rodrigo, M.; Mimura, M. Exact solutions of a competition-diffusion system. Hiroshima Math. J. 2000, 30, 257–270. [Google Scholar] [CrossRef]
- Hung, L.-C. Exact traveling wave solutions for diffusive Lotka–Volterra systems of two competing species. Jpn. J. Ind. Appl. Math. 2012, 29, 237–251. [Google Scholar] [CrossRef]
- Kudryashov, N.A.; Zakharchenko, A.S. Analytical properties and exact solutions of the Lotka–Volterra competition system. Appl. Math. Comput. 2015, 254, 219–228. [Google Scholar] [CrossRef][Green Version]
- Cherniha, R.M. Conditional symmetries for systems of PDEs: New definitions and their application for reaction–diffusion systems. J. Phys. A Math. Theor. 2010, 43, 405207. [Google Scholar] [CrossRef]
- Cherniha, R.; Davydovych, V. Nonlinear Reaction-Diffusion Systems—Conditional Symmetry, Exact Solutions and Their Applications in Biology; Springer: Cham, Switzerland, 2017. [Google Scholar]
- Cherniha, R.; Davydovych, V. New conditional symmetries and exact solutions of the diffusive two-component Lotka–Volterra system. Mathematics 2021, 9, 1984. [Google Scholar] [CrossRef]
- Polyanin, A.D.; Zaitsev, V.F. Handbook of Nonlinear Partial Differential Equations, 2nd ed.; CRC Press: London, UK; Boca Raton, FL, USA, 2012. [Google Scholar]
- Pliukhin, O. Q-conditional symmetries and exact solutions of nonlinear reaction–diffusion systems. Symmetry 2015, 7, 1841–1855. [Google Scholar] [CrossRef]
- Dorodnitsyn, V.A. On invariant solutions of the nonlinear heat equation with a source. USSR Comput. Math. Math. Phys. 1982, 22, 115–122. [Google Scholar] [CrossRef]
- Kudryashov, N.A. On exact solutions of families of Fisher equations. Theor. Math. Phys. 1993, 94, 211–218. [Google Scholar] [CrossRef]
- Clarkson, P.A.; Mansfield, E.L. Symmetry reductions and exact solutions of a class of nonlinear heat equations. Physica D 1994, 70, 250–288. [Google Scholar] [CrossRef]
- Galaktionov, V.A. Quasilinear heat equations with first-order sign-invariants and new explicit solutions. Nonlinear Anal. Theor. Meth. Appl. 1994, 23, 1595–1621. [Google Scholar] [CrossRef]
- Ibragimov, N.H. (Ed.) CRC Handbook of Lie Group Analysis of Differential Equations. Symmetries, Exact solutions and Conservation Laws; CRC Press: Boca Raton, FL, USA, 1994; Volume 1. [Google Scholar]
- Doyle, P.W.; Vassiliou, P.J. Separation of variables for the 1-dimensional non-linear diffusion equation. Int. J. Non-Linear Mech. 1998, 33, 315–326. [Google Scholar] [CrossRef]
- Hood, S. On direct, implicit reductions of a nonlinear diffusion equation with an arbitrary function—Generalizations of Clarkson’s and Kruskal’s method. IMA J. Appl. Math. 2000, 64, 223–244. [Google Scholar] [CrossRef]
- Estevez, P.G.; Qu, C.; Zhang, S. Separation of variables of a generalized porous medium equation with nonlinear source. J. Math. Anal. Appl. 2002 275, 44–59. [CrossRef]
- Kaptsov, O.V.; Verevkin, I.V. Differential constraints and exact solutions of nonlinear diffusion equations. J. Phys. A Math. Gen. 2003, 36, 1401–1414. [Google Scholar] [CrossRef]
- Galaktionov, V.A.; Svirshchevskii, S.R. Exact Solutions and Invariant Subspaces of Nonlinear Partial Differential Equations in Mechanics and Physics; Chapman & Hall: Atlanta, GA, USA; CRC Press: Boca Raton, FL, USA, 2007. [Google Scholar]
- Vaneeva, O.O.; Johnpillai, A.G.; Popovych, R.O.; Sophocleous, C. Extended group analysis of variable coefficient reaction–diffusion equations with power nonlinearities. J. Math. Anal. Appl. 2007, 330, 1363–1386. [Google Scholar] [CrossRef]
- Vaneeva, O.O.; Popovych, R.O.; Sophocleous, C. Extended group analysis of variable coefficient reaction–diffusion equations with exponential nonlinearities. J. Math. Anal. Appl. 2012, 396, 225–242. [Google Scholar] [CrossRef]
- Cherniha, R.; Serov, M.; Pliukhin, O. Nonlinear Reaction-Diffusion-Convection Equations: Lie and Conditional Symmetry, Exact Solutions and Their Applications; CRC Press: Boca Raton, FL, USA, 2018. [Google Scholar]
- Polyanin, A.D. Construction of exact solutions in implicit form for PDEs: New functional separable solutions of non-linear reaction–diffusion equations with variable coefficients. Int. J. Non-Linear Mech. 2019, 111, 95–105. [Google Scholar] [CrossRef]
- Polyanin, A.D. Functional separable solutions of nonlinear reaction–diffusion equations with variable coefficients. Appl. Math. Comput. 2019, 347, 282–292. [Google Scholar] [CrossRef]
- Polyanin, A.D.; Zhurov, A.I. Separation of variables in PDEs using nonlinear transformations: Applications to reaction–diffusion-type equations. Appl. Math. Lett. 2020, 100, 106055. [Google Scholar] [CrossRef]
- Polyanin, A.D.; Zhurov, A.I. Separation of Variables and Exact Solutions to Nonlinear PDEs; CRC Press: Boca Raton, FL, USA, 2021. [Google Scholar]
- Fang, J.; Wu, J. Monotone traveling waves for delayed Lotka–Volterra competition systems. Discret. Contin. Dyn. Syst. 2012, 32, 3043–3058. [Google Scholar] [CrossRef]
- Polyanin, A.D.; Sorokin, V.G. Reductions and exact solutions of Lotka–Volterra and more complex reaction–diffusion systems with delays. Appl. Math. Lett. 2022, 125, 107731. [Google Scholar] [CrossRef]
- Meleshko, S.V.; Moyo, S. On the complete group classification of the reaction–diffusion equation with a delay. J. Math. Anal. Appl. 2008, 338, 448–466. [Google Scholar] [CrossRef]
- Polyanin, A.D.; Zhurov, A.I. Functional constraints method for constructing exact solutions to delay reaction–diffusion equations and more complex nonlinear equations. Commun. Nonlinear Sci. Numer. Simul. 2014, 19, 417–430. [Google Scholar] [CrossRef]
- Polyanin, A.D.; Zhurov, A.I. New generalized and functional separable solutions to nonlinear delay reaction–diffusion equations. Int. J. Non-Linear Mech. 2014, 59, 16–22. [Google Scholar] [CrossRef]
- Polyanin, A.D.; Sorokin, V.G. A method for constructing exact solutions of nonlinear delay PDEs. J. Math. Anal. Appl. 2021, 494, 124619. [Google Scholar] [CrossRef]
- Polyanin, A.D.; Sorokin, V.G. Construction of exact solutions to nonlinear PDEs with delay using solutions of simpler PDEs without delay. Commun. Nonlinear Sci. Numer. Simul. 2021, 95, 105634. [Google Scholar] [CrossRef]
- Polyanin, A.D.; Sorokin, V.G. Nonlinear pantograph-type diffusion PDEs: Exact solutions and the principle of analogy. Mathematics 2021, 9, 511. [Google Scholar] [CrossRef]
- Aibinu, M.O.; Thakur, S.C.; Moyo, S. Exact solutions of nonlinear delay reaction–diffusion equations with variable coefficients. Partial Differ. Equ. Appl. Math. 2021, 4, 100170. [Google Scholar] [CrossRef]
- Prakash, P.; Choudhary, S.; Daftardar-Gejji, V. Exact solutions of generalized nonlinear time-fractional reaction–diffusion equations with time delay. Eur. Phys. J. Plus 2020, 135, 490. [Google Scholar] [CrossRef]
- Polyanin, A.D.; Zhurov, A.I. Generalized and functional separable solutions to nonlinear delay Klein–Gordon equations. Commun. Nonlinear Sci. Numer. Simul. 2014, 19, 2676–2689. [Google Scholar] [CrossRef]
- Long, F.-S.; Meleshko, S.V. On the complete group classification of the one-dimensional nonlinear Klein–Gordon equation with a delay. Math. Methods Appl. Sci. 2016, 39, 3255–3270. [Google Scholar] [CrossRef]
- Bateman, H.; Erdélyi, A. Higher Transcendental Functions; McGraw-Hill: New York, NY, USA, 1955; Volume 3. [Google Scholar]
- Polyanin, A.D.; Zaitsev, V.F. Handbook of Ordinary Differential Equations: Exact Solutions, Methods, and Problems; Chapman & Hall: Atlanta, GA, USA; CRC Press: Boca Raton, FL, USA, 2018. [Google Scholar]
- Bellman, R.; Cooke, K.L. Differential-Difference Equations; Academic Press: New York, NY, USA, 1963. [Google Scholar]
- Elsgolt’s, L.E.; Norkin, S.B. Introduction to the Theory and Application of Differential Equations with Deviating Arguments; Academic Press: New York, NY, USA, 1973. [Google Scholar]
- Polyanin, A.D.; Nazaikinskii, V.E. Handbook of Linear Partial Differential Equations for Engineers and Scientists, 2nd ed.; Chapman & Hall: London, UK; CRC Press: Boca Raton, FL, USA, 2016. [Google Scholar]
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