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Article

Existence, Uniqueness and Solutions for Diffusion and Advection Effects for Predator–Prey Model with Holling Type II Interaction Function

by
Saeed Ur Rahman
1,
José Luis Díaz Palencia
2,* and
Maria Rehman
1
1
Department of Mathematics, COMSATS University Islamabad, Abbottabad Campus, Abbottabad 22060, Pakistan
2
Department of Mathematics and Education, Universidad a Distancia de Madrid, Via de Servicio A-6, 15, 28400 Madrid, Spain
*
Author to whom correspondence should be addressed.
Mathematics 2026, 14(5), 831; https://doi.org/10.3390/math14050831
Submission received: 22 November 2025 / Revised: 14 February 2026 / Accepted: 17 February 2026 / Published: 28 February 2026
(This article belongs to the Special Issue Dynamical Systems & Partial Differential Equations)

Abstract

The present work is focused on a predator–prey model with the Holling type II interaction function, which is influenced by diffusion, advection and nonlinear reaction effects. Firstly, we show that the solutions of this dynamical model are bounded and unique. Secondly we use the Lyapunov function and then show that the equilibrium points are globally stable. Thirdly, we obtain the solution profile when the diffusion coefficient is small. For this purpose we introduce self-similar structures to convert the nonlinear partial differential equations into nonlinear ordinary differential equations and then use the singular perturbation technique to solve these equations. Fourthly, we use the Hamiltonian and Lighthill’s technique to obtain upper stationary solutions for a small coefficient of the advection term. Lastly, we consider a large diffusion coefficient and obtain the asymptotic profiles of nonstationary solutions with the help of nonlinear point scaling.

1. Introduction

The study of diffusion and advection effects in predator–prey models with Holling type II interaction functions has received significant attention from both theoretical and practical perspectives. The foundational work of Lotka [1] and Volterra [2] introduced key mathematical models to inspect predator–prey dynamics. Many other researchers have worked on these models based on trophic interactions, such as explaining the construction of predator–prey reaction–diffusion systems in [3], supplementary food sources in [4], prey refuge in [5], hunting cooperation in [6], the fear effect in [7], the Allee effect in [8], refuge and harvesting in [9,10], and dissolved oxygen–plankton in [11]. Although such models often reply on simplifying assumptions, considerable efforts have been devoted to validating their relevance in real-world contexts, particularly in assessing whether predators can regulate prey populations. Prominent frameworks like Holling’s type II functional response [12] and the Lotka–Volterra model [13] have been central in these investigations. Berryman expanded upon predation models using Holling’s type II response but did not consider the prey refuge—an ecologically important factor. This issue was later resolved by Kar [14].
The interaction between predator and prey populations in ecological systems can be principally designed in two ways: (1) one predator and multiple prey species or (2) one predator and one prey species [15]. Erbach et al. [16] examined a generalized predator–prey model, while Magal et al. [17] extended it by infusing it with historical prey population effects. Although the authors in the above three references deeply investigated trophic dynamics involving specialized or generalized predator–prey models, they did not consider prey refuges. Sen et al. [18] and Mondal et al. [19] are known as pioneers in investigating the combined influence of prey refuge and Beverton–Holt-like supplementary food on the prey and predator. The proposed model precisely analyzes stability transitions and other dynamical behaviors, incorporating an Allee effect in prey growth and cooperative hunting among predators. In addition to the Holling type II framework, providing alternative food sources for predators when prey seek refuge emerges as a biologically rational strategy. The model reveals that predators consume fewer prey relative to their population, granting prey greater potential to find protection. One possible approach is a predator–prey model where the prey manifests logistic growth. Furthermore, the predator’s food supply can be modeled using a Beverton–Holt-like functional form to account for alternative resources. As a result, one gets
u t = a 1 u 1 u K ψ ( u , v ) v v t = e ψ ( u , v ) v a 4 v + a 5 v 1 + a 6 v ,
where u ( 0 ) 0 , v ( 0 ) 0 . Here u and v denote densities of prey and predator species respectively at any time t, e  ( 0 < e < 1 ) signifies the conversion factor from prey into predator density, a 4 implies the natural mortality rate of the predator, a 1 is the logistic growth coefficient of the prey species and K is the prey species holding capacity. The predator species propagates with a Beverton–Holt-like function (adding extra food sources to the predator) of the form a 5 v 1 + a 6 v when the primary prey is absent. The predator’s maximum rate of reproduction is a 5 , which drops with density, and a 6 represents the resilience of biomass dependence. We steadily presume a 5 > 4 in order to permit the predator species to persist even in the absence of the primary prey. Ecologically, this imposed condition a 5 > 4 represents the minimum threshold for food that is required for the predator to survive and reproduce. If natural prey is insufficient, then additional food is required by the predator for their basic energy needs. When a 5 is too small, then the supplementary food cannot compensate for prey shortage and this causes the predator to decline. Therefore a 5 > 4 gives a guarantee that the supply of supplementary food is enough to sustain the predator population. The interaction function, represented as ψ ( u , v ) , specifies how the predator eats the prey species. A generalist predator–prey model is used when a 5 > a 4 . The limitation a 5 > a 4 must be secured throughout the inquiry in order to fully understand the complexity of system dynamics brought about by the Beverton–Holt-like additional food. The response function ψ ( u , v ) for predation is here concluded to be the Holling type II interaction function, with a 2 representing the maximum number of prey that the predator may eat in a given amount of time and a 3 representing the half-saturation constant. A continuous percentage of the prey refuge, m u , from the prey species is incorporated into the interacting species model (1): 0 m u < u ; m is a positive constant. Across the study, it is assumed that 0 m < 1 ; that is, for a fundamental biological system, the acceptable range of prey refuge is 0 < ( 1 m ) 1 . If m = 0 then there is no refuge and all prey are exposed to the predator; for 0 < m < 1 then some prey are protected and some are not, which is close to a real ecosystem; and if m = 1 then all prey are protected and the predator lives without food, which is unrealistic. Consequently, the model assumes the following form when this refuge phenomenon is included in the response function ψ ( u , v ) , i.e., ψ ( u , v ) = a 2 ( 1 m ) u a 3 + ( 1 m ) u . As a result, model (1) becomes (2) with the addition of the Holling type II interaction function and constant-proportion prey refuge.
u t = a 1 u 1 u K a 2 ( 1 m ) u v a 3 + ( 1 m ) u v t = e a 2 ( 1 m ) u v a 3 + ( 1 m ) u a 4 v + a 5 v 1 + a 6 v .
The addition of advection and diffusion terms in the predator–prey model remarkably enhances the consideration of the system’s dynamic. Due to environment factors or other factors, the prey and predator move directly, which is represented by the advection term. The diffusion term represents the random movement of the prey and predator. This addition of diffusion and advection terms is a modification of the previous model and obtains the conditions in which different type of solutions exist. Now we assume that the prey refuge model depends on time as well as space; then after including the diffusion and advection effects in the above-mentioned model, we have
u t = D 1 2 u x 2 + ϵ ¯ 1 u x + a 1 u 1 u K a 2 ( 1 m ) u v a 3 + ( 1 m ) u v t = D 2 2 v x 2 + ϵ ¯ 1 v x + e a 2 ( 1 m ) u v a 3 + ( 1 m ) u a 4 v + a 5 v 1 + a 6 v ,
where D 1 and D 2 are the coefficients of diffusion and ϵ ¯ 1 is the coefficient of advection, with u ( x , 0 ) = u 0 ( x ) , v ( x , 0 ) = v 0 ( x ) , u ( 0 , t ) = 0 , u ( l π , t ) = 0 , v ( 0 , t ) = 0 and v ( l π , t ) = 0 . In order to reduce the number of system parameters, the model mentioned above must be non-dimensionalized by adding u ˜ = u K , v ˜ = e v K , x ˜ = x l and t ˜ = a 1 t . After using this approach, the redesigned model looks like this (tildes dropped):
u t = d 1 2 u x 2 + ϵ 1 u x + u ( 1 u ) α ( 1 m ) u v β + ( 1 m ) u v t = d 2 2 v x 2 + ϵ 1 v x + γ ( 1 m ) u v β + ( 1 m ) u δ 1 v + ϵ v 1 + ρ v ,
where u ( x , 0 ) = u 0 ( x ) , v ( x , 0 ) = v 0 ( x ) , u ( 0 , t ) = 0 , u ( π , t ) = 0 , v ( 0 , t ) = 0 and v ( π , t ) = 0 with d 1 = D 1 2 a 1 l 2 , d 2 = D 2 2 a 1 l 2 , ϵ 1 = ϵ ¯ 1 a 1 l α = a 2 e a 1 , β = a 3 K , γ = e a 2 a 1 , δ 1 = a 4 a 1 , ϵ = a 5 a 1 , ρ = a 6 K e .
In our analysis, we take ε = 1 for simplicity, which does not affect the generality. In this paper we consider a predator–prey model with the Holling type II interaction function, under diffusion, advection and reaction effects. The governing equations of this model are nonlinear partial differential equations. In Section Two, we show that the solutions of these partial differential equations are bounded and unique. In Section Three, we consider the Lyapunov function and then show that the equilibrium points are globally stable. In Section Four, we develop the solutions for a large domain with a small coefficient of diffusion. In Section Five, we take a small coefficient of the advection term and apply the Hamiltonian and Lighthill’s technique for upper stationary solutions. In Section Six, we take a large diffusion coefficient and use the nonlinear point scaling technique to obtain the asymptotic nonstationary solutions.

2. Existence and Uniqueness

To prove the existence, we consider the following theorem.
Theorem 1.
Given that u 0 ( x ) , v 0 ( x ) L 1 ( R ) L ( R ) , then the solutions u ( x , t ) and v ( x , t ) are bounded on B γ × [ 0 , T ] with 0 < γ < , where B γ means the ball of radius γ.
Proof. 
To start with, consider the test function ξ 2 C ( Q T ) , where Q T = R × [ τ , T ] with 0 < τ < t < T . Also consider the cut-off function η ϕ 2 , with a constant ϕ 2 0 + and γ γ 0 > 0 , where γ refers to a spatial dimension, and then a cut = off function defined as follows (refer to [20] for additional insights):
η ϕ 2 C 0 ( x , t ) , 0 η ϕ 1 1 , η ϕ 2 = 1 i n B γ ϕ 2 , η ϕ 2 = 0 i n R 2 B γ ϕ 2 ,
so that
η ϕ 2 a 1 ϕ 2 , Δ η ϕ 1 a 1 ϕ 2 2 .
Multiplying the first equation of (4) by ξ 2 η ϕ 2 , integrating over B γ × [ τ , T ] , and assuming u and u x are zero at the boundary, we get
B γ u ( x , t ) ξ 2 ( x , t ) η ϕ 2 d x B γ u ( x , τ ) ξ 2 ( x , τ ) η ϕ 2 d x τ t B γ u ξ 2 s η ϕ 2 + u ξ 2 η ϕ 2 s d x d s   = τ t B γ [ d 1 u 2 ξ 2 x 2 η ϕ 2 + 2 d 1 u ξ 2 x η ϕ 2 x + d 1 u ξ 2 2 η ϕ 2 x 2 ϵ 1 u ξ 2 x η ϕ 2   ϵ 1 u ξ 2 η ϕ 2 x + u ( 1 u ) ξ 2 η ϕ 2 α ( 1 m ) u v β + ( 1 m ) u ξ 2 η ϕ 2 ] d x d s ,
which implies that
B γ u ( x , t ) ξ 2 ( x , t ) η ϕ 2 d x = B γ u ( x , τ ) ξ 2 ( x , τ ) η ϕ 2 d x + τ t B γ [ u ξ 2 s η ϕ 2 d x + u ξ 2 η ϕ 2 s + d 1 u 2 ξ 2 x 2 η ϕ 2   + 2 d 1 u ξ 2 x η ϕ 2 x + d 1 u ξ 2 2 η ϕ 2 x 2 ϵ 1 u ξ 2 x η ϕ 2 ϵ 1 u ξ 2 η ϕ 2 x + u ( 1 u ) ξ 2 η ϕ 2   α ( 1 m ) u v β + ( 1 m ) u ξ 2 η ϕ 2 ] d x d s .
Similarly multiplying the second equation of (4) by ξ 2 η ϕ 2 , integrating over B γ × [ τ , T ] , and assuming v and v x are zero at the boundary, we get
B γ v ( x , t ) ξ 2 ( x , t ) η ϕ 2 d x = B γ v ( x , τ ) ξ 2 ( x , τ ) η ϕ 2 d x + τ t B γ [ v ξ 2 s η ϕ 2 d x + v ξ 2 η ϕ 2 s + d 2 v 2 ξ 2 x 2 η ϕ 2   + 2 d 2 v ξ 2 x η ϕ 2 x + d 2 v ξ 2 2 η ϕ 2 x 2 ϵ 1 v ξ 2 x η ϕ 2 ϵ 1 v ξ 2 η ϕ 2 x + γ ( 1 m ) u v β + ( 1 m ) u ξ 2 η ϕ 2   δ 2 v ξ 2 η ϕ 2 + v 1 + ρ v ξ 2 η ϕ 2 ] d x d s .
For sufficiently large γ γ 0 > 1 , from [21] the following results hold:
τ t u d s c p τ x 2 m 1 , τ t u x d s 2 m 1 c p τ x 3 m m 1 ,
and
τ t v d s c q τ x 2 m 1 , τ t v x d s 2 m 1 c q τ x 3 m m 1 ,
where c p ( τ ) and c q ( τ ) are constants depending on fixed time τ . Admitting τ t , the defined functions ξ 2 and η ϕ 1 exhibit local stationarity. Consequently, we have
τ t B γ u ξ 2 s η ϕ 2 d x d s B γ c p τ x 2 m 1 ξ 2 s η ϕ 2 d x τ t B γ u ξ 2 η ϕ 2 s d x d s B γ c p τ ξ 2 x 2 m 1 η ϕ 2 s d x d 1 τ t B γ u 2 ξ 2 x 2 η ϕ 1 d x d s d 1 τ t B γ u 2 ξ 2 x 2 η ϕ 1 d x d s   d 1 B γ c p τ x 2 m 1 2 ξ 2 x 2 η ϕ 1 d x 2 d 1 τ t B γ u ξ 2 x η ϕ 2 x d x d s 2 d 1 B γ c p τ x 2 m m 1 ξ 2 x η ϕ 2 x d x d 1 τ t B γ u ξ 2 2 η ϕ 2 x 2 d x d s d 1 B γ c p τ x 2 m m 1 ξ 2 2 η ϕ 2 x 2 d x ϵ 1 τ t B γ u ξ 2 x η ϕ 2 d x d s ϵ 1 B γ c p τ x 2 m 1 ξ 2 x η ϕ 2 d x ϵ 1 τ t B γ u ξ 2 η ϕ 2 x d x d s ϵ 1 B γ c p τ x 2 m 1 ξ 2 η ϕ 2 x d x ,
and
τ t B γ v ξ 1 s η ϕ 2 d x d s B γ c q τ x 2 m 1 ξ 2 s η ϕ 2 d x τ t B γ v ξ 2 η ϕ 2 s d x d s B γ c q τ ξ 2 x 2 m 1 η ϕ 2 s d x d 2 τ t B γ v 2 ξ 2 x 2 η ϕ 2 d x d s d 2 τ t B γ v 2 ξ 2 x 2 η ϕ 2 d x d s   d 2 B γ c q τ x 2 m 1 2 ξ 2 x 2 η ϕ 2 d x 2 d 2 τ t B γ v ξ 2 x η ϕ 2 x d x d s 2 d 2 B γ c q τ x 2 m m 1 ξ 2 x η ϕ 2 x d x d 2 τ t B γ v ξ 2 2 η ϕ 2 x 2 d x d s d 2 B γ c q τ x 2 m m 1 ξ 2 2 η ϕ 2 x 2 d x ϵ 1 τ t B γ v ξ 2 x η ϕ 1 d x d s ϵ 1 B γ c q τ x 2 m 1 ξ 2 x η ϕ 1 d x ϵ 1 τ t B γ v ξ 2 η ϕ 2 x d x d s ϵ 1 B γ c q τ x 2 m 1 ξ 2 η ϕ 2 x d x .
For a fixed γ , the following holds:
B γ d 1 c p τ x 2 m 1 2 ξ 2 x 2 η ϕ 1 d x = d 1 c p ( τ ) x 2 m 1 ξ 2 x η ϕ 1 B γ d 1 c p ( τ ) B γ 2 m 1 x 2 m 1 1 ξ 2 x η ϕ 1 d x   d 1 c p ( τ ) B γ x 2 m 1 ξ 2 x η ϕ 1 x d x .
Note that, when γ 1 , then ( ξ 2 x η ϕ 1 ) B γ 1 . Now, considering (5) and (6), from (13) we have
B γ d 1 c p τ x 2 m 1 2 ξ 2 x 2 η ϕ 1 d x 2 m 1 B γ d 1 c p τ x 3 m m 1 ξ 2 x d x + B γ d 1 c p τ x 2 m 1 ξ 2 x a 1 ϕ 1 d x 2 m 1 d 1 c p τ B γ x 3 m m 1 ξ 2 x d x + d 1 c p τ a 1 B γ x 2 m 1 1 ξ 2 x d x .
Now considering (11) and using (5) and (6), the following assessments hold:
ϵ 1 B γ c p τ x 2 m 1 ξ 2 x η ϕ 1 d x ϵ 1 c p τ B γ x 2 m 1 ξ 2 x d x ;
B γ c p τ x 2 m m 1 ξ 2 2 η ϕ 2 x 2 d x c p τ B γ x 2 m m 1 a 1 ξ 2 ϕ 2 2 d x   a 1 c p τ B γ x 2 m m 1 2 ξ 2 d x ;
2 B γ c p τ x 2 m m 1 ξ 2 x η ϕ 2 x d x 2 c p τ B γ x 2 m m 1 ξ 2 x a 1 ϕ 2 d x   2 a 1 c p τ B γ x 2 m m 1 1 ξ 2 x d x ;
ϵ 1 B γ c p τ x 2 m 1 ξ 2 η ϕ 2 x d x ϵ 1 B γ c p τ x 2 m 1 ξ 2 ξ 2 x a 1 ϕ 2 d x   ϵ 1 B γ c p τ x 2 m 1 1 ξ 2 ξ 2 x d x .
Next, consider the following test function:
ξ 2 x , s = e g s 1 + x 2 μ ,
where g ( s ) < 0 and μ > 0 are constants specified in such a way that the integrals in (11) to (18) converge as γ . Differentiating (19) with regards to x , we get
ξ 2 x = 2 μ e g ( s ) ( 1 + x 2 ) μ + 1 x   2 μ e g ( s ) x 2 μ 1 .
After using (20), the integral in (11) can be obtained as follows:
d 1 B γ c p τ x 2 m 1 2 ξ 2 x 2 η ϕ 1 d x 4 d 1 μ m 1 c p τ B γ e g ( s ) x 3 m m 1 2 μ 1 d x   + 2 μ a 1 d 1 c p τ B γ e g ( s ) x 2 m 1 2 μ 2 d x .
The above integral is converges if 2 m 1 2 μ 2 < 1 , which implies that
μ > 3 m 2 ( m 1 ) .
Similarly, after using (20), the integral in (15) is solved as follows:
ϵ 1 B γ c p τ x 2 m 1 ξ 2 x η ϕ 1 d x 2 μ ϵ 1 c p τ B γ e g ( s ) x 2 m 1 2 μ 1 d x .
For the convergence of the above integral, we take 2 m 1 2 μ 1 < 1 and we get
μ > 1 2 ( m 1 ) .
Also, consider
τ t B γ u ( 1 u ) ξ 2 η ϕ 2 d x d s τ t B γ u ( 1 u ) ξ 2 η ϕ 2 d x   τ t B γ u ξ 2 η ϕ 2 d x   B γ c p τ x 2 m 1 ξ 2 η ϕ 2 d x   B γ c p τ e g ( s ) x 2 m 1 2 μ d x .
The convergence is achieved if we take 2 m 1 2 μ < 1 ; therefore we obtain
μ > 1 + m 2 ( m 1 ) .
After using (19), the integral in (11) can be solved as follows:
τ t B γ u ξ 2 s η ϕ 2 d x d s B γ c p τ x 2 m 1 e g ( s ) g ( s ) 1 ( 1 + x 2 ) μ d x   B γ c p τ x 2 m 1 e g ( s ) g ( s ) x 2 μ d x   = c p τ B γ e g ( s ) g ( s ) x 2 m 1 2 μ d x .
The integral converges if we take the same value of μ as that given in (26). After using (19) the integral in (11) can be solved as follows:
τ t B γ u ξ 2 η ϕ 2 s d x d s B γ c p τ x 2 m 1 e g ( s ) ( 1 + x 2 ) μ η ϕ 2 s d x   c p τ B γ x 2 m 1 e g ( s ) x 2 μ η ϕ 2 s d x   = c p τ B γ e g ( s ) x 2 m 1 2 μ η ϕ 2 s d x .
Similarly the integral converges if we take the same value of μ as that given in (26). After using (20) the integral in (18) can be solved as follows:
ϵ 1 B γ c p τ x 2 m 1 1 ξ 2 d x ϵ 1 B γ c p τ x 2 m 1 1 2 μ d x .
The integral converges if we take the same value of μ as that given in (24). After using (20) the integral in (17) can be solved as follows:
2 a 1 c p τ B γ x 2 m m 1 1 ξ 2 x d x 2 a 1 c p τ B γ x 2 m m 1 2 μ 2 d x .
The integral converges if we take the same value of μ as that given in (26). Also
τ t B γ α ( 1 m ) u v β + ( 1 m ) u ξ 2 η ϕ 2 d x d s τ t B γ α ( 1 m ) u v ( 1 m ) u ξ 2 η ϕ 2 d x d s   τ t B γ α v ξ 2 η ϕ 2 d x d s   B γ α c q τ x 2 m 1 ξ 2 η ϕ 2 d x   α c q B γ e g ( s ) x 2 m 1 2 μ d x .
The integral converges if we take the same value of μ as that given in (26), and similarly for v. Note that, when γ 1 , then ( ξ 2 x η ϕ 2 ) B γ 1 . Now, using (5), (6), (19) and (20), and after solving the integral in (12), we get
τ t B γ v ξ 2 s η ϕ 2 d x d s c q τ B γ e g ( s ) g ( s ) x 2 m 1 2 μ d x .
The integral converges if we take the same value of μ as that given in (26).
τ t B γ v ξ 2 η ϕ 2 s d x d s c q τ B γ e g ( s ) x 2 m 1 2 μ η ϕ 2 s d x .
The integral converges if we take the same value of μ as that given in (26).
d 2 τ t B γ v 2 ξ 2 x 2 η ϕ 1 d x d s 4 μ d 2 m 1 c q τ B γ e g ( s ) x 3 m m 1 2 μ 1 d x   + 2 μ a 1 d 2 c q τ B γ e g ( s ) x 2 m 1 2 μ 2 d x .
The integral converges if we take the same value of μ as that given in (22).
2 a 1 c q τ B γ x 2 m m 1 1 ξ 2 x d x 2 a 1 c q τ B γ e g ( s ) x 2 m m 1 2 μ 2 d x .
The integral converges if we take the same value of μ as that given in (26).
ϵ 1 τ t B γ v ξ 2 η ϕ 2 x d x d s ϵ 1 a 1 B γ c q τ e g ( s ) x 2 m 1 1 2 μ d x .
The integral converges if we take the same value of μ as that given in (24).
ϵ 1 τ t B γ v ξ 2 x η ϕ 2 d x d s 2 μ ϵ 1 c q τ B γ e g ( s ) x 2 m 1 2 μ 1 d x .
The integral converges if we take the same value of μ as that given in (24). Also, consider
τ t B γ δ 1 v ξ 2 η ϕ 2 d x d s τ t B γ δ 1 v ξ 2 η ϕ 2 d x d s   δ 1 c q τ B γ e g ( s ) x 2 m 1 2 μ d x .
The integral converges if we take the same value of μ as that given in (26).
τ t B γ γ ( 1 m ) u v β + ( 1 m ) u ξ 2 η ϕ 2 d x d s τ t B γ γ ( 1 m ) u v ( 1 m ) u ξ 2 η ϕ 2 d x d s   τ t B γ γ v ξ 2 η ϕ 2 d x d s   B γ γ c q τ x 2 m 1 ξ 2 η ϕ 2 d x   γ τ c q B γ e g ( s ) x 2 m 1 2 μ d x .
The integral converges if we take the same value of μ as that given in (26). Now consider
τ t B γ v 1 + ρ v ξ 2 η ϕ 2 d x d s τ t B γ v 1 + ρ v ξ 2 η ϕ 2 d x d s   τ t B γ v ρ v ξ 2 η ϕ 2 d x d s   1 ρ B γ e g ( s ) x 2 μ d x .
The integral is converges if μ > 1 2 . So for a suitable value of μ , the expressions (7) and (8) become
τ t B γ u ξ 2 η ϕ 2 d x d s A 1 ,
and
τ t B γ v ξ 2 η ϕ 2 d x d s A 2 .
Note that when B γ , then we can choose sufficiently small A 1 and A 2 . Since ξ 2 and η ϕ 2 are bounded, this allows us to establish the boundedness of solutions u and v in B γ × [ 0 , T ] . □
In the next section, we show that the solution of the system is unique. Let ( u 1 , v 1 ) and ( u 2 , v 2 ) be the solutions of (4); then we show that u 1 = u 2 and v 1 = v 2 . We introduce this result in the form of a theorem and we introduce the definitions of upper and lower solutions therein.
Theorem 2.
Assume u 1 > 0 and v 2 > 0 are upper solutions of (4). Then both u 1 and v 1 coincide with the lower solutions u 2 > 0 and v 2 > 0 , implying that the solutions are unique.
Proof. 
Assume u 1 , v 1 are the upper solutions of (4) in R × 0 , T ; then such solutions are defined in accordance with the following problem:
u 1 t = d 1 2 u 1 x 2 + ϵ 1 u 1 x + u 1 u 1 2 α ( 1 m ) u 1 v 1 β + ( 1 m ) u 1 ,
and
v 1 t = d 2 2 v 1 x 2 + ϵ 1 v 1 x + γ ( 1 m ) u 1 v 1 β + ( 1 m ) u 1 δ 1 v 1 + v 1 1 + ρ v 1 ,
with
u 1 ( x , 0 ) = u 0 ( x ) + ϵ 2 , v 1 ( x , 0 ) = v 0 ( x ) + ϵ 2 .
The variable ϵ 2 represents a positive perturbation added to the initial conditions to get the upper solutions. Also, the lower solutions u 2 , v 2 of (4) are given as solutions for the following problem:
u 2 t = d 1 2 u 2 x 2 + ϵ 1 u 2 x + u 2 u 2 2 α ( 1 m ) u 2 v 2 β + ( 1 m ) u 2
v 2 t = d 2 2 v 2 x 2 + ϵ 1 v 2 x + γ ( 1 m ) u 2 v 2 β + ( 1 m ) u 2 δ 1 v 2 + v 2 1 + ρ v 2 ,
with
u 2 ( x , 0 ) = u 0 ( x ) , v 2 ( x , 0 ) = v 0 ( x ) .
For a test function ξ 2 , the following inequalities are satisfied:
0 R u 1 u 2 x , t ξ 2 x , t d x = 0 t R { u 1 u 2 ξ 2 t + d 1 u 1 u 2 2 ξ 2 x 2 + ϵ 1 u 1 u 2 ξ 2 x + ( u 1 u 2 ) ξ 2 ( u 1 2 u 2 2 ) ξ 2 + α ( 1 m ) u 2 v 2 β + ( 1 m ) u 2 α ( 1 m ) u 1 v 1 β + ( 1 m ) u 1 ξ 2 } d x d s ,
0 R v 1 v 2 x , t ξ 2 x , t d x = 0 t R { v 1 v 2 ξ 2 t + d 2 v 1 v 2 2 ξ 2 x 2 + ϵ 1 v 1 v 2 ξ 2 x + γ ( 1 m ) u 1 v 1 β + ( 1 m ) u 1 γ ( 1 m ) u 2 v 2 β + ( 1 m ) u 2 ξ 2 δ 1 ( v 1 v 2 ) ξ 2 + v 1 1 + ρ v 1 v 2 1 + ρ v 2 ξ 2 } d x d s .
We define the following test function for the evaluation of integrals:
ξ ( | x | , s ) = e K s ( 1 + | x | 2 ) μ ,
where K > 0 and μ > 0 are constants and the following holds:
ξ 2 s = K ξ 2 ( | x | , s ) , | ξ 2 x | k 1 ( μ , d 1 ) ξ 2 ( | x | , s ) , | 2 ξ 2 x 2 | k 2 ( μ , d 1 ) ξ 2 ( | x | , s ) ,
so that
    ( u 1 u 2 ) ξ 2 t + d 1 ( u 1 u 2 ) | 2 ξ 2 x 2 | + ϵ 1 ( u 1 u 2 ) | ξ 2 x |   + ( u 1 u 2 ) ξ 2 ( u 1 + u 2 ) ( u 1 u 2 ) ξ 2 + α ( 1 m ) u 2 v 2 β + ( 1 m ) u 2 α ( 1 m ) u 1 v 1 β + ( 1 m ) u 1 ξ 2   ( u 1 u 2 ) K ξ 2 + d 1 ( u 1 u 2 ) k 2 ξ 2 + ϵ 1 ( u 1 u 2 ) k 1 ξ 2   + ( 1 ( u 1 + u 2 ) ) ( u 1 u 2 ) ξ 2 + α ( 1 m ) u 2 v 2 β + ( 1 m ) u 2 α ( 1 m ) u 1 v 1 β + ( 1 m ) u 1 ξ 2 ,
and
( v 1 v 2 ) ξ 2 s + d 2 ( v 1 v 2 ) | 2 ξ 2 x 2 | + ϵ 1 ( v 1 v 2 ) | ξ 2 x | + γ ( 1 m ) u 1 v 1 β + ( 1 m ) u 1 γ ( 1 m ) u 2 v 2 β + ( 1 m ) u 2 ξ 2   δ 1 ( v 1 v 2 ) ξ 2 + v 1 1 + ρ v 2 v 2 1 + ρ v 1 ξ 2   ( v 1 v 2 ) K ξ 2 + d 2 ( v 1 v 2 ) k 2 ξ 2 + ϵ 1 ( v 1 v 2 ) k 1 ξ 2   + γ ( 1 m ) u 1 v 1 β + ( 1 m ) u 1 γ ( 1 m ) u 2 v 2 β + ( 1 m ) u 2 ξ 2   + δ 1 ( v 1 v 2 ) ξ 2 + v 2 1 + ρ v 2 v 1 1 + ρ v 1 ξ 2 .
Now, K is chosen so that the following expressions are satisfied:
  ( u 1 u 2 ) K + d 1 ( u 1 u 2 ) k 2 ( 1 ( u 1 + u 2 ) ) ( u 1 u 2 )   + α ( 1 m ) u 2 v 2 β + ( 1 m ) u 2 α ( 1 m ) u 1 v 1 β + ( 1 m ) u 1 0 , and   ( v 1 v 2 ) K + d 2 ( v 1 v 2 ) k 2 +   + γ ( 1 m ) u 1 v 1 β + ( 1 m ) u 1 γ ( 1 m ) u 2 v 2 β + ( 1 m ) u 2   + δ 1 ( v 1 v 2 ) + v 2 1 + ρ v 2 v 1 1 + ρ v 1 0 .
Now, (49) and (50) become
R ( u 1 u 2 ) ( x , t ) ξ 2 ( x , t ) d x d s 0 t R ϵ 1 ( u 1 u 2 ) ξ 2 d x d s ,
and
R ( v 1 v 2 ) ( x , t ) ξ 2 ( x , t ) d x d s 0 t R ϵ 1 ( v 1 v 2 ) ξ 2 d x d s .
Differentiating w.r.t t we get
d d t R ( u 1 u 2 ) ( x , t ) ξ 2 ( x , t ) d x ϵ 1 R ( u 1 u 2 ) ξ 2 d x ,
and
d d t R ( v 1 v 2 ) ( x , t ) ξ 2 ( x , t ) d x ϵ 1 R ( v 1 v 2 ) ξ 2 d x .
Applying Grownwall’s inequality, we get
R ( u 1 u 2 ) ( x , t ) ξ 2 ( x , t ) d x 0 ,
and
R ( v 1 v 2 ) ( x , t ) ξ 2 ( x , t ) d x 0 .
Since ( u 1 , v 1 ) are upper solutions and ( u 2 , v 2 ) are lower solutions, we have
R ( u 1 u 2 ) ( x , t ) ξ 2 ( x , t ) d x = 0 ,
and
R ( v 1 v 2 ) ( x , t ) ξ 2 ( x , t ) d x = 0 .
Since ξ 2 is a test function, u 1 u 2 = 0 and v 1 v 2 = 0 ; i.e., u 1 = u 2 and v 1 = v 2 , which shows uniqueness of the solutions of (4). □

3. Stability Analysis

In this section we obtain the global stability for system (4) at equilibrium points ( 1 , 0 ) and 0 , 1 δ 1 ρ δ 1 .
Theorem 3.
The equilibrium point B 0 = ( 0 , 1 δ 1 ρ δ 1 ) is globally asymptotically stable.
Proof. 
Since u and v are zero at the boundary, we define the Lyapunov function as follows:
L B 0 ( t ) = Ω B 2 u 2 + B 3 v 1 δ 1 ρ δ 1 2 d x ,
where B 2 > 0 , and B 3 > 0 are constants. Differentiating (64) in t, we get
d L B 0 ( t ) d t = Ω 2 B 2 u u t + 2 B 3 v 1 δ 1 ρ δ 1 v t d x .
After using (4), we obtain
d L B 0 ( t ) d t = Ω 2 B 2 u d 1 2 u x 2 + ϵ 1 u x + u ( 1 u ) α ( 1 m ) u v β + ( 1 m ) u   + 2 B 3 v 1 δ 1 ρ δ 1 d 2 2 v x 2 + ϵ 1 v x + γ ( 1 m ) u v β + ( 1 m ) u δ 1 v + v 1 + ρ v d x .
Now employing the equilibrium point 0 , 1 δ 1 ρ δ 1 in (26), we get
d L B 0 ( t ) d t = Ω 2 B 2 d 1 u 2 u x 2 + 2 B 3 d 2 v 1 δ 1 ρ δ 1 2 v x 2 d x .
Since u and v x are zero at the boundary, the following expression holds:
d L B 0 ( t ) d t = 2 d 1 B 2 Ω u x 2 d x 2 d 2 B 3 Ω v x 2 d x ,
which implies that d L B 0 ( t ) d t 0 . So Theorem 3 is proved. □
Theorem 4.
The equilibrium point B 0 = 1 , 0 is globally asymptotically stable.
Proof. 
Since u and v are zero at the boundary, we define the Lyapunov function as follows:
L B 1 ( t ) = Ω B 2 u 1 2 + B 3 v 2 d x ,
where B 2 > 0 and B 3 > 0 are constants. Differentiating (64) in t, we get
d L B 1 ( t ) d t = Ω 2 B 2 u 1 2 u t + 2 B 3 v v t d x .
After using (4), we obtain
d L B 1 ( t ) d t = Ω 2 B 2 ( u 1 ) d 1 2 u x 2 + ϵ 1 u x + u ( 1 u ) α ( 1 m ) u v β + ( 1 m ) u   + 2 B 3 v d 2 2 v x 2 + ϵ 1 v x + γ ( 1 m ) u v β + ( 1 m ) u δ 1 v + v 1 + ρ v d x .
Now employing the equilibrium point 1 , 0 in (71), we get
d L B 1 ( t ) d t = Ω 2 B 2 d 1 u 2 u x 2 + 2 B 3 d 2 v 1 δ 1 ρ δ 1 2 v x 2 d x .
Since u and v x are zero at the boundary, the following expression holds:
d L B 1 ( t ) d t = 2 d 1 B 2 Ω u x 2 d x 2 d 1 B 2 Ω u 1 u 2 d x 2 d 2 B 3 Ω v x 2 d x ,
which implies that d L B 1 ( t ) d t 0 . So Theorem 4 is proved. □

4. Profiles of Solution

The objective of this section is to explore the behavior of solutions to equations of (4) when u ( x , t ) and v ( x , t ) are constant at x .
Theorem 5.
Assume that u ( x , t ) and v ( x , t ) are solutions to (4), with ϵ 1 α 1 and ϵ 1 α 3 ; then it holds that
u ( x , t ) = | u ( x , 0 ) | e ϵ 1 d 1 | x | t α 2 + t α 1 α 1 + 1 ( α 1 + 1 ) e ϵ 1 d 1 | x | t α 2 ,
and
v ( x , t ) = | v ( x , 0 ) | e ϵ 1 d 2 | x | t α 4 + t α 3 α 3 δ 1 + 1 ρ ( α 3 δ 1 ) + α 3 δ 1 + 1 ρ ( α 3 δ 1 ) e ϵ 1 d 2 | x | t α 4 .
with
α 1 = α 3 = 1 2 , α 2 = α 4 = 1 2 .
Proof. 
To examine the solutions to (4), we introduce self-similar structures of the form
u ( x , t ) = t α 1 f ( ζ ) , ζ = | x | t α 2 ,
and
v ( x , t ) = t α 3 g ( η ) , η = | x | t α 4 .
Substituting (74) and (75) into (4), we obtain
α 1 t α 1 1 f + α 2 ζ t α 1 1 f = d 1 t α 1 + 2 α 2 ( f ) + ϵ 1 t α 1 + α 2 ( f )   + t α 1 f ( 1 t α 1 f ) α ( 1 m ) ( t α 1 f ) ( t α 3 g ) β + ( 1 m ) ( t α 1 g ) ,
and
α 3 t α 3 1 g + α 4 η t α 3 1 g = d 2 t α 3 + 2 α 4 ( g ) + ϵ 1 t α 3 + α 4 ( g )   + γ ( 1 m ) ( t α 1 f ) ( t α 3 g ) β + ( 1 m ) ( t α 1 g ) δ 1 t α 3 g + t α 3 g 1 + ρ t α 3 g .
Additionally, considering the energy conservation throughout the evolution,
R t α 1 f ( | x | t α 2 ) d x = M 1 ,
and
R t α 3 g ( | x | t α 4 ) d x = M 2 ,
Using (78),
| x | = ζ t α 2 ,
and using (79),
| x | = η t α 4 ,
The differential unitary elements read
d x = d ζ t α 2 ,
and
d x = d η t α 4 ,
so that
R t α 1 f ( | x | t α 2 ) d x = t α 1 α 2 R f ( ζ ) d ζ = M 1 ,
and
R t α 3 g ( | x | t α 4 ) d x = t α 3 α 4 R g ( η ) d η = M 2 ,
where M 1 and M 2 are positive constants referring to the energy state of the predator–prey model. By invoking the principle of energy conservation across the domain, we assume that α 1 α 2 = 0 and α 3 α 4 = 0 . This assumption ensures that the total energy remains constant over time, reflecting the conservative nature of the system. Consequently, we can determine the specific values of α i ( i = 1 , 2 , 3 , 4 ) by analyzing the powers of the time variables in (76) and (77):
α 1 1 = α 1 + 2 α 2 α 1 α 2 = 0 ,
and
α 3 1 = α 3 + 2 α 4 α 3 α 4 = 0 .
After solving we have
α 1 = α 3 = 1 2 , α 2 = α 4 = 1 2 .
For simplicity, we take t = 1 in (76) and (77), and we have
d 1 ( f ) + ϵ 1 f α 2 ζ f + α 1 f + f 1 f α ( 1 m ) f g β + ( 1 m ) f = 0 ,
and
d 2 ( g ) + ϵ 1 g α 4 η g + α 3 g + γ ( 1 m ) f g β + ( 1 m ) f δ 1 g + g 1 + ρ g = 0 .
Choosing α = γ and after subtraction of (90) from (89), we have
d 1 ( f ) + ϵ 1 ( f ) α 2 ζ f + α 1 f + f f 2 d 2 ( g ) + ϵ 1 ( g ) α 4 η g + α 3 g δ 1 g + g 1 + ρ g = 0 ,
which is possible if
d 1 ( f ) + ϵ 1 ( f ) α 2 ζ f + α 1 f + f f 2 = 0 ,
and
d 2 ( g ) + ϵ 1 ( g ) α 4 η g + α 3 g δ 1 g + g 1 + ρ g = 0 .
For the exact solutions to (92) and (93), we take α 2 = α 1 and α 4 = α 3 ; then the above equations become
d 1 ( f ) + ϵ 1 ( f ) + α 1 ζ f + f f 2 = 0 ,
and
d 2 ( g ) + ϵ 1 ( g ) + α 3 η g δ 1 g + g 1 + ρ g = 0 .
Using a perturbation technique with a small parameter d 1 to solve the above Equation (94), neglect the highest derivative term in the outer region:
( ϵ 1 + α 1 ζ ) ( f ) + ( α 1 + 1 ) f f 2 = 0 ,
As for the outer region, ζ is large and f is constant, so assuming slow variation ( f = 0 ), solving the algebraic equation we have a non-trivial solution, which is typically dominant:
f o u t e r = α 1 + 1 ,
Now we focus on the boundary layer at ζ = 0 . Rescale ζ near the boundary layer
ζ 1 = ζ d 1
Substitute into original equation and simplify for d 1 < < 1 :
F + ϵ 1 F = 0 .
Solving for the inner region,
F ( ζ 1 ) = A + B e ϵ 1 ζ 1 .
Now we have matching ζ 1 , F ( ζ 1 ) f o u t e r , so we get A = α 1 + 1 , and from the boundary condition at ζ = 0 , we get F ( 0 ) = f ( 0 ) . Therefore, we obtain B = f ( 0 ) ( α 1 + 1 ) . So the inner solution in terms of ζ can be obtained:
f i n n e r = α 1 + 1 + ( f ( 0 ) ( α 1 + 1 ) ) e ϵ 1 ζ d 1 .
Now for the composite solution:
f c o m p ( ζ ) = f o u t e r + f i n n e r f o v e r l a p .
After computing we get
f ( ζ ) = ( α 1 + 1 ) + [ f ( 0 ) ( α 1 + 1 ) ] e ϵ 1 ζ d 1 .
Similarly solving (95) as we solved (94), we get
g ( η ) = α 3 δ 1 + 1 ρ ( α 3 δ 1 ) + g ( 0 ) + α 3 δ 1 + 1 ρ ( α 3 δ 1 ) e ϵ 1 η d 2 .
After substituting (103) and (104) into (74) and (75), we obtain
u ( x , t ) = | u ( x , 0 ) | e ϵ 1 d 1 | x | t α 2 + t α 1 α 1 + 1 ( α 1 + 1 ) e ϵ 1 d 1 | x | t α 2 ,
and
v ( x , t ) = | v ( x , 0 ) | e ϵ 1 d 2 | x | t α 4 + t α 3 α 3 δ 1 + 1 ρ ( α 3 δ 1 ) + α 3 δ 1 + 1 ρ ( α 3 δ 1 ) e ϵ 1 d 2 | x | t α 4 ,
as we intended to prove.
These solutions show that if we take an advection coefficient greater than 0.5 then the ecosystem has an initially heterogeneous state, and after an increase in time then the ecosystem becomes stable. The prey and predator populations slowly decrease but keep positive and smooth in the sense of spatial patterns, and the predator population is preserved by supplementary food. So the ecological process is realistic. □

5. Stationary Solutions

The coming idea consists of exploring upper stationary solution to (4). For this, we first look at upper solutions to u and v:
u t = d 1 2 u x 2 + ϵ 1 u x + u ( 1 u ) α ( 1 m ) u v β + ( 1 m ) u   d 1 2 u x 2 + ϵ 1 u x + u ( 1 u ) + α ( 1 m ) u v ( 1 m ) u   d 1 2 u x 2 + ϵ 1 u x + u ( 1 u ) + α v   d 1 2 u x 2 + ϵ 1 u x + u ( 1 u ) + α K 1 ˜ u ,
where K 1 ˜ u = s u p { v } and
v t = d 2 2 v x 2 + ϵ 1 v x + γ ( 1 m ) u v β + ( 1 m ) u δ 1 v + v 1 + ρ v   d 2 2 v x 2 + ϵ 1 v x + γ ( 1 m ) u v ( 1 m ) u δ 1 v + v ρ v   d 2 2 v x 2 + ϵ 1 v x + γ v δ 1 v + 1 ρ .
So, we have
u t = d 1 2 u x 2 + ϵ 1 u x + α K 1 ˜ + 1 u u 2 ,
and
v t = d 2 2 v x 2 + ϵ 1 v x + ( γ δ 1 ) v + 1 ρ .
For the stationary upper solution, the above equations can be written as follows:
d 1 d 2 u d x 2 + ϵ 1 d u d x + α K 1 ˜ + 1 u u 2 = 0 ,
and
d 2 d 2 v d x 2 + ϵ 1 d v d x + ( γ δ 1 ) v + 1 ρ = 0 .
To investigate the solution, we use the definition of a Hamiltonian (see [21]) so that
1 2 d 1 ( u ) 2 + ϵ 1 u + α K 1 ˜ + 1 u 2 2 u 3 3 = 0 ,
and
1 2 d 2 ( v ) 2 + ϵ 1 v + ( γ δ 1 ) v 2 2 + v ρ = 0 ,
which implies that
d 1 u 2 + 2 ϵ 1 u + α K 1 ˜ + 1 u 2 2 3 u 3 = 0 ,
and
d 2 v 2 + 2 ϵ 1 v + ( γ δ 1 ) v 2 + 2 v ρ = 0 .
For solving (113), we use Lighthill’s technique with small parameter ϵ 1 ; for this we introduce strained coordinates
x = s + ϵ 1 w 1 ( s ) + ϵ 1 2 w 2 ( s ) + and d d x = ( 1 ϵ 1 d w 1 d s + ) d d s .
Substituting this into (113), we have
d 1 ( 1 ϵ 1 d w 1 d s + ) 2 d u d s 2 + 2 ϵ 1 u + α K 1 ˜ + 1 u 2 2 3 u 3 = 0 .
Now we assume that
u ( s ) = u 0 ( s ) + ϵ 1 u 1 ( s ) + ϵ 1 2 u 2 ( s ) +
After substituting (117) into (116) and comparing the power of ϵ 1 , we have
d 1 d u 0 d s 2 + α K 1 ˜ + 1 u 0 2 2 3 u 0 3 = 0 ,
and
2 d 1 d u 0 d s d u 1 d s 2 d 1 d w 1 d s d u 0 d s 2 + 2 u 0 + 2 α K 1 ˜ + 1 u 0 u 1 2 u 0 2 u 1 = 0 .
After solving (118), we get
u 0 = 3 2 ( α K 1 ˜ + 1 ) s e c 2 1 2 α K 1 ˜ + 1 d 1 s .
After substituting (120) into (119) and then choosing
w 1 = d 1 ( α K 1 ˜ + 1 ) 5 2 4 3 c o t 1 2 α K 1 ˜ + 1 d 1 s + 3 s i n α K 1 ˜ + 1 d 1 s s ( α K 1 ˜ + 1 ) 2 ,
we obtain
d u 1 d s + α K 1 ˜ + 1 d 1 [ c o s 1 2 α K 1 ˜ + 1 d 1 s s i n 1 2 α K 1 ˜ + 1 d 1 s 3 s e c 2 1 2 α K 1 ˜ + 1 d 1 s 2 t a n 1 2 α K 1 ˜ + 1 d 1 s ] u 1 = 0 .
After solving we get
u 1 ( s ) = s i n 1 2 α K 1 ˜ + 1 d 1 s c o s 1 2 α K 1 ˜ + 1 d 1 s 3 .
Putting (120) and (123) into (117), we get
u ( s ) = 3 2 ( α K 1 ˜ + 1 ) s e c 2 1 2 α K 1 ˜ + 1 d 1 s + ϵ 1 s i n 1 2 α K 1 ˜ + 1 d 1 s c o s 1 2 α K 1 ˜ + 1 d 1 s 3 + O ϵ 1 2 ,
and
x = s + ϵ 1 d 1 ( α K 1 ˜ + 1 ) 5 2 4 3 c o t 1 2 α K 1 ˜ + 1 d 1 s + 3 s i n α K 1 ˜ + 1 d 1 s s ( α K 1 ˜ + 1 ) 2 + O ϵ 1 2 ,
where s n π d 1 ( α K 1 ˜ + 1 ) . Solving (114) for δ 1 > γ we have
d 2 v 2 + 2 ( ϵ 1 + 1 ρ ) v ( δ 1 γ ) v 2 = 0 ,
which implies that
v = δ 1 γ d 2 v 2 2 ( ϵ 1 + 1 ρ ) v ( δ 1 γ ) .
After solving, we get
v = ϵ 1 + 1 ρ δ 1 γ + 1 2 e δ 1 γ d 2 x + ϵ 1 + 1 ρ δ 1 γ 2 e δ 1 γ d 2 x .
These results show that if the direct movement of prey and predator populations are small then the spatial distribution of these populations are high in the main region, and away from this area the prey and predator populations decrease. This slight shift in populations does not change the balance of the system. So the ecosystem remains stable in the sense of spatial patterns.

6. Asymptotic Profiles of Solutions

In this section, we obtain the nonstationary upper solutions of (4). To this end, we consider nonlinear point scaling to develop the solution profile of (107) and (108):
u ( t , x ) = e z 2 ( t , x ) ,
and
v ( t , x ) = e z 3 ( t , x ) .
This type of scaling has been extensively utilized (see [20,22,23,24,25]). Hence, u and v satisfy the following Hamilton–Jacobi-type equations:
e z 2 z 2 t = d 1 e z 2 z 2 x 2 + d 1 2 z 2 x 2 e z 2 + ϵ 1 e z 2 z 2 x + ( 1 + α K 1 ˜ ) e z 2 e 2 z 2 ,
and
e z 3 z 3 t = d 2 e z 3 z 3 x 2 + d 2 2 z 3 x 2 e z 3 + ϵ 1 e z 3 z 3 x + ( γ δ 1 ) e z 3 + 1 ρ ,
where 2 z 2 x 2 z 2 x and 2 z 3 x 2 z 3 x . Also z 2 ( t , x ) 0 and z 3 ( t , x ) 0 at x , so e z 2 ( t , x ) = 1 and e z 3 ( t , x ) = 1 . This idea is taken from Galaktionov and Williams [22] (see Proposition 3.1 in this reference), which simplifies to
z 2 t = d 1 z 2 x 2 + ϵ 1 z 2 x + 1 + α K 1 ˜ ,
and
z 3 t = d 2 z 3 x 2 + ϵ 1 z 3 x + ( γ δ 1 ) + 1 ρ .
We assume
z 4 ( t , x ) = z 2 1 + α K 1 ˜ t ,
and
z 5 ( t , x ) = Z 3 ( γ δ 1 ) + 1 ρ t .
After substituting into (133) and (134), we get
z 4 t = d 1 z 4 x 2 + ϵ 1 z 2 x ,
and
z 5 t = d 2 z 5 x 2 + ϵ 1 z 5 x .
For the solution of (137) and (138), we assume ζ = x a t , so the above equations become
d 1 d z 4 d ζ 2 + ϵ 1 + a d z 4 d ζ = 0 ,
and
d 2 d z 5 d ζ 2 + ϵ 1 + a d z 5 d ζ = 0 .
After solving (139) in (140), we get
Z 4 = 1 + α K 1 ˜ t ( ϵ 1 + a ) d 1 ζ ,
and
Z 5 = ( γ δ 1 ) + 1 ρ t ( ϵ 1 + a ) d 2 ζ ,
where ( ϵ 1 + a ) d 2 is small and we can choose ( ϵ 1 + a ) d 1 = e ζ 2 and ( ϵ 1 + a ) d 2 = e ζ 2 , which approach zero when ζ . Putting z 4 and z 5 in (135) and (136), we get
Z 2 = ( ϵ 1 + a ) d 1 x a t ,
and
Z 3 = ( ϵ 1 + a ) d 2 x a t ,
Finally substituting (143) and (134) into (129) and (130), we get
u ( t , x ) = e ( ϵ 1 + a ) d 1 x a t .
Similarly, solving (134), we have
v ( t , x ) = e ( ϵ 1 + a ) d 2 x a t .
These solutions show the migration of prey and predator populations with a constant speed. If we take a large diffusion coefficient the populations spread slowly and widely, and if we take a small diffusion coefficient then this spread is quick. The populations slowly spreading is much more realistic than a quick spread in the ecosystem.

7. Conclusions

In this work, we present a model for describing the dynamics of the predator–prey model with the Holling type II interaction function with diffusion and advection effects. The diffusion effect means the smooth spread of prey and predator populations in space over the time, and advection effect means direct movement of prey and predator populations in space. Firstly, we demonstrate that model (4) exhibits boundedness, uniqueness and stability of solutions. For stability, we use the Lyapunov function and show the global stability at equilibrium points. Subsequently, we obtain the value of the advection coefficient, which is greater than 0.5. With these values, the ecosystem starts in a heterogeneous state and after some time it becomes stable. To obtain these results, we use the singular perturbation technique. Moreover we find that if the advection coefficient is small, this means less direct movement and then the populations in the main region are high, and away from that region populations decrease. To obtain this result, we use the definition of a Hamiltonian. Lastly, we show that for a large diffusion coefficient, the population spreads slowly and wildly, and a for small diffusion coefficient, the population spreads quickly. For this, we use nonlinear point scaling. For future work we can use the p-Laplacian operator instead of diffusion because if the populations are not spread smoothly then the p-Laplacian gives better results compared to diffusion.

Author Contributions

Conceptualization, S.U.R. and J.L.D.P.; methodology, S.U.R., J.L.D.P. and M.R.; validation, S.U.R. and J.L.D.P.; formal analysis, S.U.R., J.L.D.P. and M.R.; data curation, J.L.D.P.; writing—original draft, S.U.R. and M.R.; writing—review and editing, S.U.R., J.L.D.P. and M.R.; supervision, S.U.R. and J.L.D.P. All authors have read and agreed to the published version of the manuscript.

Funding

This research received no external funding.

Data Availability Statement

No new data were created or analyzed in this study.

Acknowledgments

We would like to thanks the referees and the editors for their careful reading and valuable comments and suggestions.

Conflicts of Interest

The authors declare no conflicts of interest.

References

  1. Lotka, A.J. Undamped oscillations derived from the law of mass action. J. Am. Chem. Soc. 1920, 42, 1595–1599. [Google Scholar] [CrossRef]
  2. Volterra, V. Variazioni e Fluttuazioni del Numero d’Individui in Specie Animali Conviventi; Memoria della Reale Accademia Nazionale dei Lincei: Rome, Italy, 1927. [Google Scholar]
  3. Han, R.; Mandal, G.; Guin, L.N.; Chakravarty, S. Dynamical response of a reaction-diffusion predator-prey system with cooperative hunting and prey refuge. J. Stat. Mech. Theory Exp. 2022, 2022, 103502. [Google Scholar]
  4. Mandal, G.; Ali, N.; Guin, L.N.; Chakravarty, S. Impact of fear on a tri-trophic food chain model with supplementary food source. Int. J. Dyn. Control 2023, 11, 2127–2160. [Google Scholar] [CrossRef]
  5. Samaddar, S.; Dhar, M.; Bhattacharya, P. Effect of fear on prey-predator dynamics: Exploring the role of prey refuge and additional food. Chaos Interdiscip. J. Nonlinear Sci. 2020, 30, 063129. [Google Scholar] [CrossRef] [PubMed]
  6. Alves, T.M.; Hilker, M.F. Hunting cooperation and Allee effects in predators. J. Theor. Biol. 2017, 419, 19–22. [Google Scholar] [CrossRef] [PubMed]
  7. Wang, J.; Cai, Y.; Fu, S.; Wang, W. The effect of the fear factor on the dynamics of a predator-prey model incorporating the prey refuge. Chaos Interdiscip. J. Nonlinear Sci. 2019, 29, 083109. [Google Scholar] [CrossRef]
  8. Courchamp, F.; Berec, L.; Gascoigne, J. Allee Effects in Ecology and Conservation; Oxford University Press: Oxford, UK, 2008. [Google Scholar]
  9. Guin, L.N.; Pal, S.; Chakravarty, S. Pattern dynamics of a reaction-diffusion predator-prey system with both refuge and harvesting. Int. J. Biomath. 2021, 14, 2050084. [Google Scholar] [CrossRef]
  10. Zhao, K. A generalized stochastic Nicholson blowfly model with mixed time-varying lags and harvest control: Almost periodic oscillation and global stable behavior. Adv. Contin. Discret. Models 2025, 2025, 171. [Google Scholar] [CrossRef]
  11. Ali, A.; Jawad, S. Stability analysis of the depletion of dissolved oxygen for the Phytoplankton-Zooplankton model in an aquatic environment. Iraqi J. Sci. 2024, 65, 2736–2748. [Google Scholar] [CrossRef]
  12. Holling, C.S. The functional response of predators to prey density and its role in mimicry and population regulation. Mem. Entomol. Soc. Can. 1965, 97, 5–60. [Google Scholar] [CrossRef]
  13. Lotka, A.J. Elements of Physical Biology; Williams and Wilkins: Baltimore, MD, USA, 1925. [Google Scholar]
  14. Kar, T.K. Stability analysis of a prey-predator model incorporating a prey refuge. Commun. Nonlinear Sci. Numer. Simul. 2005, 10, 681–691. [Google Scholar] [CrossRef]
  15. Xu, Y.; Krause, A.L.; Van Gorder, R.A. Generalist predator dynamics under Kolmogorov versus non-Kolmogorov models. J. Theor. Biol. 2020, 486, 110060. [Google Scholar] [CrossRef]
  16. Erbach, A.; Lutscher, F.; Seo, G. Bistability and limit cycles in generalist predator-prey dynamics. Ecol. Complex. 2013, 14, 48–55. [Google Scholar] [CrossRef]
  17. Mondal, S.; Samanta, G.P. Dynamics of a delayed predator-prey interaction incorporating nonlinear prey refuge under the influence of fear effect and additional food. J. Phys. A Math. Theor. 2020, 53, 295601. [Google Scholar] [CrossRef]
  18. Sen, D.; Ghorai, S.; Sharma, S.; Banerjee, M. Allee effect in prey’s growth reduces the dynamical complexity in prey-predator model with generalist predator. Appl. Math. Model. 2021, 91, 768–790. [Google Scholar] [CrossRef]
  19. Mondal, B.; Sarkar, S.; Ghosh, U. Complex dynamics of a generalist predator-prey model with hunting cooperation in predator. Eur. Phys. J. Plus 2021, 137, 43. [Google Scholar] [CrossRef]
  20. Leacock, R.A.; Padgett, M.J. Hamilton-Jacobi/action-angle quantum mechanics. Phys. Rev. D 1983, 28, 2491. [Google Scholar] [CrossRef]
  21. Bonheure, D.; Sanchez, L. Heteroclinic orbits for some classes of second and fourth order differential equations. In Handbook of Differential Equations: Ordinary Differential Equations; North Holland Publishing: Amsterdam, The Netherlands, 2006; Volume 3, pp. 103–202. [Google Scholar]
  22. Galaktionov, V.A.; Williams, J. Blow-up in a fourth-order semilinear parabolic equation from explosion-convection theory. Eur. J. Appl. Math. 2004, 14, 745–764. [Google Scholar] [CrossRef]
  23. Leacock, R.A.; Padgett, M.J. Hamilton-Jacobi theory and the quantum action variable. Phys. Rev. Lett. 1983, 50, 3. [Google Scholar] [CrossRef]
  24. Bhalla, R.S.; Kapoor, A.K.; Panigrahi, P. Quantum Hamilton-Jacobi formalism and the bound state spectra. Am. J. Phys. 1997, 65, 1187–1194. [Google Scholar] [CrossRef]
  25. Pablo, A.D.; Vazquez, J.L. Travelling waves and finite propagation in a reaction-diffusion equation. J. Differ. Equ. 1991, 93, 19–61. [Google Scholar] [CrossRef]
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MDPI and ACS Style

Rahman, S.U.; Palencia, J.L.D.; Rehman, M. Existence, Uniqueness and Solutions for Diffusion and Advection Effects for Predator–Prey Model with Holling Type II Interaction Function. Mathematics 2026, 14, 831. https://doi.org/10.3390/math14050831

AMA Style

Rahman SU, Palencia JLD, Rehman M. Existence, Uniqueness and Solutions for Diffusion and Advection Effects for Predator–Prey Model with Holling Type II Interaction Function. Mathematics. 2026; 14(5):831. https://doi.org/10.3390/math14050831

Chicago/Turabian Style

Rahman, Saeed Ur, José Luis Díaz Palencia, and Maria Rehman. 2026. "Existence, Uniqueness and Solutions for Diffusion and Advection Effects for Predator–Prey Model with Holling Type II Interaction Function" Mathematics 14, no. 5: 831. https://doi.org/10.3390/math14050831

APA Style

Rahman, S. U., Palencia, J. L. D., & Rehman, M. (2026). Existence, Uniqueness and Solutions for Diffusion and Advection Effects for Predator–Prey Model with Holling Type II Interaction Function. Mathematics, 14(5), 831. https://doi.org/10.3390/math14050831

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