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Article

Bryophytes of the Serra dos Órgãos National Park: Endemism and Conservation in the Atlantic Forest

by
Jéssica Soares de Lima
1,*,
Allan Laid Alkimim Faria
2,
Mateus Tomás Anselmo Gonçalves
1 and
Denilson Fernandes Peralta
1
1
Instituto de Pesquisas Ambientais, Avenida Miguel Stéfano, 3687, São Paulo 04301-102, SP, Brazil
2
Instituto de Biologia, Universidade de Brasília, Campus Darcy Ribeiro, Asa Norte, Brasília 70910-900, DF, Brazil
*
Author to whom correspondence should be addressed.
Plants 2025, 14(15), 2419; https://doi.org/10.3390/plants14152419
Submission received: 16 June 2025 / Revised: 7 July 2025 / Accepted: 22 July 2025 / Published: 4 August 2025
(This article belongs to the Special Issue Diversity, Distribution and Conservation of Bryophytes)

Abstract

This study presents a comprehensive inventory of bryophytes in Serra dos Órgãos National Park (PARNASO), aiming to evaluate species richness, floristic composition and threatened taxa. Despite the state of Rio de Janeiro being one of the most extensively sampled regions for bryophytes in Brazil, detailed surveys of its conservation units remain scarce. Data were obtained through bibliographic review, herbarium specimen analysis, and new field collections. A total of 504 species were recorded, belonging to 202 genera and 76 families. The park harbors three locally endemic species, eight endemic to Rio de Janeiro, and sixty-nine species endemic to Brazil. Additionally, eleven species were identified as threatened, comprising seven Endangered (EN), two Critically Endangered (CR), and two Vulnerable (VU) according to the IUCN guidelines. PARNASO includes four distinct ecosystems along an altitudinal gradient: sub-montane forest (up to 500 m), montane forest (500–1500 m), upper-montane forest (1500–2000 m), and high-altitude fields (above 2000 m). Montane Forest showed the highest species richness, followed by high-altitude fields, upper-montane forest, and sub-montane forest. The findings highlight PARNASO’s importance in preserving bryophyte diversity in a highly diverse yet understudied region. This work contributes valuable baseline data to support conservation strategies and future ecological studies in Atlantic Forest remnants.

1. Introduction

Serra dos Órgãos National Park (PARNASO), established on November 30th, 1939, holds the distinction of being the third national park created in Brazil. This protected area is situated within the Atlantic Forest domain, specifically in the Serra do Mar massifs and called “Órgãos” in Portuguese because its peaks resemble and are named as “pipe organs”. The Serra do Mar is an extensive mountain range approximately 1500 km along the Brazilian coast, from the northern Santa Catarina State to Rio de Janeiro State [1,2].
PARNASO currently represents one of the most important remaining biodiversity hotspots in Rio de Janeiro State, since its ecosystems are one of the largest centers of flower plant endemism in the Atlantic Forest [3,4]. It also plays a vital role in preserving essential water sources that feed two main watersheds—Paraíba do Sul and several streams of the Baía de Guanabara [5,6,7,8].
The Atlantic Forest domain is one of the most species-rich forests in the tropics [4,9,10]. It is also recognized for having high levels of biodiversity and endemism, expressed via its complex structural environments [11,12]. Tropical forests harbor the highest diversity of bryophyte species in the world, not only due to their structural complexity and microhabitat heterogeneity, but also because of favorable climatic conditions such as high humidity, frequent precipitation, and relatively stable temperatures [13,14]. Some authors have reported that this diversity may vary along the altitudinal range, especially concerning structural and floristic composition [15,16,17,18].
The Atlantic Forest within PARNASO comprises well-preserved forests characterized by four distinct vegetation formations based on altitude, climate, and soil conditions, among other factors [6,19,20,21,22]. In the lower parts around Sede Guapimirim, sub-montane forest prevails, while near Sede at Teresópolis, the forest transitions from montane forest to upper-montane forest, culminating above 2000 m.a.s.l. and high-altitude fields along the trail from Teresópolis to Petrópolis [19].
The species list of the vascular plants in Flora Organensis [19] is strongly based on the work of 18th-century botanists and on collections made especially by A.C. Brade (1881–1971) and C.T. Rizzini (1921–1992) between the 1930s and 1950s of the 20th century. These collectors also recorded bryophytes, and the specimens were studied by Bartram (1954) [19], who identified 146 species. Most bryophyte specimens from Serra dos Órgãos are deposited in the Herbarium of Jardim Botânico do Rio de Janeiro (RB).
Bryophytes from PARNASO were not sufficiently explored, and even considering that, though several researchers collected and published new taxa, the study was dispersed as examined material in taxonomical revisionary work. For example, Faria et al. (2021) [23] attempted to re-collect species founded by Glaziou in Serra dos Órgaos and the taxonomic reclassification of an Atlantic Forest endemic, and Peralta et al. (2020) [24] reported species of the Andreaceae family from the region.
There is no other information on bryophytes in PARNASO; thus, there is no detailed data on the floristic structure and composition of bryophytes. A comprehensive species list will contribute to filling a significant knowledge gap for the state of Rio de Janeiro and support the assessment of threatened bryophyte species. In addition, the revision of herbarium specimens and the incorporation of new collections will help stimulate further research on the ecology and biogeography of bryophytes within this important national park. This work aims to provide an inventory of the bryophyte species in PARNASO and discuss the following topics: (i) species richness and floristic composition; (ii) substrate preferences; and (iii) endemism and conservation status.

2. Results

2.1. Species Richness and Floristic Composition

The present study documented 504 species in 202 genera and 76 families (Table 1). The Serra dos Órgaos National Park preserves 57% of the known bryophyte flora of Rio de Janeiro State and 31% of Brazil. A total of 3000 specimens were analyzed, resulting in the identification of 428 species and an additional 79 species (indicated by * in Table 1) that remain known only from the literature.
Three species are locally endemic, being known only from the type specimens collected over 100 years ago: Southbya organensis (Lützelburg 6003 p.p. [holotype JE]), Plagiochila pseudoradicans (Lützelburg 6029b [holotype JE]) and Dicranodontium pulchroalare ssp. brasiliense (Lützelburg 1519 [holotype JE]). Additional field research is urgently needed in order to obtain new records and data of these three endemic species.
The most species-rich families of liverworts are Lejeuneaceae (65 spp.) and Lepidoziaceae (16 spp.), while for moss they are Leucobryaceae (30 spp.) and Sphagnaceae (25 spp.). A total of 8 species of hornworts were found, distributed among the families Dendrocerotaceae (4 spp.), Anthocerotaceae (2 spp.) and Nothotyladaceae (2 spp.). The most species-rich genera were Campylopus (Leucobryaceae, 23 spp.); Plagiochila (Plagiochilaceae, 21 spp.); Fissidens (Fissidentaceae, 20 spp.); Lejeunea (Lejeuneaceae, 16 spp.); and Metzgeria (Metzgeriaceae, 14 spp.). Considering the samples analyzed, the most common species with the largest number of records were Campylopus arctocarpus (7 specimens); Lejeunea acanthogona (7); Lejeunea flava (7); Radula nudicaulis (7); Bryum capillare (6); Fissidens wallisii (6); Lejeunea serpillifolioides (6); Rigodium toxarion (6); Trichocolea brevifissa (6); and Zelometeorium ambiguum (6).

2.2. Substrate Preferences

The bryophyte species predominantly colonized rocks (210 spp.), followed by tree trunks (209 spp.), soil (205 spp.), logs (50 spp.), and leaves (29 spp.) (Figure 1).
The availability of trees within the forests of PARNASO clearly influences the bryophyte species occurrence. Mosses predominantly colonize tree trunks, with only four species being found on leaves: Paraleucobryum longifolium (Dicranaceae); Philophyllum tenuifolium (Hookeriaceae); Toloxis imponderosa (Meteoriaceae); and Schlotheimia tecta (Orthotrichaceae).
The lowest bryophyte colonization was for leaves (epiphyllous). The leaf-colonizing species most frequent were the leafy liverworts of the family Lejeuneaceae (21 spp.), followed by Radulaceae (2 spp.) and the thallose liverwort family Metzgeriaceae (2 spp.). According to Gradstein et al. (2001) [17] and Ilkiu-Borges & Lisboa (2004) [57], species of Lejeuneaceae can colonize different types of substrates, but their occurrence on leaves is indicative of preserved native areas.

2.3. Endemism and Conservation Status

The Serra dos Órgaos National Park protects three species locally endemic, eight from Rio de Janeiro state, and 69 Brazilian (Table 2). Endemism and species richness are high with an increase in altitude, as was also found in Itatiaia National Park, located in Serra da Mantiqueira [18].
Of the 28 species of Brazilian endemic bryophytes exclusive to high-altitude fields, 12 belong to the family Sphagnaceae (42%). The species Sphagnum luetzelburgii and S. sehnemii are endemic to Brazil, occurring in the states of Paraná, Santa Catarina, Rio Grande do Sul, and Rio de Janeiro. However, in Rio de Janeiro State, these species are restricted to mountains of the Atlantic Forest and have only been reported for PARNASO by Costa (2021) [58] and not by the present work.
The lower number of endemic species recorded in sub-montane forest probably reflects the effects of ecological and biogeographic processes, as well as human interference [18]. Among the 19 endemic species exclusively found in montane forest, 13 are liverworts and 6 are mosses. The predominant liverwort family is Lejeuneaceae, with nine species, followed by Metzgeriaceae with three and Jamesoniellaceae with two.
We have re-collected some species considered rare, including Bryum limbatum. Ochi (1980) [59] cites a collection from Metzgeria furcata, collected by Bartram (1954) [19]; Sphagnum cuspidatum, collected by Luetzelburg in 1923; and Andreaea acutifolia, collected by Mueller in 1845. Andreaea acutifolia is always mistakenly identified as A. rupestris, the two differing mainly by perichaetial leaves [24]. The species S. cuspidatum is widespread in Brazil [60], but has been collected only once in PARNASO.
The Serra dos Orgaos Park also protects eleven of the species indicated as threatened by CNCFlora and Flora e Funga do Brasil (Figure 2 and Table 2) in the Rio de Janeiro state: seven as Endangered (EN), two as Critically Endangered (CR) and two as Vulnerable (VU). Among these threatened species, three are known only from the type material, and were not found in the field trips. Southbya organensis was described by Herzog (1949) [61] and is classified as Critically Endangered (CR) by CNCFlora (2012) [58] due to its distribution being restricted to Serra dos Órgãos, specifically Morro do Açu peak, which has excessive tourism and recurrent arson instead. Plagiochila pseudoradicans and Dicranodontium pulchroalare ssp. brasiliense were described by Herzog (1949) [61], and neither was assessed by CNCFlora (2012) [62] because they were considered as Data Deficient (DD) at that time.

3. Discussion

The Serra dos Órgãos National Park (PARNASO) exhibits remarkable bryophyte diversity. This richness highlights the park as a key conservation area for bryophytes in the Atlantic Forest. These findings suggest that even in areas already considered well-studied, significant knowledge gaps remain, highlighting the dynamic nature of bryophyte diversity and distribution.
One important factor contributing to this diversity is the presence of forested areas with abundant tree trunks, which serve as critical substrates for epiphytic bryophytes. The architectural, morphological, phenological, and chemical traits of different tree species create varied microhabitats that support a wide range of bryophyte taxa [57,63]. Shade-loving epiphytes, in particular, are commonly found in these forest environments but are highly sensitive to environmental disturbances. As such, they are often among the first species to disappear following canopy opening or habitat degradation [64], making them reliable indicators of forest health.
In contrast, mosses exhibit a broader ecological amplitude due to their desiccation tolerance and structurally complex adaptations, such as costa development, hyaline leaf apices, papillae, and specialized conducting tissues like hydroids and leptoids [29,65,66]. These features enable many mosses to colonize exposed and drier substrates, including rocky outcrops. Similar patterns of moss dominance on rock surfaces have been observed in other highland regions of Brazil, such as Chapada Diamantina [67,68], Serra do Caraça [69], and Serra da Canastra [70].
The high-altitude ecosystems of PARNASO further enhance bryophyte diversity by creating isolated and unique habitats. Mountain summits, often compared to ecological islands due to their environmental isolation, support specialized flora adapted to harsh conditions such as strong winds, low temperatures, and nutrient-poor soils [71,72,73]. These rupestrian fields form natural refugia, acting as barriers to typical forest species and favoring the emergence of narrow endemics and rare taxa [74,75,76].
Overall, the findings of this study underscore the ecological importance and conservation value of Serra dos Órgãos National Park. Alongside Itatiaia National Park, PARNASO remains one of the most critical remnants of Atlantic Forest in Rio de Janeiro and Brazil, particularly in terms of bryophyte conservation. The park’s diverse topography, wide range of microhabitats, and climatic heterogeneity—marked by high altitudes, low temperatures, and high light availability—create optimal conditions for bryophyte diversity. Furthermore, the presence of endemic and threatened species reinforces the park’s role as a conservation priority. Such studies are valuable as they reveal the significance of endemism and species richness, crucial factors in determining conservation priorities [77], and emphasize the importance of preserving Serra dos Órgãos National Park.

4. Materials and Methods

4.1. Study Area

Serra dos Órgãos National Park (PARNASO) is located in the highlands region of Rio de Janeiro State, Brazil, ranging the municipalities of Teresópolis, Petrópolis, Guapimirim and Magé (22°23′36.96″–22°34′57.72″ S and 43°10′57.72″–42°58′43.68″ W). It is a protected area with 20,024 hectares of Atlantic Forest in a central position in the Serra do Mar Ecological Corridor. Ranging in altitude from 80 to 2275 m.a.s.l. (Figure 3), it is the highest portion of the entire Serra do Mar [2]. The climate is tropical and super humid (80–90% relative humidity), with the annual average temperature ranging from 13 to 23 °C, although minimums can reach −5 °C during winter above 800 m.a.s.l. [6,8,74]. The average annual rainfall ranges from 1500 mm to 3000 mm, with higher concentrations of rain during summer (December to March) and a dry season during winter (June to August), while heat is well distributed throughout the year [78,79].
The vegetation classification adopted follows Veloso et al. (1991) [20] and Safford (1999) [80]: Sub-montane Forest (below 600 m.a.s.l., typically evergreen, with a high canopy and rich floristic diversity; dominant tree species include Protium spp., Virola spp., and members of the Myrtaceae and Fabaceae families); montane forest (600–1500, characterized by shorter trees and more frequent presence of cloud cover; common tree species include Clethra spp. and various species of Melastomataceae); upper-montane forest (above 1500, vegetation here is more stunted, often with gnarled trees adapted to frequent mist and lower temperatures). Dominant elements include Weinmannia paulliniifoli and Drimys brasiliensis, and high-altitude fields (above 1800–2000) are open grassland formations interspersed with shrubs, herbs, and occasional low woody elements. Common species include Vellozia spp., Paepalanthus spp., and Eryngium spp. (Figure 4).

4.2. Studied Taxa

The species list developed here is based on (i) a bibliographic survey conducted by consulting published volumes of major Brazilian botanical journals, as well as international bibliographic databases (Web of Science, SciELO, and Google Scholar), in search of peer-reviewed articles; the search focused on article titles containing combinations of the keywords “Serra dos Órgãos,” “bryophytes,” “mosses,” and “liverworts,” followed by a careful and detailed reading of each selected publication; (ii) herbarium specimens collected in the study area; and (iii) new collections (Figure 4).
Herbarium specimens were obtained from online sources like (GBIF [81], jabot, JStorTypes and SpeciesLink accessed several times). The specimens are requested on loan and re-determined and carefully examined for small species associations, with one of each species cited as voucher in the Table 1. The authors conducted four field excursions to the PARNASO, each lasting one week, two during the rainy season and two during the dry season, with the aim of locating species not previously recorded in the herbarium. The collection, preservation and herborization methodologies follow Frahm (2003) and employ free walks [82] for covering all available substrates like tree trunks, logs, soil, rocks and leaves.
Field trips in the Park intended to check if species collected in the past are yet present, to collect new specimens and to record new taxa. Collected samples were deposited in the herbarium of the University of Brasília (UB), with duplicates sent to the herbarium of Instituto de Pesquisas Ambientais (SP). The classification system used here follows Crandall-Stotler et al. [83] for Marchantiophyta, Goffinet et al. [65] for Bryophyta and Renzaglia et al. [84] for Anthocerotophyta. Data referring to endemic species of Brazil and of Rio de Janeiro State came from Flora e Funga do Brasil (floradobrasil.jbrj.gov.br), while conservation status followed CNCFlora (cncflora.jbrj.gov.br).

5. Conclusions

The Serra dos Órgãos National Park stands out as a key refuge for bryophyte diversity in the Atlantic Forest, hosting numerous endemic and threatened species. The park’s altitudinal variation, diverse microhabitats, and unique environmental conditions favor high species richness and endemism. These findings reinforce the ecological value of PARNASO and underscore the urgency of conserving this unique mountainous ecosystem.

Author Contributions

Methodology, M.T.A.G. and D.F.P.; Validation, D.F.P.; Investigation, J.S.d.L., A.L.A.F., M.T.A.G. and D.F.P.; Resources, D.F.P.; Data curation, J.S.d.L., A.L.A.F. and D.F.P.; Writing—original draft, A.L.A.F.; Writing—review & editing, J.S.d.L., M.T.A.G. and D.F.P. All authors have read and agreed to the published version of the manuscript.

Funding

This research was funded by the Fundação de Amparo à Pesquisa do Distrito Federal (FAP-DF).

Data Availability Statement

The original contributions presented in this study are included in the article; further inquiries can be directed to the corresponding author.

Acknowledgments

We thank the Fundação de Amparo à Pesquisa do Distrito Federal (FAP-DF) for financial support to this project and the park coordinator PARNASO (Jorge Nascimento). We fully thank the support in field collections of the colleagues Amanda Leal, Anderson Eustáquio, Clapton Moura, Cosme Ponte and Flávio Rick.

Conflicts of Interest

The authors declare no conflicts of interest.

Abbreviation

PARNASOSerra dos Órgãos National Park

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Figure 1. Substrates colonized by bryophytes in Serra dos Órgãos National Park.
Figure 1. Substrates colonized by bryophytes in Serra dos Órgãos National Park.
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Figure 2. Pictures of some species indicated as threatened in Serra dos Orgaos National Park. (A) Gongylanthus liebmannianus (Lindenb. & Gottsche) Steph., (B) Metzgeria hegewaldii Kuwah., (C) Plagiochila boryana Gottsche ex Steph., (D) Atractylocarpus longisetus (Hook.) E.B.Bartram, (E) Leptodontium wallisii (Müll.Hal.) Kindb., (F) Sphagnum perforatum Warnst.
Figure 2. Pictures of some species indicated as threatened in Serra dos Orgaos National Park. (A) Gongylanthus liebmannianus (Lindenb. & Gottsche) Steph., (B) Metzgeria hegewaldii Kuwah., (C) Plagiochila boryana Gottsche ex Steph., (D) Atractylocarpus longisetus (Hook.) E.B.Bartram, (E) Leptodontium wallisii (Müll.Hal.) Kindb., (F) Sphagnum perforatum Warnst.
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Figure 3. Location of the Serra dos Órgãos National Park, Rio de Janeiro State, Brazil.
Figure 3. Location of the Serra dos Órgãos National Park, Rio de Janeiro State, Brazil.
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Figure 4. Collection sites in Serra dos Órgãos National Park. (A) Trail of the “Mirante”, (B) aspect of the montane forest with streams, (C) “Véu da Noiva” waterfall, (D) aspect of the upper-montane forest, (E) aspect of high-altitude fields, with Sphagnum among Chusquea sp.
Figure 4. Collection sites in Serra dos Órgãos National Park. (A) Trail of the “Mirante”, (B) aspect of the montane forest with streams, (C) “Véu da Noiva” waterfall, (D) aspect of the upper-montane forest, (E) aspect of high-altitude fields, with Sphagnum among Chusquea sp.
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Table 1. Richness and diversity of bryophyte species. Symbols on the species voucher: species only known from the literature (*).
Table 1. Richness and diversity of bryophyte species. Symbols on the species voucher: species only known from the literature (*).
GroupFamilyTaxonVoucher
LiverwortsAcrobolbaceaeTylimanthus laxus (Lehm. & Lindenb.) SprucePeralta 27761 (SP)
AneuraceaeAneura pinguis (L.) Dumort.Peralta 27768 (SP)
AneuraceaeRiccardia chamedryfolia (With.) GrolleSilva, AL 515
AneuraceaeRiccardia digitiloba (Spruce ex Steph.) PagánPeralta 27496 (SP)
AneuraceaeRiccardia emarginata (Steph.) HellPeralta 27618 (SP)
AneuraceaeRiccardia fucoidea (Sw.) Schiffn.Peralta 20395 (SP)
AneuraceaeRiccardia glaziou (Spruce) MeenksGaspar 121 (R)
AneuraceaeRiccardia regnellii (Aongström.) HellPeralta 27487 (SP)
ArnelliaceaeGongylanthus liebmannianus (Lindenb. & Gott.) StephGonçalves 358 (SP)
ArnelliaceaeSouthbya organensis Herzog *Ph. v. Lützelburg 6003 p.p. (Tipo JE)
BalantiopsidaceaeIsotachis aubertii (Schwägr.) Mitt.Gaspar 148 (R)
BalantiopsidaceaeIsotachis erythrorhiza (Lehm. & Lindenb.) Steph.Peralta 27572 (SP)
BalantiopsidaceaeIsotachis multiceps (Lindenb. & Gottsche) GottscheLima 783 (SP)
BalantiopsidaceaeIsotachis serrulata (Sw.) GottscheGonçalves 396 (SP)
BalantiopsidaceaeNeesioscyphus carneus (Nees) GrollePeralta 20398 (SP)
BalantiopsidaceaeNeesioscyphus homophyllus (Nees) GrollePeralta 27722 (SP)
CalypogeiaceaeCalypogeia peruviana Nees & Mont.Peralta 27491 (SP)
CephaloziaceaeCephaloziella divaricada (Sm.) Schiffn.Gaspar 112 (R)
CephaloziaceaeCephaloziella granatensis (J.B.Jack) FulfordPeralta 27638 (SP)
CephaloziaceaeCylindrocolea planifolia (Steph.) R.M.Schust.Peralta 27621 (SP)
CephaloziaceaeKymatocalyx dominicensis (Spruce) VánaPeralta 27755 (SP)
CephaloziaceaeOdontoschisma denudatum (Nees) Dumort.Peralta 20506 (SP)
CephaloziaceaeOdontoschisma longiflorum (Taylor) Trevis. *Fulford (1976)
CephaloziellaceaeCephaloziopsis intertexta (Gottsche) R.M. Schust.Peralta 27488 (SP)
CephaloziellaceaeFuscocephaloziopsis crassifolia (Lindenb. & Gottsche) Váňa & L. Söderstr.Dias 1414 (SP)
DumortieraceaeDumortiera hirsuta Nees & Mont.Peralta 27710 (SP)
FrullaniaceaeFrullania atrata NeesFaria 1560 (SP)
FrullaniaceaeFrullania beyrichiana (Lehm. & Lindenb.) Lehm. * & Lindenb.Beyrich s.n. (holótipo W)
FrullaniaceaeFrullania brasiliensis RaddiPeralta 27631 (SP)
FrullaniaceaeFrullania cuencensis TaylorPeralta 27551 (SP)
FrullaniaceaeFrullania ericoides (Nees) Mont. *Nees (1833, as Jungermannia)
FrullaniaceaeFrullania kunzei (Lehm. & Lindb.) Lehm. & Lindb.Peralta 20492 (SP)
FrullaniaceaeFrullania obcordata (Lehm. & Lindenb.) Lehm. & Lindenb. in Gott. et al. *Winter & Schaefer-Verwimp (2020) [25]
FrullaniaceaeFrullania obscura (Sw.) Dumort.Peralta 20453 (SP)
FrullaniaceaeFrullania riojaneirensis (Raddi) Spruce *Gottsche et al. (1844) [26]
FrullaniaceaeFrullania schaefer-verwimpii Yuzawa & Hatt.Schafer-Verwimp 7375 (SP)
FrullaniaceaeFrullania setigera Steph.Peralta 27501 (SP)
FrullaniaceaeFrullania vitalii Yuzawa & Hatt.Yano 13575 (SP)
GeocalycaceaeSaccogynidium caldense (Ångstr.) GrollePeralta 27620 (SP)
HerbertaceaeHerbertus acanthelius SprucePeralta 27534 (SP)
HerbertaceaeHerbertus juniperoideus (Sw.) GrolleGissi 339 (SP)
HerbertaceaeHerbertus pensilis (T. Taylor) SprucePeralta 27612 (SP)
HerbertaceaeHerbertus sendtneri (Nees) A.Evans *Fulford (1963 as Herbertus grossispinus) [27]
HerbertaceaeTriandrophyllum subtrifidum (Hook.f. & Taylor) Fulford & Hatch. *Grolle (1964) [28]
JamesoniellaceaeSyzygiella anomala (Lindenb. & Gottsche) Steph.Lima 789 p.p. (SP)
JamesoniellaceaeSyzygiella colorata (Lehm.) K. Feldberg et al.Peralta 27575 (SP)
JamesoniellaceaeSyzygiella concreta (Gottsche) SprucePeralta 20397 (SP)
JamesoniellaceaeSyzygiella contigua Steph.Peralta 20509 (SP)
JamesoniellaceaeSyzygiella manca (Mont.) J. B. Jack & Steph.Gonçalves 389 (SP)
JamesoniellaceaeSyzygiella perfoliata (Sw.) SprucePeralta 20401 (SP)
JamesoniellaceaeSyzygiella sonderi (Gottsche) K. Feldberg, Váňa, Hentschel & HeinrichsPeralta 27502 (SP)
JamesoniellaceaeSyzygiella trigonifolia (Steph.) HerzogPeralta 27613 (SP)
JamesoniellaceaeSyzygiella uleana Steph.Peralta 20428 (SP)
JungermanniaceaeAnastrophyllum piligerum (Nees) Steph.Peralta 20380 (SP)
JungermanniaceaeAnastrophyllum tubulosum (Nees) GrollePeralta 27565 (SP)
JungermanniaceaeJungermannia amoena Lindenb. & GottscheGaspar 131 (R)
JungermanniaceaeJungermannia hyalina LyellLima 792 (SP)
JungermanniaceaeJungermannia sphaerocarpa Hook.Gaspar 122 (R)
LejeuneaceaeAcanthocoleus aberrans (Lindenb. & Gottsche) KruijtPeralta 27465 (SP)
LejeuneaceaeAnoplolejeunea conferta (C.F.W.Meissn. ex Spreng.) A.EvansPeralta 27645 (SP)
LejeuneaceaeBlepharolejeunea incongrua (Lindenb. & Gottsche) van Slageren & KruijtPeralta 27577 (SP)
LejeuneaceaeBrachiolejeunea phyllorhiza (Nees) Kruijt & Gradst. *Gottsche et al. (1844)
LejeuneaceaeBryopteris diffusa (Sw.) Nees *Stotler & Crandall (1974) [29]
LejeuneaceaeBryopteris filicina (Sw.) NeesPeralta 27530 (SP)
LejeuneaceaeCeratolejeunea rubiginosa Gottsche ex Steph.Peralta 27681 (SP)
LejeuneaceaeCheilolejeunea acutangula (Nees) GrollePeralta 27523 (SP)
LejeuneaceaeCheilolejeunea beyrichii (Lindenb.) E. ReinerPeralta 27492 (SP)
LejeuneaceaeCheilolejeunea filiformis (Sw.) W. Ye, R.L. Zhu & Gradst. *Gottsche et al. (1844, as Omphalanthus) [26]
LejeuneaceaeCheilolejeunea grosseoleosa C.J.Bastos & Schäf.-Verw.Peralta 27549 p.p. (SP)
LejeuneaceaeCheilolejeunea holostipa (Spruce) Grolle & R.-L.ZhuAlkimim-Faria 1486
LejeuneaceaeCheilolejeunea unciloba (Lindenb.) MalombeLima 750 (SP)
LejeuneaceaeCheilolejeunea xanthocarpa (Lehm. & Lindenb.) MalombePeralta 27504 (SP)
LejeuneaceaeCololejeunea cardiocarpa (Mont.) A.EvansPeralta 27687 p.p. (SP)
LejeuneaceaeCololejeunea diaphana A.EvansPeralta 27687 p.p. (SP)
LejeuneaceaeCololejeunea jamesii M.E.Reiner & PócsPeralta 20511 p.p. (SP)
LejeuneaceaeCololejeunea microscopica (Taylor) Schiffn.Peralta 27694 (SP)
LejeuneaceaeCololejeunea papilliloba (Steph). Steph.Peralta 27695 (SP)
LejeuneaceaeColura itatyana Steph.Gaspar 138 (R)
LejeuneaceaeColura tenuicornis (A.Evans) Steph. *Jovet-Ast (1953) [30]
LejeuneaceaeCyclolejeunea luteola (Spruce) Grolle & R.-L.ZhuSchafer-Verwimp 7416 (SP)
LejeuneaceaeDibrachiella parviflora (Nees) X.Q. Shi, R.L. Zhu & Gradst.Silva, AL 500
LejeuneaceaeDicranolejeunea axilaris (Nees & Mont.) Schiffn.Peralta 27735 (SP)
LejeuneaceaeDiplasiolejeunea alata Jovet-AstPeralta 27593 p.p. (SP)
LejeuneaceaeDiplasiolejeunea unidentata (Lehm. & Lindenb.) Schiffn. *Gradstein & Costa (2003) [31]
LejeuneaceaeDrepanolejeunea anoplanta (Spruce) Steph.Peralta 27659 (SP)
LejeuneaceaeDrepanolejeunea araucariae Steph.Gaspar 135 (R)
LejeuneaceaeDrepanolejeunea bidens (Steph.) A.EvansGaspar 117 (R)
LejeuneaceaeDrepanolejeunea fragilis Bischl.Peralta 27593 p.p. (SP)
LejeuneaceaeDrepanolejeunea lichenicola (Spruce) Steph. *Bischler (1964) [32]
LejeuneaceaeDrepanolejeunea mosenii (Steph.) Bischl.Peralta 27639 (SP)
LejeuneaceaeFrullanoides tristis van SlagerenPeralta 27646 (SP)
LejeuneaceaeHaplolejeunea umbrosa Gradst. & Ilkiu-BorgesPeralta 27495 (SP)
LejeuneaceaeHarpalejeunea oxyphylla (Nees & Mont.) Steph.Gaspar 124 (R)
LejeuneaceaeHarpalejeunea schiffneri S.W. ArnellPeralta 27462 (SP)
LejeuneaceaeHarpalejeunea stricta (Lindenb. & Gottsche) Steph.Peralta 27526 (SP)
LejeuneaceaeHarpalejeunea subacuta A.EvansPeralta 20450 p.p. (SP)
LejeuneaceaeLejeunea acanthogona SprucePeralta 27463 (SP)
LejeuneaceaeLejeunea aphanes SpruceLima 750 (SP)
LejeuneaceaeLejeunea bermudiana (A.Evans) R.M.Schust.Peralta 27719 (SP)
LejeuneaceaeLejeunea beyrichiana (Steph) Gradst. & C.J.Bastos *Beyrich s.n. (Tipo NY)
LejeuneaceaeLejeunea cerina (Lehm. & Lindenb.) GottscheSchaefer-Verwimp 7369 (SP)
LejeuneaceaeLejeunea flaccida Lindenb. & GottschePeralta 27512 (SP)
LejeuneaceaeLejeunea flava (Sw.) NeesPeralta 27505 (SP)
LejeuneaceaeLejeunea glaucescens GottschePeralta 27476 (SP)
LejeuneaceaeLejeunea grossitexta (Steph.) E.Reiner & Goda *Schäfer-Verwimp & Vital (1989) [33]
LejeuneaceaeLejeunea laetevirens Nees & Mont.Peralta 27464 (SP)
LejeuneaceaeLejeunea longidentata C.J.Bastos, Gradst., S. Vilas Bôas-Bastos & Schäf.-Verw.Peralta 27543 (SP)
LejeuneaceaeLejeunea oligoclada SprucePeralta 27498 p.p. (SP)
LejeuneaceaeLejeunea pterigonia (Lehm. & Lindenb.) Mont.Peralta 27525 (SP)
LejeuneaceaeLejeunea raddiana Lindenb.Peratla 20484 (SP)
LejeuneaceaeLejeunea serpillifolioides (Raddi) Gradst.Peralta 27611 (SP)
LejeuneaceaeLejeunea subplana (Steph.) C.J.BastosGonçalves 306 p.p. (SP)
LejeuneaceaeMarchesinia brachiata (Sw.) Schiffn.Peralta 27508 (SP)
LejeuneaceaeMetalejeunea cuculata (Reinw. et al.) GrolleYano 13558 (SP)
LejeuneaceaeMicrolejeunea cystifera HerzogPeralta 27482 (SP)
LejeuneaceaeMicrolejeunea epiphylla Bischl.Gonçalves 306 p.p. (SP)
LejeuneaceaeMicrolejeunea squarrosa J. Heinrichs, A. Schäfer-Verwimp, Pócs & S.S. DongPeralta 27671 p.p. (SP)
LejeuneaceaeMicrolejeunea stricta (Lindenb. & Gottsche) Steph.Lima 782 p.p. (SP)
LejeuneaceaeNeurolejeunea breutelii (Gottsche) A.EvansLima 767 p.p. (SP)
LejeuneaceaeOtigoniolejeunea huctumalcensis (Lindenb. & Gottsche) Y.M. Wei, R.L. Zhu & Gradst.Peralta 27474 (SP)
LejeuneaceaePrionolejeunea aemula (Gottsche) A.EvansPeralta 27727 (SP)
LejeuneaceaeSchiffneriolejeunea polycarpa (Nees) Gradst.Peralta 20468 (SP)
LejeuneaceaeVitalianthus bischelerianus (Porto & Grolle) R.M.Schust. & Giancotti *Gottsche et al. (1844, as Phragmicoma juliformis)
LepidoziaceaeBazzania aurescens SprucePeralta 20394 (SP)
LepidoziaceaeBazzania hookeri (Lindenb.) Trevis.Fulford (1963)
LepidoziaceaeBazzania longistipula (Lindenb.) Trevis.Peralta 27480 (SP)
LepidoziaceaeBazzania nitida (Weber) GrollePeralta 27529 (SP)
LepidoziaceaeBazzania stolonifera (Sw.) Trevis. *Nees (1833, as Jungermannia) [34]
LepidoziaceaeKurzia capillaris (Sw.) GrolleLima 780 (SP)
LepidoziaceaeLepidozia coilophylla TaylorPeralta 27532 (SP)
LepidoziaceaeLepidozia cupressina (Sw.) Lindenb.Gonçalves 311 (SP)
LepidoziaceaeLepidozia inaequalis (Lehm. & Lindenb.) Lehm. & Lindenb.Peralta 20402 (SP)
LepidoziaceaeLepidozia incurvata Lindenb.Lima 784 (SP)
LepidoziaceaeLepidozia macrocolea SprucePeralta 27539 (SP)
LepidoziaceaeMicropterygium leiophyllum Spruce *Gottsche et al. (1844, as M. vulgare)
LepidoziaceaeMicropterygium pterygophyllum (Nees) Trevis.Nees (1833, as Jungermannia)
LepidoziaceaeParacromastigum dusenii R.M.Schust. *Dias 1310 (SP)
LepidoziaceaeParacromastigum pachyrhizum (Nees) FulfordSchafer-Verwimp 7421 (SP)
LepidoziaceaeTelaranea nematodes (Gottsche ex Austin) M.A.HoweLima 754 (SP)
LophocoleaceaeLophocolea muricata (Lehm.) Nees in Gottsche et al.Peralta 27508 (SP)
LophocoleaceaeClasmatocolea vermicularis (Lehm.) GrollePeralta 27566 (SP)
LophocoleaceaeCryptolophocolea martiana (Nees) L.Soderstr. et al. subsp. martianaLima 768 (SP)
LophocoleaceaeHeteroscyphus combinatus (Nees) Schiffn.Peralta 27556 (SP)
LophocoleaceaeLeptoscyphus amphibolius (Nees) GrollePeralta 27583 (SP)
LophocoleaceaeLeptoscyphus porphyrius (Nees) GrollePeralta 27628 (SP)
LophocoleaceaeLeptoscyphus spectabilis (Steph.) GrollePeralta 27658 (SP)
LophocoleaceaeLophocolea bidentata (L.) Dumort.Peralta 20420 (SP)
LophocoleaceaeLophocolea connata (Sw.) Nees *Nees (1833, as Jungermannia)
LophocoleaceaeLophocolea mandonii Steph.Peralta 20379 (SP)
MarchantiaceaeMarchantia chenopoda L.Vital 1197 (SP)
MarchantiaceaeMarchantia polymorpha L.Schwacke, C.A.W. s.n. (R)
MarchantiaceaeMarchantia paleacea Bert.Peralta 20475 (SP)
MetzgeriaceaeMetzgeria acuminata Steph.Peralta 27471 (SP)
MetzgeriaceaeMetzgeria albinea SprucePeralta 27475 (SP)
MetzgeriaceaeMetzgeria brasiliensis Schiffn.Dias 1234 (SP)
MetzgeriaceaeMetzgeria ciliata Raddi *Raddi (1823) [35]
MetzgeriaceaeMetzgeria conjugata Lindb.Silva, AL 517 (UB)
MetzgeriaceaeMetzgeria consanguinea Schiffn.Peralta 27528 (SP)
MetzgeriaceaeMetzgeria convoluta Steph. *Costa (2008) [36]
MetzgeriaceaeMetzgeria cratoneura Schiffn. *Costa (2008) [36]
MetzgeriaceaeMetzgeria decipiens (C. Massal.) Schiffn.Eiten 7105 (SP)
MetzgeriaceaeMetzgeria fruticola SprucePeralta 27715 (SP)
MetzgeriaceaeMetzgeria furcata (L.) Dumort. *Bartram (1954) Gonçalves 326 (SP)
MetzgeriaceaeMetzgeria hegewaldii Kuwah.Peralta 27466 (SP)
MetzgeriaceaeMetzgeria lechlerii Steph. *Costa et al. (2005) [37]
MetzgeriaceaeMetzgeria myriopoda Lindb.Yano 13561 (SP)
NoterocladaceaeNoteroclada confluens (Hook.f. & Taylor) SprucePeralta 20471 (SP)
PallaviciniaceaeJensenia spinosa (Lindenb. & Gottsche) GrolleGonçalves 400 (SP)
PallaviciniaceaePallavicinia lyelii (Hook.) S.F.GrayGonçalves 186 (SP)
PallaviciniaceaeSymphyogyna brasiliensis (Nees) Nees & Mont.Peralta 27750 (SP)
PallaviciniaceaeSymphyogyna podophylla (Thunb.) Mont. & NeesSilva, AL 502
PlagiochilaceaePlagiochila amicta Steph. *Heinrichs (2002) [38]
PlagiochilaceaePlagiochila bicuspidata GottschePeralta 10518 (SP)
PlagiochilaceaePlagiochila bifaria (Sw.) Lindenb.Peralta 27516 (SP)
PlagiochilaceaePlagiochila boryana Gottsche ex Steph.Peralta 27749 (SP)
PlagiochilaceaePlagiochila corrugata (Nees) Nees & Mont.Peralta 20376 (SP)
PlagiochilaceaePlagiochila crispabilis Lindenb.Peralta 20511 (SP)
PlagiochilaceaePlagiochila cristata (Sw.) Lindenb.Schafer-Verwimp 7401 (SP)
PlagiochilaceaePlagiochila deflexirama TaylorPeralta 27721 (SP)
PlagiochilaceaePlagiochila exigua (Taylor) TaylorPeralta 20468 (SP)
PlagiochilaceaePlagiochila flabelliflora Steph.Gonçalves 334 (SP)
PlagiochilaceaePlagiochila gymnocalycina (Lehm. & Lindenb.) Lindenb.Dias 1397 (SP)
PlagiochilaceaePlagiochila loriloba Herzog ex CarlPerlata 27519 (SP)
PlagiochilaceaePlagiochila macrotachya Lindenb.Peralta 27700 (SP)
PlagiochilaceaePlagiochila martiana (Nees) Lindenb.Peralta 20385 (SP)
PlagiochilaceaePlagiochila montagnei Nees & Mont. *Heinrichs & Gradstein (2000) [39]
PlagiochilaceaePlagiochila pseudoradicans Herzog *Ph. v. Lützelburg 6029b (type JE)
PlagiochilaceaePlagiochila punctata (Taylor) TaylorLima 783 p.p. (SP)
PlagiochilaceaePlagiochila rutilans Lindenb. *Heinrichs (2002) [38]
PlagiochilaceaePlagiochila simplex (Sw.) Lindenb.Peralta 27780 (SP)
PlagiochilaceaePlagiochila subbidentata TaylorPeralta 27586 (SP)
PlagiochilaceaePlagiochila superba d’OrbignyPeralta 20457 (SP)
PorellaceaePorella brasiliensis (Raddi) Schiffn.Peralta 27724 (SP)
PorellaceaePorella reflexa (Lehm. & Lindenb.) Trevis.Yano 13580 (SP)
RadulaceaeRadula angulata Steph.Peralta 27697 (SP)
RadulaceaeRadula javanica GottschePeralta 27527 (SP)
RadulaceaeRadula ligula Steph.Peralta 20407 (SP)
RadulaceaeRadula nudicaulis Steph.Gonçalves 408 (SP)
RadulaceaeRadula pallens (Sw.) Nees ex MontLima 759 (SP)
RadulaceaeRadula recubans TaylorGaspar 142 (R)
RadulaceaeRadula schaefer-verwimpii K.YamadaPeralta 27489 (SP)
RadulaceaeRadula stenocalyx Mont.Peralta 27691 (SP)
RadulaceaeRadula subinflata Lindenb. & GottscheGonçalves 322 (SP)
RadulaceaeRadula tectiloba Steph.Peralta 27507 (SP)
RadulaceaeRadula voluta Taylor ex Gottsche, Lindenb. & NeesPeralta 20429 (SP)
ScapaniaceaeScapania portoricensis Hampe & GottschePeralta 27541 (SP)
TrichocoleaceaeTrichocolea brevifissa Steph.Peralta 27503 (SP)
MossesAdelotheciaceaeAdelothecium bogotense (Hampe) Mitt.Peralta 27692 (SP)
AndreaceaeAndreaea acutifolia Hook.f. & Wilson *Müller (1845) Gonçalves 401 (SP)
AndreaceaeAndreaea rupestris Hedw.Gaspar 137 (R)
AndreaceaeAndreaea subulata Harv. *Lützelburg, P. von 7041 (tipo JE of Andreaea hamulata)
BartramiaceaeBartramia mathewsii Mitt.Peralta 27606 (SP)
BartramiaceaeBreutelia grandis (Hampe) ParisPeralta 27559 (SP)
BartramiaceaeBreutelia tomentosa (Sw. ex Brid.) A.JaegerLima 798 (SP)
BartramiaceaeLeiomela bartramioides (Hook.) ParisPeralta 27500 (SP)
BartramiaceaeLeiomela piligera (Hampe) Broth. *Brotherus (1924) [40]
BartramiaceaePhilonotis cernua (Wilson) Griffin & W.R.BuckPeralta 20426 (SP)
BartramiaceaePhilonotis hastata (Duby) Wijk & Margad.Alkimim-Faria, 1513
BartramiaceaePhilonotis uncinata (Schwägr.) Brid. *Brotherus (1894)
BrachytheciaceaeAerolindigia capillacea (Hornsch.) M.MenzelPeralta 20418 (SP)
BrachytheciaceaeMeteoridium remotifolium (Müll.Hal.) ManuelPeralta 27763 (SP)
BrachytheciaceaeRhynchostegium scariosum (Taylor) A.JaegerPeralta 20419 (SP)
BrachytheciaceaeRhynchostegium serrulatum (Hedw.) A.JaegerGonçalves 333 p.p. (SP)
BrachytheciaceaeSquamidium brasiliense Broth. *Allen & Crosby (1986) [41]
BrachytheciaceaeSquamidium leucotrichum (Taylor) Broth.Peralta 27486 (SP)
BrachytheciaceaeZelometeorium ambiguum (Hornsch.) Manuel *Manuel (1977) [42]
BrachytheciaceaeZelometeorium patulum (Hedw.) ManuelGonçalves 332 (SP)
BruchiaceaeTrematodon longicollis Michx. *Brotherus (1924b) [43]
BryaceaeAnomobryum julaceum (Schrad. ex P.Gaertn. et al.) Schimp.Peralta 20377 (SP)
BryaceaeBrachymenium acuminatum Harv.Dias 1372 (SP)
BryaceaeBrachymenium consimile (Mitt.) A.JaegerPeralta 27654 (SP)
BryaceaeBrachymenium hornschuchianum Mart. *Brotherus (1894) [44]
BryaceaeBrachymenium radiculosum (Schwägr.) Hampe *Brotherus (1924b) [43]
BryaceaeBrachymenium systilium (Müll.Hal.) A.JaegerPeralta 27673 (SP)
BryaceaeBryum argenteum Hedw.Vital 1191 (SP)
BryaceaeBryum atenense WilliamsAlkimim-Faria 1246 (UB)
BryaceaeBryum billarderii Schwaegr.Dias 1306a (SP)
BryaceaeBryum capillare Hedw.Peralta 20465 (SP)
BryaceaeBryum densifolium Brid. *Bartram (1954, as B. gracilescens)
BryaceaeBryum huillense Welw. & DubyPeralta 20415 (SP)
BryaceaeBryum limbatum Müll.Hal *Gonçalves 384 (SP)
BryaceaeBryum pabstianum Müll.Hal. *Hampe (1879) [45]
BryaceaeBryum subapiculatum HampePeralta 20462 (SP)
BryaceaeRhodobryum aubertii (Schwägr.) Thér.Peralta 27762 (SP)
BryaceaeRhodobryum beyrichianum (Hornsch.) Müll.Hal.Peralta 20514 (SP)
BryaceaeRhodobryum roseum (Hedw.) Limpr.Peralta 27582 (SP)
BryaceaeRhodobryum subverticillatum Broth. *Bartram (1952) [46]
BryaceaeSchoenobryum concavifolium (Griff.) Gangulee *Bartram (1954) [19]
CalymperaceaeCalymperes nicaraguense Renauld & CardotAlkimim-Faria 1256
CalymperaceaeSyrrhopodon gaudichaudii Mont. *Bartram (1954) [19]
CalymperaceaeSyrrhopodon helicophyllus Mitt.Alkimim-Faria, AL 1254
CalymperaceaeSyrrhopodon ligulatus Mont.Peralta 27490 (SP)
CalymperaceaeSyrrhopodon prolifer Schwägr.Peralta 27473 (SP)
CatagoniaceaeCatagonium brevicaudatum Müll.Hal. ex Broth.Peralta 27630 (SP)
DaltoniaceaeCalyptrochaeta albescens (Hampe) W.R.BuckLima 775 (SP)
DaltoniaceaeCalyptrochaeta setigera (Mitt.) W.R.BuckPeralta 27702 (SP)
DaltoniaceaeDaltonia splachnoides (Sm.) Hook. & TaylorPeralta 27546 (SP)
DaltoniaceaeLeskeodon aristatus (Geh. & Hampe) Broth.Peralta 27517 (SP)
DicranaceaeAongstroemia julacea (Hook.) Mitt.Peralta 27550 (SP)
DicranaceaeAtractylocarpus brasiliensis (Müll.Hal.) R.S.WilliamsPeralta 20490 (SP)
DicranaceaeAtractylocarpus longisetus (Hook.) E.B.Bartram *Bartram (1954) [19]
DicranaceaeDicranella harrisii Müll.Hal.) Broth.Schafer-Verwimp 13101 (SP)
DicranaceaeDicranella hilariana (Mont.) Mitt. *Luetzelburg (1923, as D. trematodontifolia) [47]
DicranaceaeDicranella subsulcata (Hampe) HampeSchafer-Verwimp 7422 (SP)
DicranaceaeDicranodontium pulchroalare Broth. *Lützelburg 1519 (holótipo JE)
DicranaceaeDicranoloma billardieri (Brid. ex Anon) ParisPeralta 27597 (SP)
DicranaceaeDicranum frigidum Müll.Hal.Dias 1361 (SP)
DicranaceaeHolomitrium crispulum Mart.Peralta 27599 (SP)
DicranaceaeLeucoloma cruegerianum (Müll.Hal.) A.JaegerPeralta 27678 (SP)
DicranaceaeLeucoloma serrulatum Brid.Peralta 27701 (SP)
DicranaceaeLeucoloma triforme (Mitt.) A.JaegerGonçalves 190 (SP)
DicranaceaeMicrocampylopus curvisetus (Hampe) Giese & J.-P.FrahmSchafer-Verwimp 7422 p.p. (SP)
DicranaceaeParaleucobryum longifolium (Hedw.) LoeskeDias 1325 (SP)
DicranaceaePilopogon guadalupensis (Brid.) J.-P.FrahmPeralta 1558 (SP)
DiphysciaceaeDiphyscium longifolium Griff.Peralta 27723 (SP)
DitrichaceaeCeratodon purpureus (Hedw.) Brid.Dias 1315 (SP)
DitrichaceaeCladastomum robustum Broth.Lima 485 (SP)
DitrichaceaeCladastomum ulei Müll.Hal.Peralta 27635 (SP)
DitrichaceaeCrumuscus vitalis W.R.Buck & SniderLima 791 (SP)
DitrichaceaeDitrichum crinale (Taylor) KuntzeLima 801 (SP)
DitrichaceaeDitrichum paulense Geh. ex HampePeralta 27656 (SP)
EntodontaceaeEntodon hampeanus Müll.Hal.Peralta 27614 (SP)
EntodontaceaeEntodon jamesonii (Taylor) Mitt.Peralta 27580 (SP)
EntodontaceaeErythrodontium longisetum (Hook.) Paris *Vattimo-Gil & Vattimo (1980) [48]
EntodontaceaeErythodontium squarrosum (Hampe) Paris *Brotherus (1924)
EphemeraceaeMicromitrium tenerum (Bruch & Schimp.) CrosbyAlkimim-Faria, AL 1379 (UB)
FabroniaceaeDimerodontium pellucidum Schwägr.Schafer-Verwimp 13113 (SP)
FabroniaceaeFabronia ciliaris (Brid.) Brid.Gonçalves 182 (SP)
FissidentaceaeFissidens anguste-limbatus Mitt.Peralta 27665 (SP)
FissidentaceaeFissidens asplenioides Hedw.Peralta 27709 (SP)
FissidentaceaeFissidens bryoides Hedw.Lima 787 (SP)
FissidentaceaeFissidens crispus Mont.Gonçalves 180 (SP)
FissidentaceaeFissidens elegans Brid.Peralta 27479 (SP)
FissidentaceaeFissidens flabellatus Hornsch. *Beyrich s.n. (lectótipo BM)
FissidentaceaeFissidens intromarginatus (Hampe) Mitt. *Ule 1225 (lectotype H, tipo de F. hemibryoides)
FissidentaceaeFissidens lagenarius Mitt. *Bartram (1954)
FissidentaceaeFissidens oblongifolius Hook.f. & WilsonPeralta 20464 (SP)
FissidentaceaeFissidens oediloma Müll.Hal. ex Broth.Peralta 27571 (SP)
FissidentaceaeFissidens pellucidus Hornsch.Peralta 27538 (SP)
FissidentaceaeFissidens prionodes Mont.Peralta 27740 (SP)
FissidentaceaeFissidens radicans Mont.Peralta 27737 (SP)
FissidentaceaeFissidens rigidulus Hook.f. & WilsonPeralta 27756 (SP)
FissidentaceaeFissidens saprofilus Broth.Lima 754 (SP)
FissidentaceaeFissidens scariosus Mitt.Lima 756 (SP)
FissidentaceaeFissidens semicompletus Hedw.Alkimim-Faria 1228
FissidentaceaeFissidens taxifolius Hedw.Alkimim-Faria 1367
FissidentaceaeFissidens wallisii Müll.Hal. ex Broth.Peralta 27675 (SP)
FissidentaceaeFissidens weirii Mitt.Peralta 27484 (SP)
FunariaceaeEntosthodon bonplandii (Hook.) Mitt.Schaefer-Verwimp 7405 (SP)
FunariaceaeFunaria hygrometrica Hedw. *Müller (1845) [49]
GrimmiaceaeGrimmia laevigata (Brid.) Brid.Dias 1298 (SP)
GrimmiaceaeGrimmia longirostris Hook.f. & WilsonGaspar 110 (R)
GrimmiaceaeRacomitrium subsecundum (Hook. & Grev. ex Harv.) Mitt.Dias 1405 (SP)
HedwigiaceaeHedwigidium rhabdocarpum (Hampe) A.JaegerFaria 1547 (SP)
HookeriaceaeHookeria acutifolia Hook. & Grev.Peralta 27739 (SP)
HookeriaceaePhilophyllum tenuifolium (Mitt.) Broth.Peralta 27581 (SP)
HypnaceaeChrysohypnum diminutivum (Mitt.) Broth.Peralta 20478 (SP)
HypnaceaeEctropothecium leptochaeton (Schwägr.) W.R.BuckPeralta 20461 (SP)
HypnaceaeMittenothamnium reduncum (Mitt.) OchyraPeralta 27624 (SP)
HypnaceaeMittenothamnium reptans (Hedw.) CardotEiten 7133 (SP)
HypnaceaeTaxiphyllum taxirameum (Mitt.) M. Fleisch.Gonçalves 194 (SP)
HypnaceaeVesicularia vesicularis (Schwägr.) Broth.Alkimim-Faria, 1364
HypopterygiaceaeHypopterygium tamariscinum (Hedw.) Brid.Yano 13560 (SP)
HypopterygiaceaeLopidium concinnum (Hook.) WilsonPeralta 27712 (SP)
LembophyllaceaeOrthostichella versicolor (Müll.Hal.) B.H.Allen & W.R.BuckPeralta 27520 (SP)
LeucobryaceaeCampylopus aemulans (Hampe) A.Jaeger *Frahm (1975) [50]
LeucobryaceaeCampylopus arctocarpus (Hornsch.) Mitt.Peralta 27547 (SP)
LeucobryaceaeCampylopus cryptopodioides Broth. *Frahm (1991) [15]
LeucobryaceaeCampylopus cuspidatus (Hornsch.) Mitt. *Frahm (1975) [50]
LeucobryaceaeCampylopus densicoma (Müll.Hal.) ParisGonçalves 409 (SP)
LeucobryaceaeCampylopus dichrostis (Müll.Hal.) ParisGonçalves 330 (SP)
LeucobryaceaeCampylopus filifolius (Müll.Hal.) ParisLima 789 (SP)
LeucobryaceaeCampylopus fragilis (Brid.) Bruch & Schimp.Peralta 27610 (SP)
LeucobryaceaeCampylopus gemmatus (Müll.Hal.) ParisVital 7634 (SP)
LeucobryaceaeCampylopus heterostachys (Hampe) A.JaegerDias 1299 (SP)
LeucobryaceaeCampylopus introflexus (Hedw.) Brid.Alkimim-Faria 1510
LeucobryaceaeCampylopus julicaulis Broth.Gaspar 104 (R)
LeucobryaceaeCampylopus lamellinervis (Müll.Hal.) Mitt.Müller 1845
LeucobryaceaeCampylopus occultus Mitt.Frahm sn (SP147028)
LeucobryaceaeCampylopus pilifer Brid.Peralta 27605 (SP)
LeucobryaceaeCampylopus pyriformis (Schultz) Brid.Gaspar 114 (R)
LeucobryaceaeCampylopus richardii Brid.Dias 1324 (SP)
LeucobryaceaeCampylopus savannarum (Müll.Hal.) Mitt.Gaspar 139 (R)
LeucobryaceaeCampylopus subcuspidatus (Hampe) A.JaegerPeralta 27641 (SP)
LeucobryaceaeCampylopus thwaitesii (Mitt.) A.JaegerGonçalves 183 (SP)
LeucobryaceaeCampylopus trachyblepharon (Müll.Hal.) Mitt.Paula sn (SP463831)
LeucobryaceaeCampylopus uleanus (Müll.Hal.) Broth.Dias 1384 (SP)
LeucobryaceaeCampylopus widgrenii (Müll.Hal.) Mitt.Vital 1189 (SP)
LeucobryaceaeLeucobryum albicans (Schwägr.) Lindb.Peralta 27522 (SP)
LeucobryaceaeLeucobryum clavatum Hampe *Luetzelburg 1923 [47]
LeucobryaceaeLeucobryum crispum Müll.Hal.Dias 1235 (SP)
LeucobryaceaeLeucobryum giganteum Müll.Hal. *Herzog (1926) [51]
LeucobryaceaeLeucobryum sordidum Ångström *Luetzelburg (1923) [47]
LeucobryaceaeOchrobryum gardneri (Müll.Hal.) Lindb.Alkimim-Faria 1364 (UB)
LeucobryaceaeOctoblepharum albidum Hedw. *Bartram (1954) [19]
LeucomiaceaeLeucomium strumosum (Hornsch.) Mitt.Peralta 27769 (SP)
MeteoriaceaeMeteorium deppei (Hornsch.) Mitt.Peralta 27573 (SP)
MeteoriaceaeMeteorium nigrescens (Hedw.) Dozy & Molk. *Brotherus (1924)
MeteoriaceaeMeteorium pseudoteres W.R.BuckPeralta 27668 (SP)
MeteoriaceaePilotrichella flexilis (Hedw.) ÅngströmPeralta 20491 (SP)
MeteoriaceaeToloxis imponderosa (Taylor) W.R.BuckPeralta 20517 (SP)
MniaceaeEpipterygium immarginatum Mitt.Peralta 20432 (SP)
MniaceaePlagiomnium rhynchophorum (Hook.) T.J.Kop.Peralta 20411 p.p. (SP)
MniaceaePohlia camptotrachela (Renauld & Cardot) Broth.Peralta 20433 (SP)
MniaceaePohlia elongata Hedw.Lima 790 (SP)
MniaceaePohlia papilosa (Müll.Hal. ex A.Jaeger) Broth.Faria 1554 (SP)
MniaceaePohlia tenuifolia (A.Jaeger) Broth.Peralta 27579 (SP)
MniaceaeSchizymenium campylocarpum (Arn. & Hook.) A.J.ShawLima 761 (SP)
MniaceaeSchizymenium pusillum (Hook.f. & Wilson) A.J.ShawDias 1382 (SP)
NeckeraceaeHomalia glabella (Hedw.) Schimp.Gonçalves 310 (SP)
NeckeraceaeHomaliodendron flabellatum (Sm.) M.Fleisch.Peralta 27569 (SP)
NeckeraceaeNeckera ehrenbergii Müll.Hal.Peralta 27588 (SP)
NeckeraceaePorotrichum korthalsianum (Dozy & Molk.) Mitt.Dias 1420 (SP)
NeckeraceaePorotrichum leucocaulon Müll.Hal.Peralta 27674 (SP)
NeckeraceaePorotrichum longirostre (Hook.) Mitt.Peralta 20387 p.p. (SP)
NeckeraceaePorotrichum superbum (Taylor) HampePeralta 27585 (SP)
NeckeraceaePorotrichum thieleanum (Müll.Hal.) Mitt.Peralta 20383 (SP)
NeckeraceaeThamnobryum fasciculatum (Hedw.) I.SastrePeralta 27703 (SP)
OrthotrichaceaeMacrocoma orthotrichoides (Raddi) Wijk & Margad.Schafer-Verwimp 7449 (SP)
OrthotrichaceaeMacromitrium catharinense Paris *Li et al. (2019) [52]
OrthotrichaceaeMacromitrium cirrosum (Hedw.) Brid. *Bartram (1954)
OrthotrichaceaeMacromitrium microstomum (Hook. & Grev.) Schwägr.Gonçalves 395 (SP)
OrthotrichaceaeOrthotrichum papillosum (Hampe) Lesquereux & JamesAlkimim-Faria, 1236
OrthotrichaceaeSchlotheimia appressifolia Mitt.Atwood 2007
OrthotrichaceaeSchlotheimia capillaris HampePeralta 20376 p.p. (SP)
OrthotrichaceaeSchlotheimia rugifolia (Hook.) Schwägr.Lima 753 (SP)
OrthotrichaceaeSchlotheimia tecta Hook.f. & WilsonGonzatti 4003 (SP)
OrthotrichaceaeZygodon reiwardtii (Hornsch.) A.BraunPeralta 27629 (SP)
PhylogoniaceaePhyllogonium viride Brid.Gonçalves 320 (SP)
PilotrichaceaeCallicostella martiana (Hornsch.) A.JaegerGonçalves 197 (SP)
PilotrichaceaeCallicostella pallida (Hornsch.) ÅngströmPeralta 27689 (SP)
PilotrichaceaeCyclodictyon albicans (Hedw.) KuntzePeralta 27587 (SP)
PilotrichaceaeCyclodictyon marginatum (Hook. & Wilson) KuntzeSchafer-Verwimp 13124 (SP)
PilotrichaceaeCyclodictyon varians (Sull.) KuntzePeralta 27619 (SP)
PilotrichaceaeLepidopilidium brevisetum (Hampe) Broth.Silva, AL 481A (UB)
PilotrichaceaeLepidopilidium nitens (Hornsch.) Broth.Peralta 27764 (SP)
DaltoniaceaeLepidopilum affine Müll.Hal. ex Broth.Alkimim-Faria, 1360 (UB)
PilotrichaceaeLepidopilum muelleri (Hampe) Mitt.Peralta 20409 (SP)
PilotrichaceaeLepidopilum scabrisetum (Schwägr.) SteerePeralta 27666 (SP)
PilotrichaceaeLepidopilum subsubulatum Geh. & HampePeralta 27521 (SP)
PilotrichaceaeThamniopsis incurva (Hornsch.) W.R.BuckYano 13564 (SP)
PilotrichaceaeThamniopsis langsdorffii (Hook.) W.R.BuckGonçalves 307 (SP)
PilotrichaceaeTrachyxiphium guadalupense (Brid.) W.R.BuckPeralta 27497 p.p. (SP)
PlagiotheciaceaePlagiothecium novogranatense (Hampe) Mitt.Peralta 27570 (SP)
PolytrichaceaeAtrichum androgynum (Müll.Hal.) A.JaegerGonçalves 195 (SP)
PolytrichaceaeItatiella ulei (Broth. ex Müll.Hal.) G.L.Sm.Peralta 27634 (SP)
PolytrichaceaeOligotrichum riedelianum Mitt.Peralta 27594 (SP)
PolytrichaceaePogonatum campylocarpum (Müll.Hal.) Mitt.Peralta 20439 (SP)
PolytrichaceaePogonatum pensilvanicum (E.B.Bartram ex Hedw.) P.Beauv.Paula, J.C. sn (SP463829)
PolytrichaceaePogonatum perichaetiale (Mont.) A.JaegerAlkimim-Faria 1236 (UB)
PolytrichaceaePolytrichadelphus pseudopolytrichum (Raddi) G.L.Sm.Eiten 7156 (SP)
PolytrichaceaePolytrichum angustifolium Mitt.Peralta 20378 (SP)
PolytrichaceaePolytrichum commune L. ex Hedw.Goeldi sn (SP219763)
PolytrichaceaePolytrichum juniperinum Willd. ex Hedw.Faria 1562 (SP)
PottiaceaeBarbula indica (Hook.) Spreng.Peralta 27774 (SP)
PottiaceaeChionoloma angustatum (Mitt.) M.MenzelPeralta 27677 (SP)
PottiaceaeChionoloma fractum M.J.Cano, J.A.Jiménez & M.AlonsoPeralta 27589 (SP)
PottiaceaeHyophila involuta (Hook.) A.JaegerSilva, AL 499 (UB)
PottiaceaeHyophiladelphus agrarius (Hedw.) R.H.Zander *Bartram (1954)
PottiaceaeLeptodontium araucarieti (Müll.Hal.) ParisGonçalves 413 (SP)
PottiaceaeLeptodontium filicicola HerzogPeralta 27669 (SP)
PottiaceaeLeptodontium flexifolium (Dicks.) HampeDias 1276 (SP)
PottiaceaeLeptodontium pungens (Mitt.) Kindb.Dias 1377 (SP)
PottiaceaeLeptodontium stellatifolium (Hampe) Broth.Peralta 27595 (SP)
PottiaceaeLeptodontium viticulosoides (P.Beauv.) Wijk & Margad.Peralta 27607 (SP)
PottiaceaeLeptodontium wallisii (Müll.Hal.) Kindb.Gonçalves 364 (SP)
PottiaceaeStreptopogon calymperes Müll.Hal.Peralta 27450 (SP)
PottiaceaeTortella humilis (Hedw.) Jenn.Peralta 20417 (SP)
PottiaceaeTortella tortuosa (Hedw.) Limpr.Lima 806 (SP)
PriodontaceaePrionodon densus (Hedw.) Müll.Hal.Yano 13579 (SP)
PterobryaceaeOrthostichopsis tortipilis (Müll.Hal.) Broth.Peralta 27515 (SP)
PtychomitriaceaePtychomitrium sellowianum (Müll.Hal.) A.JaegerYano 12555 (SP)
PylaisiadelphaceaeIsopterygium tenerifolium Mitt.Peralta 20516 (SP)
PylaisiadelphaceaeIsopterygium tenerum (Sw.) Mitt.Eiten 7100
RacopilaceaeRacopilum tomentosum (Hedw.) Brid.Alkimim-Faria 1402 (UB)
RhacocarpaceaeRhacocarpus inermis (Müll.Hal.) Lindb.Vital 1192 (SP)
RhacocarpaceaeRhacocarpus purpurascens (Brid.) Müll.Hal.Peralta 27643 (SP)
RhizogoniaceaePyrrhobryum spiniforme (Hedw.) Mitt.Gonçalves 314 (SP)
RhizogoniaceaeRhizogonium novae-hollandiae (Brid.) Brid.Schaefer-Verwimp 13122 (SP)
RigodiaceaeRigodium toxarion (Schwägr.) A.Jaeger.Peralta 27537 (SP)
SeligeriaceaeBrachydontium notorogenes W.R.Buck & Schaf.-Verw.Peralta 20441 (SP)
SematophyllaceaeAptychella proligera (Broth.) HerzogPeralta 27650 (SP)
SematophyllaceaeAptychopsis pungifolia (Hampe) Broth.Gonçalves 200 (SP)
SematophyllaceaeAptychopsis pyrrhophylla (Müll.Hal.) Wijk & Margad.Peralta 27485 (SP)
SematophyllaceaeBrittonodoxa subpinnata (Brid.) W.R.Buck, P.E.A.S.Câmara & Carv.-Silva *Müller (1845)
SematophyllaceaeDonnellia commutata (Müll.Hal.) W.R.BuckAlkimim-Faria 1342 (UB)
SematophyllaceaeDonnellia lagenifera (Müll.Hal.) W.R.BuckGonzatti 4005 (SP)
SematophyllaceaeMicrocalpe subsimplex (Hedw.) W.R.Buck *Müller (1845)
SematophyllaceaeKuerschneria laevigata (Herzog) Ochyra & Bedn.-Ochyra *Ochyra & Bednarek Ochyra (2010) [53]
SematophyllaceaeSematophyllum beyrichii (Hornsch.) Broth.Gonçalves 331 (SP)
SematophyllaceaeSematophyllum cuspidiferum Mitt.Peralta 20495 (SP)
SematophyllaceaeSematophyllum decumbens Mitt.Peralta 27494 (SP)
SematophyllaceaeSematophyllum galipense (Müll.Hal.) Mitt.Peralta 27748 (SP)
SematophyllaceaeSematophyllum lithophilum (Hornsch.) Ångström *Buck (1998) [54]
SematophyllaceaeSematophyllum subdepressum (A.Jaeger) Broth. *Brotherus (1924)
SematophyllaceaeSematophyllum subpinnatum (Brid.) E.BrittonPeralta 27743 (SP)
SematophyllaceaeTrichosteleum glaziovii (Hampe) W.R.BuckPeralta 27514 (SP)
SematophyllaceaeTrichosteleum sentosum (Sull.) A.JaegerPeralta 27497 p.p. (SP)
SematophyllaceaeVitalia cuspidifera (Mitt.) P.E.A.S.Câmara, Carv.-Silva & W.R.BuckSampaio, A.J. 2380
SematophyllaceaeVitalia galipensis (Müll.Hal.) P.E.A.S.Câmara, Carv.-Silva & W.R.BuckLima 768 p.p. (SP)
SphagnaceaeSphagnum aciphyllum Müll.HalPeralta 27592 (SP)
SphagnaceaeSphagnum brasiliense Warnst.Santos & Costa 759, 760 (RB)
SphagnaceaeSphagnum brevirameum HampeGonçalves 167 (SP)
SphagnaceaeSphagnum capillifolium (Ehrh.) Hedw.Gonçalves 170 (SP)
SphagnaceaeSphagnum costae H.A.Crum & D.P.CostaPeralta 20494 (SP)
SphagnaceaeSphagnum cuspidatum Ehrh. ex Hoffm. *Luetzelburg (1923, as S. serratum) Gonçalves 369 (SP)
SphagnaceaeSphagnum divisum H.A.CrumGonçalves 419 (SP)
SphagnaceaeSphagnum exquisitum H.A.CrumLima 822 (SP)
SphagnaceaeSphagnum geraisense H.A.CrumGonçalves 421 (SP)
SphagnaceaeSphagnum gracilescens Hampe ex. Müll.Hal.Peralta 27651 (SP)
SphagnaceaeSphagnum luetzelburgii H.K.G.Paul ex H.A.CrumCosta 4173 (RB)
SphagnaceaeSphagnum longicomosum Müll.Hal. ex Warnst.Gonçalves 335 (SP)
SphagnaceaeSphagnum longistolo Müll.Hal.Peralta 27553 (SP)
SphagnaceaeSphagnum magellanicum Brid.Lima 824 (SP)
SphagnaceaeSphagnum palustre L. *Andrews (1941) [55]
SphagnaceaeSphagnum perforatum Warnst.Costa et al. 5017 (RB)
SphagnaceaeSphagnum perichaetiale HampePeralta 27652 (SP)
SphagnaceaeSphagnum platyphylloides Warnst.Gonçalves 415 (SP)
SphagnaceaeSphagnum pseudoramulinum H.A.CrumLima 817 (SP)
SphagnaceaeSphagnum ramulinum Warnst.Lima 815 (SP)
SphagnaceaeSphagnum recurvum P.Beauv.Peralta 27560 (SP)
SphagnaceaeSphagnum sanguinale Warnst.Gonçalves 168 (SP)
SphagnaceaeSphagnum sehnemii H.A.CrumLuetzelburg 6162 (F, HBR as Sphagnum densum)
SphagnaceaeSphagnum subsecundum NeesGonçalves 353 (SP)
SphagnaceaeSphagnum sucrei H.A.CrumPeralta 27598 (SP)
StereophyllaceaeEntodontopsis leucostega (Brid.) W.R.Buck & IrelandAlkimim-Faria 1456 (UB)
StereophyllaceaeEntodontopsis nitens (Mitt.) W.R.Buck & IrelandAlkimim-Faria 1263 (UB)
StereophyllaceaeEulacophyllum cultelliforme (Sull.) W.R.Buck & Ireland *Costa et al. (2005) [37]
ThuidiaceaeThuidium pseudoprotensum (Müll.Hal.) Mitt.Peralta 27616 (SP)
ThuidiaceaeThuidium recognitum (Hedwig) LindbergLima 797 (SP)
ThuidiaceaeThuidium tamariscinum (Hedw.) Schimp.Gonçalves 359 (SP)
ThuidiaceaeThuidium tomentosum Schimp.Peralta 20510 (SP)
ThuidiaceaePelekium involvens (Hedw.) A. TouwLima 751 (SP)
ThuidiaceaePelekium minutulum (Hedw.) A.TouwLima 746 (SP)
ThuidiaceaePelekium muricatulum (Hampe) A.TouwAlkimim-Faria 1376 (UB)
ThuidiaceaePelekium scabrosulum (Mitt.) A.TouwAlkimim-Faria 1388 (UB)
HornwortsAnthocerotaceaeAnthoceros hispidus Steph.Yano 13588 (SP)
AnthocerotaceaeAnthoceros punctatus L. *Hooker & Wilson (1844) [56]
DendrocerotaceaeDendroceros crispatus (Hook.) NeesPeralta 27459 (SP)
DendrocerotaceaeDendroceros crispus (Sw.) Nees *Gottsche et al. (1844)
DendrocerotaceaeNothoceros minarum (Nees) J.C.VillarrealGonçalves 178 SP)
DendrocerotaceaeNothoceros vincentianus (Lehm. & Lindenb.) J.C. VillarealPeralta 27705 (SP)
NotothyladaceaePhaeoceros bulbiculosus (Broth.) Prosk.Peralta 20452 (SP)
NothotyladaceaePhaeoceros laevis (L.) Prosk.Peralta 27542 (SP)
Table 2. List of endemic bryophyte species found in PARNASO. (EB = endemic to Brazil; ERJ = endemic to Rio de Janeiro State; and CS = conservation status category.
Table 2. List of endemic bryophyte species found in PARNASO. (EB = endemic to Brazil; ERJ = endemic to Rio de Janeiro State; and CS = conservation status category.
GroupTáxonEBERJCS
LiverwortsCheilolejeunea grosseoleosa C.J.Bastos & Schäf.-Verw.X
Cololejeunea diaphana A.EvansX
Cololejeunea papilliloba (Steph). Steph.X
Colura itatyana Steph.XX
Frullania schaefer-verwimpii Yuzawa & Hatt.X
Frullania vitalii Yuzawa & Hatt.X
Fuscocephaloziopsis crassifolia (Lindenb. & Gottsche) Váňa & L. Söderstr.X
Gongylanthus liebmannianus (Lindenb. & Gottsche) Steph. EN
Haplolejeunea umbrosa Gradst. & Ilkiu-BorgesX
Harpalejeunea schiffneri S.W. ArnellX
Harpalejeunea subacuta A.EvansX
Lejeunea beyrichiana (Steph) Gradst. & C.J.BastosXX
Lejeunea longidentata C.J.Bastos, Gradst., S. Vilas Bôas-Bastos & Schäf.-Verw.X
Lejeunea oligoclada SpruceX
Leptoscyphus spectabilis (Steph.) GrolleX
Metzgeria brasiliensis Schiffn.X
Metzgeria convoluta Steph.X
Metzgeria cratoneura Schiffn.X
Metzgeria hegewaldii Kuwah. XEN
Microlejeunea squarrosa J. Heinrichs, A. Schäfer-Verwimp, Pócs & S.S. DongX
Micropterygium pterygophyllum (Nees)X
Neesioscyphus carneus (Nees) GrolleX
Paracromastigum dusenii (Steph.) R.M.Schust. XEN
Plagiochila amicta Steph.X
Plagiochila boryana Gottsche ex Steph. EN
Saccogynidium caldense (Ångstr.) GrolleX
Southbya organensis Herzog XCR
Syzygiella uleana Steph.X
Vitalianthus bischlerianus (Porto & Grolle) R.M.Schust. & GiancottiX
MossesAtractylocarpus brasiliensis (Müll.Hal.) R.S.WilliamsX EN
Atractylocarpus longisetus (Hook.) E.B.Bartram EN
Brachydontium notorogenes W.R.Buck & Schaf.-Verw.XXCR
Brachymenium hornschuchianum Mart.X
Breutelia grandis (Hampe) ParisX
Callicostella martiana (Hornsch.) A.JaegerX
Calyptrochaeta albescens (Hampe) W.R.BuckX
Campylopus densicoma (Müll.Hal.) Paris EN
Campylopus dichrostis (Müll.Hal.) ParisX
Campylopus gemmatus (Müll.Hal.) ParisX
Campylopus julicaulis Broth.X
Campylopus thwaitesii (Mitt.) A.JaegerX
Campylopus uleanus (Müll.Hal.) Broth.X
Campylopus widgrenii (Müll.Hal.) Mitt.X
Cladastomum robustum Broth.X
Cladastomum ulei Müll.Hal.X
Crumuscus vitalis W.R.BuckX
Cyclodictyon marginatum (Hook. & Wilson) KuntzeX
Daltonia splachnoides (Sm.) Hook. & TaylorX
Dicranodontium pulchroalare Broth.XX
Dimerodontium pellucidum Schwägr.X
Ditrichum paulense Geh. ex HampeX
Itatiella ulei (Broth. ex Müll.Hal.) G.L.Sm.X
Leiomela piligera (Hampe) Broth.X
Lepidopilidium brevisetum (Hampe) Broth.X
Lepidopilum scabrisetum (Schwägr.) SteereX
Leptodontium stellatifolium (Hampe) Broth.X
Leptodontium wallisii (Müll.Hal.) Kindb. VU
Leskeodon aristatus (Geh. & Hampe) Broth.X
Leucobryum clavatum HampeX
Leucoloma triforme (Mitt.) A.JaegerX
Microcalpe subsimplex (Hedw.) W.R.BuckX
Polytrichum angustifolium Mitt.X
Rhacocarpus inermis (Müll.Hal.) Lindb.X
Sphagnum brasiliense Warnst.X
Sphagnum costae H.A.Crum & D.P.CostaX
Sphagnum divisum H.A.CrumX
Sphagnum exquisitum H.A.CrumX
Sphagnum geraisense H.A.CrumX
Sphagnum gracilescens HampeX
Sphagnum longicomosum Müll.Hal. ex Warnst.X
Sphagnum luetzelburgii H.K.G.Paul ex H.A.CrumXX
Sphagnum perforatum Warnst.X VU
Sphagnum pseudoramulinum H.A.CrumX
Sphagnum ramulinum Warnst.X
Sphagnum sehnemii H.A.CrumX
Sphagnum sucrei H.A.CrumX
Trichosteleum glaziovii (Hampe) W.R.BuckX
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de Lima, J.S.; Faria, A.L.A.; Gonçalves, M.T.A.; Peralta, D.F. Bryophytes of the Serra dos Órgãos National Park: Endemism and Conservation in the Atlantic Forest. Plants 2025, 14, 2419. https://doi.org/10.3390/plants14152419

AMA Style

de Lima JS, Faria ALA, Gonçalves MTA, Peralta DF. Bryophytes of the Serra dos Órgãos National Park: Endemism and Conservation in the Atlantic Forest. Plants. 2025; 14(15):2419. https://doi.org/10.3390/plants14152419

Chicago/Turabian Style

de Lima, Jéssica Soares, Allan Laid Alkimim Faria, Mateus Tomás Anselmo Gonçalves, and Denilson Fernandes Peralta. 2025. "Bryophytes of the Serra dos Órgãos National Park: Endemism and Conservation in the Atlantic Forest" Plants 14, no. 15: 2419. https://doi.org/10.3390/plants14152419

APA Style

de Lima, J. S., Faria, A. L. A., Gonçalves, M. T. A., & Peralta, D. F. (2025). Bryophytes of the Serra dos Órgãos National Park: Endemism and Conservation in the Atlantic Forest. Plants, 14(15), 2419. https://doi.org/10.3390/plants14152419

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