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Article

The Greuter Herbarium in Palermo: An Inventory of Its Type Specimens Available Online, with Some Thoughts on Type Terminology (Occasional Papers from the Herbarium Greuter, N° 5) †

1
Botanischer Garten, 14195 Berlin, Germany
2
Herbarium Mediterraneum Panormitanum, 90133 Palermo, Italy
I take pleasure in dedicating this paper to my friend Francesco Maria Raimondo on the occasion of his 80th birthday. The single prominent contemporary botanist of his home island, he has steered the fates of the Palermo Botanical Garden as its Director for 19 years and has restored it to its flagship role among Italian botanical gardens. It is sad to note that his heirloom does no longer receive the care and admiration it deserves, and that he himself must suffer the ingratitude of many of his pupils and former friends. May he live to see his dream come true of Palermo as the reborn hub of Mediterranean plant science, a role that its central position in the area predestines it to play.
Plants 2024, 13(8), 1086; https://doi.org/10.3390/plants13081086
Submission received: 8 March 2024 / Revised: 26 March 2024 / Accepted: 7 April 2024 / Published: 12 April 2024

Abstract

:
The separately stored type herbarium of the Herbarium Greuter in Palermo is comprised of 339 sheets, corresponding to 336 specimens pertaining to 328 different names. Material from the Mediterranean area, especially Greece, predominates, followed by that from the Caribbean (Cuba) and Australia. The list includes transcribed label data and links to the digital specimen images and to the protologue texts. A new type of terminology is introduced, with the terms “first-step holotype” and “second-step holotype” designating type categories parallel to the similar terms already in use for lectotypes, and the phrase “detailed here” is used as an equivalent to “designated here” in second-step typification.

1. Introduction

In December 2008, the author donated his personal herbarium and botanical library to the Università degli Studi in Palermo to become an autonomous part of the university’s Herbarium Mediterraneum Panormitanum (PAL), kept separately under the official designation PAL-Gr. The herbarium was then estimated to comprise c. 95,000 specimens and included the historical French herbaria of Father Pierre Gave (1846–1916) and Dr. Antoine Bras (1802–1883). Subsequently, by exchange, purchase, and gift, c. 150,000 specimens have been added that are currently owned by the Foundation Herbarium Greuter and are destined to be transferred eventually to and gradually included in the PAL-Gr herbarium.
PAL-Gr hosts specimens from all over the globe, with a neat predominance of material from the Mediterranean countries (Greece in the first place) and including sizeable portions of Australian, South African, and Caribbean material, reflecting the donor’s main areas of interest. Type specimens, in so far as they were identified as such, are kept separately for their better safeguarding. This type herbarium has been digitized, and the high-resolution images are being made freely available and searchable and the high-resolution images are being made freely available and searchable via the University of Palermo Botanical Garden’s Virtual Herbarium (https://herbarium.unipa.it/herbarium_vsimple_en_ma.asp, accessed on 6 March 2024).

2. Material and Methods

This paper lists the contents of the PAL-Gr type herbarium. The list provides the following information: (1) the name typified, with reference to its place of valid publication, and, when the protologue is available online, a link (URL) to it; protologues that could not be traced on the internet have been digitized anew and can be similarly accessed through an apposite link; (2) a limited number of synonyms, especially when the currently accepted name differs from the one being typified; (3) the kind of type (such as holotype, neotype, etc.); (4) the PAL-Gr accession number; (5) the country of origin of each specimen, followed, between quotation marks, by the collecting-site data as they appear on the label; (6) the collection date, collector name, and collecting number; and (7) the whereabouts and status of any duplicates of which the location is known.

3. Discussion

Type category terms. The International Code of Nomenclature, in its current edition (Turland & al. 2018 [1]; hereafter the Code), recognizes and defines the following type categories: holotype, syntype, lectotype, paratype, neotype, and epitype, plus the duplicates of each of these types, using the prefix “iso-”. Paratypes having no immediate relevance for nomenclatural purposes are not taken into consideration, nor even mentioned, in the present inventory.
The system of type categories thus defined is, in a general way, consistent and has been working in a satisfactory way for many years and through several successive editions of the Code. There is, however, one major illogicality embedded in it. The types of various categories, as defined in Art. 9, all consist of one element: a single specimen (or sometimes illustration). However, as stated in Art. 40.2, a type may also consist of more than one specimen, provided they all belong to the same gathering. That provision applies in the case of names published from 1958 onward, when an indication of the type became a prerequisite for the valid publication of names of new genera and infrageneric taxa. However, even before 1958, cases abound in which reference to multiple specimens of a single gathering is referred to in a protologue, and these specimen runs have then been generally (if inaccurately) accepted as constituting the holotype.
Analogous situations frequently arise with types of other categories, but these are explicitly covered in Art. 9.17. The Code, in Art. 9 Ex. 14, in such cases uses the phrase “first-step” lectotype for the multiple-specimen “type” initially designated and permits subsequent selection of a “second-step” type from among the elements of the “first-step” [lecto-]type. This introduces a disparity of treatment between holotypes and other type categories, in so far as with the former, when the original type consists of more than one specimen, all of these are considered to be syntypes (Art. 40 Note 1). As there is no provision parallel to Art. 9.17, restricting subsequent lectotype designation to one of these syntypes, other isosyntypes, even if not seen or cited by the original author, are under Art. 9.12, eligible as types on equal footing with the syntypes proper.
One of the purposes of the following list has been to establish the correct categories of the types concerned. Surprisingly, it turned out that close to one-third of the names concerned had “holotypes” consisting of more than one specimen. As a consequence, I felt it was very desirable to devise a satisfactory way to address this situation in simple and easily understood terms. I, therefore, propose to designate such non-unitary “holotypes” as “first-step holotypes” by analogy to what the Code has been using for lectotypes of a similar kind and, by the same analogy, to designate as “second-step holotypes” the lectotypes obtained by selecting one of the first-step holotype elements by analogy to what Art. 9.17 (and not just 9.12) provides. Also, in this kind of situation, I shall use the phrase “specified here” as a direct equivalent of “designated here” to meet the requirements of Art. 7.11. These suggestions are not covered by the provisions of the current Code, but they are compatible with them. If found to be useful and generally acceptable, they may hopefully be reflected in the text of some future Code edition (the date for submitting proposals for the next edition has unfortunately passed).

4. Results: The PAL-Gr Type Inventory

Isotype: PAL-Gr 52738—Albania, “Korca, Mali i Moravës, bei Drenova, ca. 1100–1200 m, Serpentin,” 12. Sep 1961, F. K. Meyer 6151 (Holotype: JE) (other isotypes: JE [x3]).
Isotype: PAL-Gr 15480—Greece, “Epirus orientalis. In rupestribus calcareis regionis alpinae inferioris mt. Tsumerka supra pagum Vulgarelion loco dicto Spathes. Alt. 1500 m,” 8 Jul 1893, E. de Halácsy, Iter graecum secundum a. 1893 (holotype [specified here]: WU in Halácsy Greek hb.) (other isotypes: B [×3], JE, K, LD, S).
Holotype: PAL-Gr 24749—Greece, “Epirus/Macedonia (distr. Konitsa/Kastoria), montes Grammos in vertice orientali, alt. 2350–2442 m. In petrosis graminosis, solo flyscheo,” 14 Aug 1978, W. Greuter 14360 (Isotypes: ATH, B, C, G, M, UPA).
Isotype: PAL-Gr 123563—Israel, “Philistean Plain, hill near kibbutz Giv’at-Brenner, Batha and Garigue on calcified sandstone, 31.867130 N, 34.807520 E, Alt. 70 m.”, 2 Apr 2013, O. Fragman-Sapir (holotype: HUJ; other isotypes: HUI, OSBU).
  • Allium antonii-bolosii (‘A. Bolosii’) P. Palau in Anales Inst. Bot. Cavanilles 11: 485. 1953 [? = Allium hirtovaginatum Kunth]. [25723].
Isotype: PAL-Gr 25723—Spain, Balearic Islands, “Cabrera (Baleares), c.d.” [Protologue: Habitat in fis[s]uris angstissimis rupestribus calcareis ad N. vel ad W orientatis. Floret Jul-Sept.], Palau (dupl. ex BC; holotype [specified here]: herbarium, Col·legi Oficial de Farmacèutics de les Illes Balears, Palma de Mallorca).
Holotype [specified here]: PAL-Gr 35235—Libya, “Wadi el-Bab,” 16 May 1981, Brullo and Furnari (isotypes: B, CAT [2×], FI, G [2×], MPU).
  • Allium integerrimum Zahar. in Ann. Mus. Goulandris 3: 90. 1977 [19137].
Holotype: PAL-Gr 19137—Greece, “Macedonia occ., prov. et distr. Pieria: in colle arci (katro) Platamon, alt. 50–100 m. In rarioribs dumetorum Quercus cocciferae. Flores rosei,” 7 Jul 1971, Greuter 9192 (Isotype: ATH).
Isotype: PAL-Gr 35757—Italy, Sicily, “Lampedusa, Vallone Madonna,” 25 Jun 1985, Brullo (holotype: CAT; other isotypes: B, CAT, FI).
Holotype: PAL-Gr 45677—Greece, “Nomos Evros, Eparhia Soufli, 1 km S of Dhadhia, 100 m, 41°07′20″ N, 26°13′ E. Grassland on micaschist hill. Tepals cream with pale green midnerve, anthers light yellow, ovary purplish black,” 13 Jun 1992, Greuter 23311, Zimmer, Phitos, Kamari, Anagnostopoulos, and Athanasiou (Isotypes: B, SEV).
Holotype: 51792 PAL-Gr—Greece, “Nom. Lakonia, Ep. Epidhavros Limiras: Bay of Palea Monemvasia, alt. 0–2 m. 36°44′00″ N, 23°02′00″ E. Sandy beach and salt marshes,” 8 Jun 1995, Iatrou & al. in OPTIMA Iter Mediterraneum VII, #1792. (Isotype: none).
Lectotype (designated here): 66270 p.p. [excl.sterile leafy plant] PAL-Gr—Greece, “in arenosis maritimis Phaleri, flor. d. 21 Jun 1873, Heldreich in Baenitz, Herbarium europaeum.” (Isolectotypes: numerous; paralectotypes: same data but “folia d. 26 Mar 1874,” mixed with the foregoing).
  • Allium tardans Greuter and Zahar. in Biol. Gallo-Hellen. 6(1): 51. 1975 [17519].
Holotype: PAL-Gr 17519—Greece, Crete, “Ep. Temenos: ad cacumen montis Júktas. 700 m. In praeruptis rupestribus calcareisadoccidentem spectantibus parcissime,” 14 Sep 1966, Greuter 7519 (Isotype: none).
Isotype: PAL-Gr 63251—Cuba, “Prov. Villa Clara, Sagua la Grande. Mogotes de Jumagua. Vegetación cársica de mogotes. 50–88 m s.m.,” 10 Nov 2011, Noa, Pérez-Lobregón and Oliver in UCLV 10024 (holotype: ULV [specimen shown in Revista Jard. Bot. Nac. Univ. Habana 40: 93, in Fig. 5]; other isotypes: B, HAC, HAJB, JE, ULV).
Holotype: PAL-Gr 15974—Greece, “Sterea, prov. Viotia, distr. Levadhia: in latere austro-occidentali montis Parnassos supra Arahova, alt. 1400 m. In clivis asperis rupestribus calcareis meridiem spectantibus. In scansilibus sole illustribus,” 15 Jun 1963, Greuter 5974 (Isotypes: G, MEL, SALA, UPA, Z).
Holotype: PAL-Gr 14667—Greece, Crete, “Ep. Kissamoas: W-Teil der Insel Elafonìsi, Kuppe beim Leuchtfeuer, 10 m.ü.M., Kalk. Auf Flugsand auf felsigem Grund; stark abweichende neue Sippe des Rassenkreises,” 4 Jun 1962, Greuter 4667 (Isotypes: G, HUJ, LD, W, Z).
Isotype: PAL-Gr 58585—Italy, Sicily, “Cavagrande del Cassibile (Siracuse), 36°97′89″ N, 15°07′95″ E, 400 m a.s.l., on vertical calcarenite cliffs,” 2 May 2012, GuarinoRaimondo (holotype [to be specified]: PAL; isotypes: B, FI, PAL).
Isotype: PAL-Gr 30612—Greece: “insula Antikythera: Potamos, in parte boreali insulae, in saxosis calc. litoreis,” 8 May 1964, Rechinger 24367a (Holotype [specified here]: G; isotype: W?).
Holotype PAL-Gr 13642—Greece, Crete, “Ep. Ierapetra/Sitia: N-Hang des Berges Afèndis Kavoùsi, 1200 m.ü.M., Kalk. Auf Schutt an offenen Stellen in der Phrygana stellenw. sehr hfg.,” 19 Jun 1961, Greuter 3642 (Isotypes: G, W, Z).
Isotype: PAL-Gr 133293—Cuba, “Parque Nacional Viñales, Sierra de la Caoba, ladera hacia Cañadores, Pinar del Río. Individuo único sobre carso de mogote. 225 m smn,” 3 Aug 2011, Chaves in HAC 43069 (holotype: HAC [specimen with flowers]; other isotypes: B, HAC, HAJB [2×], JE).
Isotype: PAL-Gr 128078—Cuba, “Isla de la Juventud (Isla de Pinos) (Mun. especial), llanura cársica del sur: camino de Cayo Piedra (Cayo las Piedras) a Punta del Este, monte seco, calizo,” 4 Nov 1981, Álvarez & al. In HFC 45586 (holotype: HAJB [specimen #53, shown in the protologue, Figure 1]; other isotypes: B, JE [2×]).
Isotype: PAL-Gr 30751—Greece: “Messenia, Methoni, in ruinis castelli,” 11 May 1964, Rechinger 24861 (Holotype [specified here]: G; isotype: W?):
  • Aristida neglecta subsp. breviglumis Catasús in Greuter and Rankin, Fl. Rep. Cuba, Ser. A, Pl. Vasc. 21A: 81. 2015 [63250].
Isotype: PAL-Gr 63250—Cuba, “Prov. Granma, municipio de Buey Arriba, Palmarito, área protegida Refugio de Fauna ‘Bosque de Palmarito’, al SE de Yara,” 15 Dec 2014, Catasús and Arjona (Holotype: HAJB #451; other isotypes: B, HAJB ##449–450, JE).
  • Aristolochia merxmuelleri Greuter and E. Mayer in Bot. Jahrb. Syst. 107: 321. 1985 [35745].
Isotype: PAL-Gr 35745—“Yugoslavia,” SW-Serbia (Kosovo): Miruša sub m. Koznik—in declivibus glareosis fruticosis, solo serpentin., cca 500 m s.m.,” 20 Apr 1985, Mayer 11071 Holotype: LJU; other isotypes: B, M).
  • Arrhenatherum elatius subsp. braun-blanquetii P. Monts. and L. Villar in Doc. Phytosoc. 78: 13. 1974 [=Arrhenatherum elatius (L.) P. Beauv.] [7110].
Isotype: PAL-Gr 7110—Spain, “ditione oscensis (Huesca), Aragonia, in glareosis cum Cochlearia aragonensis ad sepemtrionem Sierra de Guara, 1550 m alt., lapides minutas, calcareas, 1–3 cm diam.,” 4 Jul 1968, Montserrat 2848/68 (Holotype: ?JACA).
Holotype: PAL-Gr 48357—Greece, Crete, “Ep. Sitia N-u. NO-Seite des Kap Màvro nwl. Erimùpolis, 20–160 m.ü.M., Kalk u. pleistozänes Konglomerat. Offene Lygeum-Fluren, Felsritzen etc. vebr. U. stellenw. Hfg.; neue Art aus der Verwandtschaft v. A. turnefortii, hierher auch “A. crassifolia L.” b. Gandog.,” 12 May 1962, Greuter 4446 (Isotypes: ATH, G, HUJ, LD, UPA, W, Z, hb. Zaffran).
Isotype: PAL-Gr 66334—Italien: “supra Bordighera in sax. M. Nero [‘Alles nur überwinterte Formen von neuem blühend! Daher kleinblumiger’, Bornmüller],” 25 Apr 1893, Haussknecht (holotype [specified here]: JE #16649).
Holotype: PAL-Gr 53217—Greece “Nom. Arkadia, Ep. Megalopolis: NW foothills of the Taijetos range, c. 0.5 km W of Neochori, alt. c. 100 m, 37°11′00″ N 22!13′45″ E.” Mixed fir, pone and chestnut wood by a stream, on mixed substrate,” 11 Jun 1995, Kamari & al. in OPTIMA Iter Mediterraneum VII, #2217. (isotypes: B, BEO, BRNM, MA, SALA, UPA, W).
Isotype: PAL-Gr 6014—Greece: “Saria ins.,” 26 May 1886, Major 544 (holotype [specified here]: FI #15355; other isotypes: FI ##15353–15354, 15356–15361 [8×], G, K, M, MPU, NY [2×], RSA, US).
Isotype: PAL-Gr 39662—Australia, [Western Australia: “Murchison River”] (see Chapman, Austral. Pl. Name Index: 305. 1991) [Oldfield in Drummond VI] 121 (holotype [specified here]: K #277584; isotypes: BM, G, K, L, MEL).
Isotype: PAL-Gr 133292—Cuba, “Prov. Villa Clara. Santa Clara. En los exteriores de la sala de oncología del hospital infantil ‘José Luis Miranda’. Vegetación ruderal,” 9 Jan 2022, Puente and Castañeda (holotype: HAJB #1256; other isotypes: B, JE, ULV [2×]).
  • Aubrieta albanica F. K. Mey. and J. E. Mey. in Haussknechtia, Beih., 15: 62. 2011 [52737].
Isotype: PAL-Gr 52737—Albania: “Nemërçka, über Permet, südlich Qafa e Dhembelit, ca. 1750 m, Westhang,” 12 Jul 1959, Meyer 3952 (holotype [specified here]: JE #16678; other isotype: JE #16679).
Holotype: PAL-Gr 53554—Greece, “Nom. Messinia, Ep. Kalamata: 6–8 km NE of Ano Amfía along the road to Poliana, alt. 600–800 m, 37°08′20″ N, 22°07′30″ E. Dry river bed and rocky limestone slopes with maquis,” 14 Jun 1995, Kamari & al. in OPTIMA Iter Mediterraneum VII, #2554 (isotypes: B, BEO, BRNM, MA, SALA, UPA, W).
Isotype: PAL-Gr 66672—Greece, “Agrapha (Dolopia veterum): in reg. inf. Pindi circa monasterium Corona, in nem. querc., alt. 3500–3370′, substr. Schist.,” 20–28 Jun 1885, Haussknecht (holotype [specified here]: JE #3794; isotypes: JE #3792–3793, 3795–3796, K, BR).
Isotype: PAL-Gr 39670—Australia, [Western Australia, “Nov. Holland. Swan River (from holotype label), [Drummond VI] 85″ (holotype [specified here]: G #434218; other isotypes: K #959438, MEL #114150, 114151, PERTH?).
Holotype: PAL-Gr 13733—Greece, Crete, “Ep. Pirjotissi: NW-Hang des Berges Mavri ob der Quelle karonero, 1800–1900 m.ü.M., Tripolitsa-Kalk. Auf Feischuttflecken am felsigen Abhang, nicht selten,” 30 Jun 1961, Greuter 3733 (isotypes: G, W, Z).
  • Bolanthus intermedius Phitos in Bot. Chron. 1: 39. 1981 [=Bolanthus thymifolius (Sm.) Phitos] [29534].
Isotype: PAL-Gr 29534—Greece, “Ins. Euboea: in saxosis serpentinicis et magnesiticis litoreis ad pagum Mantudi,” 23 Jun 1978, Georgiadis 1657 (holotype: UPA); other isotypes: none traced).
Isotype: PAL-Gr 35236—Greece, Crete: “eparch. Sfakia: in monte Volakias, alt. 1700 m,” 3 Aug 1966, Zaffran 3223 (Holotype: B).
Holotype: PAL-Gr 15518—Greece, “Regio aegaea austr., insula Karpathos: in faucibus Kounoupa ad septemtriones oropedii Lastos, alt. 600 m. In scansilibus rupium calcarearum et in herbidis inter saxa,” 23 May 1963, Greuter 5518 (isotypes: G, HUJ, LD, M, W, Z).
Isotype: PAL-Gr 35236—Montenegro, “In asperis calcareis ad Njegus,” Jul 1898, Sagorski (holotype: JE?).
  • Bumelia neglecta Bisse and J. E. Gut. in Revista Jard. Bot. Nac. Univ. Habana 6: 21. 1985, homon. illeg. [non Bumelia neglecta (Cronquist) Lundell 1975] [=Sideroxylon cubense (Griseb.) T. D. Penn.] [https://www.jstor.org/stable/42596610?seq=3 (accessed on 6 March 2024)].
Isotype: PAL-Gr 123176—Cuba, “Prov. Pinar del Río, Loma Peluda de Cayajabos, 200–400 m, IX? 1972, Bisse in HFC 23422 (holotype [specified here]: HAJB #273; other isotypes: B, FR, HAJB #274–278 [5×], JE).
Holotype: PAL-Gr 17714—Greece, Crete, “Ep. Sfakia: supra hiatum Kakískala prope fontem Linoséli, 1450 m. Ad rupes praeruptas calcáreas sole illustres haud raro, sed nil praeter folia vidi,” 11 Oct 1966, Greuter 7714 (isotypes: E, G, K, LD, M, W).
Isotype: PAL-Gr 121244—Cuba, “Prov. Holguín, Moa. Alrededores del Campamento Arroyo Limones. Suelo laterita,” 18 Apr 1985, Álvarez & al. in HFC 56156 (holotype [specified here]: HAJB #1067; other isotypes: B [2×], HAJB ##1068–1069 [2×], JE [2×]).
Isotypes: PAL-Gr 121243 and 126208—Cuba, “Prov. Holguín, Mun. Moa. Cuchillas de Moa, Alrededores del Aserrío La Melba. Suelo arciloso ácido, charrascales altos con Bonetia cubensis, 450–500 m s.m.,” 28 Apr 1980, Álvarez & al. in HFC 42165 (holotype [specified here]: HAJB #1065; other isotypes: B [2×], HAJB ##1064, 1066 [2×], JE [2×]).
  • Buxus sclerophylla Eg. Köhler in Wiss. Z. Friedrich-Schiller Univ. Jena, Mat.-Naturwiss. Kl. 31(2): 239. 1982. [123180].
Isotype: PAL-Gr 123180—Cuba, “Prov. Guantánamo. Mun. San Antonio del Sur: Abra Mariana, loma al este del barranco,” 9 Feb 1979, Berazaín & al. in HFC 39083 (holotype [specified here]: HAJB #239; other isotypes: B, HAC, HAJB ##240–242 [3×], JE ##25874–25875 [2×]).
Isotype: PAL-Gr 66703—Montenegro, “zwischen Njegus u. Cetinje,” Sagorski (holotype [specified here]: JE #5419; isotypes: JE #5418, MEL).
  • Calendula suffruticosa subsp. greuteri Ohle in Feddes Repert. 85: 269. 1974 [17133].
Holotype: PAL-Gr 17133—Spain, “Andalusien, Prov. Granada: unterste Talenge des Rio Guadalfeo nwl. Motril, 70 m.ü.M. Steile w-exp. Felshänge, Kalk; auf Schutt nicht selten zwischen Gebüsch,” 5 May 1965, Greuter 7133 (isotypes: HAL, W).
  • Campanula aizoides Zaffran in Greuter, Florist. Report Cretan Area: 15. 1972 [3427].
Holotype: PAL-Gr 3427—Greece, Crete, “Crête à l’E des gorges de Samaria. Calcaire. Altitude 2000 m,” 16 Jul 1965, Zaffran (isotype: hb. Zaffran).
Second-step holotype (specified here): PAL-Gr 14190—Greece, Crete, “Ep. Temenos: Insel Dìa, Bucht Panajìa an der Südküste, 1–2 m.ü.M., Kalk. Ritzen und Löcher der Küstenfelsen, hfg. mit Bellium minutum,” 20 Apr 1962, Greuter 4190 (isotypes: G, HUJ, LD, W, Z).
  • Campanula flagellaris Halácsy in Denkschr. Kaiserl. Akad. Wiss., Wien. Math.-Naturwiss. Kl. 61: 246, 1894 [non Kunth 1819] [=Campanula tymphaea Hausskn.] [Gr].
Isolectotype: PAL-Gr 5487—Greece, “Epirus boreali-orientalis. In herbidis regionis superioris m. Peristeri. Alt. 1700 m. Solo calcareo,” 14 Jul 1893, Halácsy (lectotype [designated here]: WU in Halácsy Greek hb.).
Isotype: PAL-Gr 122719—Greece, “Nomos Lakonias, Ep. Gythiou: 1.5 km N-NW of Areopolis, east entrance of Limeni village, above the road: on limestone rocks. Alt. 20–30 m. Lat. 36°40′ N, Long. 22°22′ E,” 30 Apr 2017, Kyriakopoulos 1321 (holotype: UPA; other isotype: ACA).
Second-step holotype (specified here): PAL-Gr 44153; and isotype: PAL-Gr 27034—Greece, “ex insula Kastellorizo (Megiste). Insula Ro, alt. 2–100 m. Profusius in totâ insulâ,” 12 Apr 1974, Greuter 11829 (other isotype: ATH).
  • Campanula korabensis F. K. Mey. in Haussknechtia, Beih., 15: 149. 2011 (≡Campanula versicolor subsp. korabensis (F. K. Mey.) I. Jankovic and D. Lakusic [52736].
Isotype: PAL-Gr 52736—Albania, “Korab, Wiesen oberhalb Radomir, ca. 1400 m, an Felsen,” 5 Aug 1959, Meyer 4759 (holotype [specified here]: JE #16705; isotype: JE #16706).
Second-step holotype (specified here): PAL-Gr 15532—Greece, “Regio aegaea austr., insula Karpathos: in latere occidentali montis Kali Limni prope fontem Holetra, alt. 800 m. In clivis dumulosis calcareis, solo rupestri, abunde,” 25 May 1963, Greuter 5532 (isotypes: LJU, M, W, UPA, Z, hb. Grims).
Isolectotype: PAL-Gr 66358—Greece, “(Agrapha (Dolopia veterum): in reg. infer. m. Pindi circa monasterium Koróna, in nemorosis quercinis alt 3500–3700′ ssubstratu schistoso,” 20–28 Jun 1885, Haussknecht (holotype [designated here]: JE #5947; other isolectotype: JE #5948).
Holotype: PAL-Gr 12885—Greece, Crete, “Ep. Pirjotissi: S-Hang des Berges Màvri ob Kamàres, ca. 1400 m.ü.M., Kalk. Kermeseichenwald, bei einer Quelle,” 3 May 1960, Greuter 2885 (isotypes:none).
Isotype: PAL-Gr 129939—Cuba, “Prov. Guantánamo, Baracoa, parte alta del Yunque de Baracoa. Bosque siempre verde mesófilo,” 17 Apr 1986, Arias & al. in HFC 58862 (holotype: HAJB #1260; other isotypes: B, HAJB #1261–1263 [3×], JE [2×], ULV).
Isotype: PAL-Gr 59075—Cuba, “Las Tunas, Jesús Menéndez, al este de Playa Herradura, complejo de vegetción de costa arenosa, 76°25′44″ W, 16°49′ N, alt. 5 m,” 27 Mar 2013, S. Fuentes & al. 749 (holotype: ULV (specimen shown in the protologue, Figure 9); other isotypes: B, HAJB #1266–1271 [6×], JE, ULV).
  • Centaurea aegusae Domina, Greuter, and Raimondo in Israel J. Pl. Sci. 64: 51. 2017 (≡Centaurea panormitana subsp. aegusae (Domina, Greuter, and Raimondo) Guarino and Pignatti 2019) [https://www.researchgate.net/publication/315694332 (accessed on 6 March 2024)].
Isotype: PAL-Gr 122018—Italy, Sicily, “Island of Favignana, Mt. Santa Caterina, Scindo Passo, 37.920730° N, 12.307299° E, 100 m a.s.l., maritime carbonate cliffs,” 25 May 2016, Domina (holotype [to be specified]: PAL; isotypes: B, FI, PAL, RO).
  • Centaurea chrysocephala Phitos and T. Georgiadis in Biol. and Ecol. Médit. 4(1): 4. 1978 [25628].
Isotype: PAL-Gr 25628—Greece, “Prov. Trikala: supra oppidum Kalampaka, ad monasterium Varlaam-Meteora; in conglomeratis,” 4 Jul 1974, Phitos 12563 (holotype: UPA).
Isotype: PAL-Gr 38807—Greece, “Nomos “Viotias, Ep. Megaridos/Thivon: Mt. Kitheron, W of an abandoned military camp; open, rocky, grazed ground, alt. 1290 m; limestone, 38°11′ N, 23°15′ E. Flowers yellow,” 12 Jul 1992, Constantinidis 2690 (holotype: UPA; other isotype: B).
  • Centaurea cycladum Heldr., Herb. Graec. Norm. 1902: #1652. 1902 (=Centaurea spinosa subsp. cycladum (Heldr.) Hayek) [66454].
Isolectotype: PAL-Gr 66454—Greece, “Flora Aegaea: legimus in Cycladum ins. Mykonos ubi vulgatissima in collibus et montibus graniticis,” 15 Jul 1901, Heldreich in Herbarium Graecum Normale #1652 (Lectotype [designated here]: WU #67939; other isolectotypes: GB, JE [2×], LD, M, PI, W).
Holotype: PAL-Gr 19638—Greece, “Regio Aegaea sept., penins Sithonia: in regione Aliki inter pagos Neos Marmaras et Toroni, alt. 20 m. In fissuris et at pedes rupium marmoreo-schistosorum praeruptarum secus viam,” 18 Jun 1972, Greuter 10514 and Rechinger 44684 (isotypes: ATH, BSB, ELVE, G, LTR, UPA, hb. Damboldt, hb. Grims).
  • Centaurea grisebachii subsp. occidentalis in Strid and Kit Tan, Mount. Fl. Greece 2: 500. 1991 [20855].
Isotype: PAL-Gr 20855—Greece, “Macedonia occ., prov. and distr. Grevena: ad occidentem pagi Kranea, in latere boreo-orientali montis Simandro, loco ‘Baltsa’ vocato, alt. 1500 m. In regione sillvae Fagi, Pini leucodermis et pallasianae, solo ophiolithico,” 16 Aug 1974, Charpin 11126, Dittrich, Greuter 12295 and van Auw (holotype: C; other isotypes: G, LD, DUIS, UPA, W, hb. Dittrich).
  • Centaurea mantoudii T. Georgiadis, Contrib. Phylogén. Centaurea L. (sect. Acrolophus (Cass.) DC.) en Grèce: 56. 1980 [=Centaurea laureotica Heldr. ex Halácsy 1898] [29715].
Isotype: PAL-Gr 29715—Greece, “Ins. Euboea: in saxosis maritimis, propre pagum Mantoudi,” 23 Jun 1978, Georgiadis 1723 (holotype: UPA; other isotypes: none known).
  • Centaurea musakii T. Georgiadis in Bot. Not. 132: 311. 1979 [29709].
Isotype: PAL-Gr 29709—Greece, “Prov. Trikala: prope pagum Pyli, in rupestribus calc. alt. c. 200 m,” 22 Sep 1977, Georgiadis 1732 (holotype: UPA; other isotypes: none known).
Isotype: PAL-Gr 32045—Greece, “Macedonia or., prov. Kavala, eparhia Pangeo: in crista orientali montis Pangeo, ad occidentem cacuminis Kara Göl tepe, alt. 1300 m (40°54′30″ N/24°08′ E). In clivis rupestribus marmoreis praeruptis septemtriones spectantibus,” 18 Jul 1978, Greuter 16055 (holotype: B; other isotypes: AAU, ATH, C, DUIS, G, GOET, M, UPA).
  • Centaurea panormitana Lojac., Fl. Sicul. 2(1): 137. 1903 [xxx].
isoneotype: PAL-Gr 67524—Italy, Sicily, “Punta Mastrangelo, Monreale, prov. of Palermo. Coordinates: 38°03′52″ N, 13°14′37″ E, alt. 850 m a.s.l. Habitat: carbonate cliffs facing the sea,” 3 Aug 2015, Domina (neotype (Domina & al. in Willdenowia 46: 25. 2016 [https://www.jstor.org/stable/24753265 (accessed on 6 March 2024)]: PAL; other isoneotypes: B, BM, FI, RO).
Isotype: PAL-Gr 25642—Greece, “Prov. Ipiros: prope vicum Monodendrion, ad muros monast. Hagia Paraskevi, ca 1100 m,” 13 Jun 1972, Phitos 11716 (holotype: UPA; other isotypes: none known).
  • Centaurea pelia var. trigona T. Georgiadis, Contrib. Phylogén. Centaurea L. (sect. Acrolophus (Cass.) DC.) en Grèce: 59. 1980.
Isotype: PAL-Gr 29737—Greece, “Prov. Trikala: inter vicos Metsovo et Kalambaka, prope pagum Trigona,” 4 Jul 1974, Phitos 12564 and Tsanoudakis (holotype: UPA; other isotypes: none known).
Holotype: PAL-Gr 17689—Greece, Crete, “Ep. Sfakia: in faucibus infra pagum Asféndos, 280–320 m. In rupium calcarearum praeruptarum fissuris sole illustribus frequens. Flores desunt, sed ex verbis pastoris indigenae certe rosei,” 8 Oct 1966, Greuter 7689 (isotypes: ATH, B, E, G, GB, K, LD, M, P, W, hb. Phitos, hb. Zaffran).
  • Centaurea prespana Rech. f. in Ann. Mus. Goulandris 2: 55. 1975 [? = Centaurea soskae Hayek 1926] [30762].
Isotype: PAL-Gr 30762—Greece, “Macedonia occidentalis: distr. Phlorina: ad occidentem pagi Vrondera ad occidentem lacus Prespa ad fines Albaniae. In saxosis calc., 1300–1400 m,” 23 Jun 1972, Stamatiadou (holotype: W; other isotype: ATH).
Isotype: PAL-Gr 20089—Greece, “Regio Aegaea orientalis, insula Psara: in colle ‘Karajianni’ ad sept. sinûs Arhondiki litoris occidentalis, alt. 50–100 m. In dumulosis spinescentibus, solo rupestri eruptivo,” 24 Apr 1973, Greuter 11064 (holotype: ATH; other isotypes: none).
  • Centaurea thasia Hayek in Repert. Spec. Nov. Regni Veg. Beih. 30(2): 762. 1931 [20433].
Neotype [designated here]: PAL-Gr 20433—Greece, “Regio aegaea sept., insula Thasos: in vertice montis Ipsario, alt. 1100 m. In praeruptis rupestribus schistosis septemtriones spectantibus regionis silvaticae Pini nigrae,” 6 Jul 1973, Greuter 11431 (isoneotypes: ATH, C, G, H).
Holotype: PAL-Gr 16558—Greece, “Aegäis; nom. Attikí, Ep. Kíthira. Bei der Kirche Odijìtria söl. Livàdi, 170 m.ü.M., sandig verwitterndes Neogen. Cistus-phrygana, vereinzelt,” 12 May 1964, Greuter 6558 (isotypes: none).
  • Cerastium epiroticum Möschl and Rech.f. in Bol. Soc. Brot., ser. 2, 36: 41. 1962 [30586].
Isotype: PAL-Gr 30586—Greece, “Epirus: in saxosis calc. vallis fluvii Thiamis prope pagum Vrusina, 51 km W Ioannina,” 15 May 1961, Rechinger 23385 (holotype: W #12853; other isotypes: B, E, G, GZU, MA, MO, S, WU).
  • Cerastium grandiflorum subsp. serpentini F. K. Mey. in Haussknechtia Beih., 15: 47. 2011 [52735].
Isotype: PAL-Gr 52735—Albania, “Kolgecaj (Bajram Curri), an der Straße zwischen Kam und Kolgecaj, ca. 400 m. Serpentin,” 13 May 1960, Meyer 5609 (holotype [specified here]: JE #16669; other isotype: JE #16670).
Holotype: PAL-Gr 13273—Greece, Crete, “Ep. Kidonia: H.-I. Akrotìri, NW-Grat des Hügels Sklòcha, 430 m.ü.M., Kalk. Rasenfg. eine flache, erdgefüllte Gesteinsmulde ausfüllend, streng lokal; Zwergform mit auffällig verkürzten Btn.-Stielen,” 17 Mar 1961, Greuter 3273 (isotype: Z).
  • Chamaecrista lineata var. imiasensis (‘iniasensis’) A. Barreto and Yakovlev in Novosti Sist. Vysš. Rast. 28: 93. 1991. [128079].
Isotype: PAL-Gr 128079—Cuba, “Prov. Guantánamo. Mun. Imías: Alto de Cotilla (al norte de Cajobabo), pinares y charrascos, suelo ultrabásico y laterita, 400–500 m,” 29 May 1982, Bässler & al. in HFC 47172 (holotype [specified here]: HAJB #330); other isotypes: B, HAJB #331–334 [4×], JE).
  • Chamaepeuce atropurpurea Boiss. [& Heldr. in Boissier, Diagn. Pl. Orient., ser. 2, 3: 47. 1856, pro syn.] ex Pančić, Fl. Princip. Serbiae: 432. 1874 [6130] [=Ptilostemon strictus (Ten.) Greuter].
Isolectotype: PAL-Gr 6130—Serbia, “In dumetis Neresina Serb. austr.,” Aug 1872, Pančić (lectotype (Greuter in Boissiera 22: 79. 1973): LAU; other isolectotype: BEO?).
Isolectotype: PAL-Gr 6006—Morocco, “Hab. In decivibus montis Iguermalez, versus Tizzi Iffri (Atlante rhiphaeo), 1500 m alt., solo schistoso,” 24 Jun 1927, Font Quer in It. Marocc. 1927 #691 (lectotype [Greuter in Boissiera 22: 127. 1973]: G; other isolectotypes: BCN, BM, FI, LAU, MA, MPU [2×], RO, S).
Holotype: PAL-Gr 14848—Greece, Crete, “Ep. Sfakia: Karst-Kraterlandschaft in N-Teil des Talkessels Ammudzarà ndl. ob Anòpolis, 1800 m.ü.M., Tripolitsa-Kalk. In grossen Büschen mehrf. in sonst kahlen senkrechten Karren u. Schründen, intensiv duftend; neue Art,” 2 Jul 1962, Greuter 4848 (isotypes: ATH, G, HUJ, LD, W, Z).
Isotype: PAL-Gr 122003—Cuba, “Provincia Santiago de Cuba. Altiplanicie de Santa María del Loreto. Finca los Monieles, al este de la subestación eléctrica con paneles fotovoltaicos. Vegetación secundaria sobre suelos derivados de areniscas y conglomerados. Al borde de los farallones,” 1 Oct 2016, Méndez, Rifá and Regalado in HPC 12016 (holotype: HIPC; other isotypes: B, HAC, HAJB, JE, ULV).
Isotype: PAL-Gr 63252—Cuba, “Prov. Holguín. Municipio Moa. “Concesión minera Santa Teresita” al este de Yamanigüey. Pinares y charrascales,” 31 Mar 2011, Borsch & al. 4624 (holotype: ULV [specimen shown in the protologue, Figure 1]; other isotypes: B, FR, HAJB, JE, ULV).
Holotype: PAL-Gr 17738—Greece, Crete, “Ep. Kidonia: circa refugium Vólika supra Kámbi, 1350 m. In apricis graminosisque, solo saxoso calcareo (ad 2100 m. ascendens),” 15 Oct 1966, Greuter 7738 (isotypes: ATH, B, E, FI, G, GB, K, LD, LE, M, P, S, UPA, W, hb. Zaffran).
Isotype: PAL-Gr 39667—Australia, [Western Australia, “between Moore and Murchison Rs”] (from holotype label), [Drummond VI] 98 (holotype [specified here]: K #190019; other isotypes: MEL, NSW, PERTH).
Holotype: PAL-Gr 17802—Greece, Crete, “Ep. Ierapetra: in faucibus inter Christòs et Metaxochóri, 450 m. In rupium calcarearum praeruptarum sole illustrium fissuris vix accessibilibus rarissima,” 27 Oct 1966, Greuter 7802 (isotypes: E, G, LD, W).
Isotype: PAL-Gr 20660—Greece, “Epirus, prov. Ioannina, distr. Dhodhoni: montes Timfi, in latere boreo-orientali montis Astraka, alt. 1900–2000 m. Ad rupes calcareas praeruptas,” 21 Aug 1974, Charpin 11336, Dittrich, Greuter 12508 and von Auw (holotype: G; other isotypes: LD, UPA, W, hb. Dittrich).
Holotype: PAL-Gr 4055—Libya, “Marwah, Gebel Akhdar,” 23 Jan 1967, Boulos 1380 (isotypes: CAI?, K).
Lectotype (Greuter in Fl. Medit. 25 [Spec. Iss.]: 161. 2015) [https://www.herbmedit.org/flora/FL25SI_157-166.pdf]): PAL-Gr 62676):—original watercolor drawing of Cynoglossum marifolium by Roxburgh (isotype: K).
Holotype: PAL-Gr 27081—Greece, “ex insula Kastellorizo (Megiste) Dodekanesi, in collibus inter planos Barpouti et Parboutti (in parte austro-occidentali insulae), alt. 50–100 m. In regione dumuletorum sempervirentium, solo lapidoso vel rupestri calcareo,” 18 Apr 1974, Greuter 11964 (isotypes: ATH, C, ERE, G, MA).
  • Dianthus fruticosus subsp. sitiacus Runemark in Bot. Not. 133: 488. 1980 [17601].
Isotype: PAL-Gr 17601—Greece, Crete, “Ep. Sitia: in insulae Psíra parte austro-occidentali, 150 m. In praeruptis rupestribus calcareis ad septentriones spectantibus frequens,” 27 Sep 1966, Greuter 7601 (holotype: E; other isotypes: K, LD, M, W).
Second-step holotype (specified here): PAL-Gr 38797—Greece, “Nomos Viotias, Eparchia Thivon. C. 6.6 km S of Xironomi village, along the road to Aliki. An ophiolithic area W of the road. Slopes with phrygana and low sparse shrubs, stony places. Alt. c. 320–350 m. Lat. 38°12′ N, Long. 23°02′ E,” 22 May 1998, Constantinidis 7693 (first-step holotype: UPA [2×]; other isotypes: B, C, UPA).
Isotype (holotype fragment): PAL-Gr 1848—Greece, Crete, “in rupe Asprokremnos dicta Sphakia [in fauce Kalous Lakkous, 2000′],” [15] July 1846, Heldreich [1640] (holotype: G-BOIS; other isotypes: G, G-BOIS, G-BU, FI, K [2x], WU [Halácsy, Greek hb.]).
Isotype: PAL-Gr 1847—Greece, Crete: “N. Rethymnis: cliffs N of Kria Vrisi (NW of Melabes),” 1 Jun 1964, Snogerup, Strid and Bothmer 20955 (holotype: LD; other isotypes: none).
Holotype and isotype: PAL-Gr 14669 and 14669a—Greece, Crete: “Ep. Sfakia: unterer Teil der Schlucht Dòmata wl. Aj. Rùmeli, 20–40 m.ü.M., Plattenkalk. Hfg. In Ritzen buschiger O-exp. Schluchtwände,” 5 Jun 1962, Greuter 4669 (holotype: LD; other isotypes: G, LD, W, Z).
Holotype: PAL-Gr 126731—Cuba, “Prov. Villa Clara, Munic. Corralillo, entre las Cañas y el arroyo Clarita, alt. 85 m. 22°50′55″ N, 80°28′22″ W. Sabana en suelo mocarrero (con capa superficial de glomérulos ferralíticos),” 4 Mar 2019, Greuter 29687 & al. (isotypes: B, HAJB, JE, ULV).
  • Erysimum candicum subsp. carpathum Snogerup in Opera Bot. (Lund) 13: 38. 1967 [15516].
Isotype: PAL-Gr 15516—Greece: “Regio aegaea austr., insula Karpathos: in faucibus Kounoupa ad septemtriones oropedii Kato Lastos, alt. 600 m. In fissuris rupium calcarearum praeruptarum,” 23 May 1963, Greuter 5516 (holotype: LD; other isotype: Z).
Isolectotype: PAL-Gr 2772—Greece: “In m. Panassi reg. media,” Aug 1855, Guicciardi in Heldreich, Fl. Graec. Exs. (lectotype [designated here]: G #754289; other isolectotypes: C, G #147474, K, LD, MPU, WU).
Holotype: PAL-Gr 14849—Greece, Crete: “Ep. Sfakia: Hügel zwischen den Alpen Màvro Korfàli u. Eklissìdia am N-Fuss des Berges Pàchnes, 1800–1900 m.ü.M., Kalk. Nicht selten, doch nur noch spärl. mit Früchten im Verwitterungsschutt auf schwarzem Fels; abweichend duch punktierte Kapseln, glatte Samen und Blattränder,” 3 Jul 1962, Greuter 4849 (isotypes: G, HUJ, LD, W, Z).
Isotype: PAL-Gr 35829—Greece, Crete: “Nom. Chanion, Ep. Sfakion: Middle and lower parts of the Imbros ravine, NE of Chora Sfakion, 200–400 m. Forming loose cushions up to 1 m diam. in rock crevices. Lvs turning yellow,” 12 May 1988, Strid 26697 (holotype [to be specified]: C [2×]; other isotypes: ATH, B, E, G, LD [3×], UPA).
  • Exostema lucidum Borhidi and M. Fernández Fernández in Acta Bot. Hung. 35: 302. 1989. (≡Exostema lancifolium var. lucidum (Borhidi and M. Fernández) Borhidi) [http://real-j.mtak.hu/3649/ (accessed on 6 March 2024)].
Isotype: PAL-Gr 126209—Cuba, “Prov. Holguín, Mun. Moa: Cuchillas de Moa, barranco del arroyo Jaragua cerca de la Mina Jaragua, charrascales sobre roca ultrabásica, 200 m,” 3 May 1980, Álvarez de Zayas & al. in HFC 42659 (holotype [specified here]: HAJB #453; other isotypes: B, HAJB ##454–459 [6×], JE [2×]).
  • Festuca armoricana Huon ex Kerguélen in Lejeunia, ser. 2, 75: 9. 1975 [9416].
Isotype: PAL-Gr 9416—France: “Hillion (France, Côtes-du-Nord), Pointe de Lermaot, sables maritimes fixés et pelouses sur falaises basses, avec Thymus drucei, Dactyis glomerata et Foeniculum vulgare. 2n = 28,” 27 May 1972, Kerguélen in Soc. Échange Pl. Vasc. Europe Occid. Bassin Médit. #7211 (holotype [specified here]: P #748568; other isotypes: CHAPA, G, GENT, MA [2×], MICH, MSB, P #748567, SEV).
Isotype: PAL-Gr 30709—Greece: “Macedonia: Distr. Grevena: Montes Pindus: In declivibus boreali-orientalibus montis Aphtia, in valle Arkudolaka (Valea Kalda) ditionis pagi Perivoli, substr. serpent., ca. 1700 m,” 30–31 Jul 1956, Rechinger 18489 (holotype [specified here]: W #197900012707; other isotypes: B, GZU, MA, MO, RSA, S, W #197200001938).
Isotype: PAL-Gr 30710—Greece: “Macedonia: Penins. Chalkidike. Montes Cholomonda, in quercetis, ca.500–800 m,” 5 Jun 1955; Rechinger 17253 (holotype: [specified here]: W #197900012638; other isotype: W #197200001966).
Isotype: PAL-Gr 9406—France: “Kerloc’h, près Crozon (France, Finistère), falaises maritimes au S de la plage, avec Dactyis glomerata, Armeria maritima, Hieracium pilosella, Scilla verna, Sanguisorba minor, Hypochoeris radicata, Anagallis parviflora, Thymus drucei, Sedum acre, Euphorbia portlandica, Daucus gummifer et Anthyllis vulneraria, pH = 5.8,” 26 May 1972, Kerguélen and Authier 790 in Soc. Échange Pl. Vasc. Europe Occid. Bassin Médit. #7221 (holotype: LG; other isotypes: BR, CHAPA, G, K, MA, MSB, P [3×], SEV).
  • Festuca ophioliticola Kerguélen in Lejeunia, ser. 2, 75: 13. 1975 (≡Festuca ovina subsp. ophioliticola (Kerguélen) M. Wilk. 1985) [9398].
Isotype: PAL-Gr 9398—France: “Peumérit (France, Finistère), entre Peumérit et Lespurit-Quélenn, sur les rochers de serpentines (ancienne carrière) et dans la lande à Erica cinerea, Ulex europaeus, Thesium humifusum, Sieglingia decumbens, Serratula tinctoria, Festuca arundinacea, , pH = 6, 2n = 28,” 24 May 1972, Kerguélen in Soc. Échange Pl. Vasc. Europe Occid. Bassin Médit. #7229 (holotype [specified here]: P #1887862; other isotypes: AV, CHAPA, G, GENT, K, L, MA [2×], P #1887293, SEV).
  • Festucopsis serpentini subsp. lurensis F. K. Mey. in Haussknechtia Beih. 15: 26. 2011 [52734].
Isotype: PAL-Gr 52734—Albania, “Lura, Kunora e Lures, ca. 1900 m,” 2 Aug 1959, Meyer 4682 (Holotype [specified here]: JE #16665; other isotype: JE #16666).
  • Gymnospermium altaicum subsp. peloponnesiacum Phitos in Strid and Tan, Fl. Hellenica 2: 81. 2002 (≡Gymnospermium peloponnesiacum (Phitos) Strid 2009) [42787].
Isotype: PAL-Gr 42787—Greece, “Prov. Achaia: mons Panachaikon, in declivibus occidentalibus, 1400–1500 m; in petrosis,” 13 May 1987, Phitos and Kamari 19922 (holotype: UPA; other isotype: C).
Isoneotype: PAL-Gr 62429—Italy, Sicilia: “Monte Gallo, near Palermo, 38°12′57″ N, 18°36′ E, 450 m a.s.l., on rocky limestone habitat,” 9 Jun 2014, Domina (neotype [see Iamonico and Domina in Pl. Biosyst. 149: 725. 2015]: PAL; other isoneotypes: FI, G, RO).
Isotype: PAL-Gr 39673—Australia, [Western Australia, “between Moore and Murchison Rs”] (from holotype label), [Drummond VI] 82 (holotype [specified here]: K #704050; other isotypes: MEL [2×], PERTH [fragment]).
Isotype: PAL-Gr 128204—Cuba, “Prov. Holguín, municipio Rafael Freyre, al este de la Bahía de Naranjo, camino al restaurante ‘conuco de Mongo Viña’, matorral xeromorfo espinoso sobre serpentina. 10 m s.m., 21°06′00″ N, 75°52′14.6″ W,” 30 May 2019, González Gutiérrez 1905-1 (holotype: HAJB #1216; other isotype: B).
Isotype: PAL-Gr 133251—Cuba, “Prov. Guantánamo, San Antonio. Abra de Mariana, en el barranco 5 km al NO de San Antonio del Sur,” 11 May 1980, Álvarez & al. in HFC 43077 (holotype: JE #28983; other isotypes: B, HAJB, JE #28984).
Isotype: PAL-Gr 55254—Spain, “Palencia: Cevico Navero, 30TVM3602, 850 m, en matorrales sobre Margas y suelos de costra yesífera, en comunidades de Aphyllanthion,” 11 Jun 1982, Fdez. Díez (holotype: SEV #93257; other isotypes: BC, G, MAF, SALA).
Second-step holotype (specified here): PAL-Gr 16559—Greece, “Aegäis, Nom. Attikì, Ep. Kìthira: bei der Kirche Odijìtria söl. Livàdi, 170 m.ü.M., sandig verwitterndes Neogen. Nicht selten in der Cistusphrygana,” 12 May 1964, Greuter 6559 (isotypes: G, W).
Isotype: PAL-Gr 45143—Germany, “Thüringen, Landkreis Hildburghausen, Gleichamberg, Großer Gleichberg, 650 m s.m., ehemaliger Steinbruch, Schutthalden, MTB 5629/21, 50°22′ N, 10°35′ E,” 30 May 1998, Gottschlich 35725 and Meierott in Hieracia Europ. Sel. #112 (holotype: M; isotypes: G, FI, GOET).
Isotype: PAL-Gr 125290—Italy, “Friaul-Julisch Venetien, Prov. Udine, Comeglians, M. Crostis, (), 1883 m s.m., Feinschutt (Kalk?), MTB 9443/2, 46°33′21″ N 12°55′32″ E,” 22 Jul 2016, Gottschlich-65761 in Hieracia Europ. Sel. #706 (holotype: FI; isotypes: FR, H, IBF, LI, M, PRA, W, Hb. Brandstätt., Hb. Dunkel, Hb. Gottschl.).
Isotype: PAL-Gr 52625—Italy, “Friaul-Julisch Venetien, Prov. Pordenone, Erto e Casso, untere Kehren im Aufstieg zum Val Zemola, 1000 m s.m., Kalkschutt und Gebüsche, MTB 9740/1, 46°16′ N, 12°23′ E,” 8 Aug 2009, Gottschlich-55066 in Hieracia Europ. Sel. #362 (holotype: FI; Isotypes: B, GOET, H, IBF, LI, M, PRC, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 62680—Italy, “Campania, Montevergine, Mercogliano (AV). Coordinate: 40°56′34.52″ N, 14°43′35.63″ E. Alt.: 1290 m. Esposiz.: NO. Habitat: carbonate rocks,” 07 Jul 2014, Di Gristina [101733] (holotype: PAL!; other isotypes: B!,hb. Gottschl.).
Isotype: PAL-Gr 45237—Italy, “Abruzzen, Prov. L’Áquila, Gran Sasso, Calascio: S “Piano Buto,” 1050 m, Kalkfelsböschung, MTB 3648/3, 42°18′ N, 13°40′ E,” 3 Jun 2004, Gottschlich 48828 in Hieracia Europ. Sel. #208 (holotype [to be specified]: FI; Isotypes: B, FI, GOET, H, IBF, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45236—Italy, “Abruzzen, Prov. Téramo, Gran Sasso, Sent. 2: zwischen Rif. Duca degli Abruzzi und M. Portella, 2300–2320 m, nordexponierte Kalkschutthänge, MTB 3547/3, 42°26′ N, 13°32′ E,” 31 Jul 2001, Gottschlich 44649 and Dunkel in Hieracia Europ. Sel. #207 (holotype [to be specified]: FI; isotypes: B, FI, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 58829—Greece, “Prefecture of Chalcidice, C. Chalcidice, 40°30′13,6″ N, 23°33′13,5″ E, 547 m, At the slope of road. Rocky position near at heathland,” 6 Jun 2011, Melikoki 6.13.3 (holotype: TAUF; other isotypes: ATHU, B, C, FI, LD, UPA, W, hb. Gottschl.).
  • Hieracium boreoapenninum Gottschl. in Webbia 64: 3. 2009 [48439].
Isotype: PAL-Gr 48439—Italy, “Ligurien, Prov. Génova, Montebruno: Passo della Scoglina, 920–930 m s.m., Kammweg. MTB 1521/1, 44°28′ N, 9°14′ E, Gottschlich 52362 in Hieracia Europ. Sel. #275 (holotype: FI; other isotypes: B, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 121155—Italy, “Toskana, Prov. Lucca, Alpi Apuane, Arni, aufgelassener Marmor-Steinbruch bei “Tre Fiumi,” 761 m, Marmorfeinschutt mit Weidenbüschen, MTB 1927/2, 44°03′18″ N, 10°16′21″ E,” 1 Jul 2015, Gottschlich 64325 in Hieracia Europ. Sel. #611 (holotype: FI; other isotypes: B, FR, H, IBF, LI, M, PRA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 129006—Greece, “Ostmakedonien, Nom. Xanthi, Lydodromio → Ag. Georgios, 550 m s.m., Gebüsch, 41°14′22″ N, 24°45′51″ E,” 24 Jun 2019, Gottschlich 72651 and Dunkel in Hieracia Europ. Sel. #801 (holotype: B; other isotypes: FI, FR, H, IBF, LI, M, MSTR, UPA, W, Hb. Brandstätt., Hb. Dunkel, Hb. Gottschl.).
  • Hieracium calodon subsp. rhenovulcanicum Gottschl. and Heinrichs in Decheniana 154: 8. 2001 (≡Pilosella rhenovulcanica (Gottschl. and Heinrichs) Bomble 2020) [45056].
Isotype: PAL-Gr 45056—Germany, “Rheinland-Pfalz, Landkreis Mayen-Koblenz, Plaidt, Autobahn-Abfahrt Plaidt, 146 m s.m., Straßenrand, MTB 5610/12, 50°23′ N, 7°24′ E,” 25 May 1995, Gottschlich 27383, Heinrichs and Raabe in Hieracia Europ. Sel. #25 (holotype: B #100024153; other isotypes: BONN, GOET, HBG, M).
Isotype:PAL-Gr 48433—Italy, “Abruzzen, Prov. Chieti, Majella: Sent. 1: Blockhaus → Monte Cavallo → Bivacco Fusco, 2045–2100 m s.m., Pinus mugo-Gebüsche. MTB 3850/4, 42°07′ N, 14°06′ E” 31 Jul 2000, Gottschlich-41661 and Dunkel in Hieracia Europ. Sel. #269 (holotype [to be specified]: FI; other isotypes: B, FI, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
  • Hieracium cepitinum Gottschl. in Stapfia 89: 63. 2009 [≡Pilosella cepitina (Gottschl.) Gottschl. 2010; =Pilosella arnoserioides (Nägeli and Peter) Soják in Preslia 43: 186. 1971] [https://www.zobodat.at/pdf/STAPFIA_0089_0001-0328.pdf (accessed on 6 March 2024)].
Isotype: PAL-Gr 45190—Italy, “Abruzzen, Prov. Téramo: Gran Sasso, Pietracamela, Straße Richtung Prati di Tivo, Pietracamela: E C.le Cepito, 1220 m s.m., Kalkmagerrasen, MTB 3447/3, 42°31′ N, 13°33′ E,” 6 Jun 2001, Gottschlich 44066 and Dunkel in Hieracia Europ. Sel. #159 (holotype [to be specified]: FI; other isotypes: B, FI, GOET, H, IBF, LI, M, PR, W, hb.Brandstätt., hb.Dunkel, hb.Gottschl.).
Isoepitype: PAL-Gr 66254—Spain, “Lleida: Tredòs (Vall d’Aran), cap al Lac de Baish de Bacivèr, 31TCH3329, datum ED50, 1920 m, roques calcàries,” 24 Jul 2012, Mateo, Rosselló, Sàez and del Egido (epitype [Ferrer-Gallego & al. in Willdenowia 45: 388. 2015: https://www.jstor.org/stable/24753246?seq=3]: VAL #210645; other isoepitypes: B, LEB, VAL #227665 [interpreting the lectotype: Linn. No. 954.28 (LINN) [http://linnean-online.org/8846]).
Isotype: PAL-Gr 121152—Italy, “Toskana, Prov. Lucca, Alpi Apuane, Umgebung Rif. Val Serenaia, 1040–1060 m s.m., Schuttflächen, Abbruchkanten, Bachbett, MTB 1827/3, 44°08′09″ N, 10°11′49″ E,” 17 Jun 2014, Gottschlich 62663 in Hieracia Europ. Sel. #608 (holotype: FI; other isotypes: B, FR, H, IBF, LI, M, PRA, TO, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45215—Italy, “Prov. L’Áquila, Val Roveto, Rendinara: NNE-exponiertes Kar des M. Pratillo, 1900–1970 m s.m., Kalkschutt, Felsrasen. MTB 4146/4, 41°48′ N 13°29′ E,” 30 Jul 2004, Gottschlich 48977 and Dunkel in Hieracia Europ. Sel. #186 (holotype [to be specified]: FI; isotypes: (B, FI, GOET, IBF, H, LI, M, PR, W, hb. Brandstätter, hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45241—Italy, “Prov. L’Áquila, Gran Sasso,: Barisciano: N-Hang des unbenannten Berges “1516 m” zwischen M. della Selva und M. Capanello, 1350 m s.m., lichter Kiefernforst, MTB 3647/2, 42°21 N, 13°35′ E,” 9 Jun 2003, Gottschlich 47509, Dunkel, Conti, Tinti and Londrillo” in Hieracia Europ. Sel. #212 (holotype [to be specified]: FI; isotypes: B, FI, GOET, IBF, H, LI, M, PR, W, hb. Brandstätter, hb. Dunkel, hb. Gottschl.).
  • Hieracium dasycraspedum Buttler in Strid, Mount. Fl. Greece 2: 630. 1991 [20638].
Holotype: PAL-Gr 20638—Greece, “Epirus, prov. Ioannina, distr.Dhodhoni: montes Timfi, in latere boreali montis Astraka supra refugium, alt. 2200 m. In scansilibus graminosis rupium calcarearum,” 21 Aug 1974, Charpin 11358, Dittrich, Greuter 12530 and von Auw (isotypes: G, LD, M, UPA, hb. Dittrich).
  • Hieracium dentatum subsp. montiscatriae Gottschl. in Webbia 66: 216. 2011 [52608].
Isotype: PAL-Gr 52608—Italy, “Marche, Prov. Pésaro-Urbino, M. Cátria, NW-Hang, S Rif. della Vernosa, 1500–1600 m s.m., offene Kalkfelsrasen, MTB 2542/1, 43°27′56″ N, 42°14′ E,” 1 Aug 2000, Gottschlich 55854 and Dunkel in Hieracia Europ. Sel. #355 (holotype: FI; other isotypes: B, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
  • Hieracium dentatum subsp. setuligerum Gottschl. in Webbia 66: 218. 2011 [52606].
Isotype: PAL-Gr 52606—Italy, “Marche, Prov. Macerata, Monti Sibillini, Ussita, Frontignano, S-Hang M. Bicco oberhalb Sent. 158, 1820–1880 m s.m., Kalkschutt. MTB 3045/3, 42°54′52″ N, 13°11′07,”E” 12 Jul 2009, Gottschlich 54828 in Hieracia Europ. Sel. #353 Holotyppe: FI; other isotypes: B, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 52606—Germany, “Niedersachsen, Landkreis Friesland, Urwaldstraße westlich Bockhorn, 5 m s.m., Straßenböschung, MTB 2614/11 (53°23′47′′ N, 8°0′15′′ E), 10 Jun 2015, Schabelreiter 1444 in Hieracia Europ. Sel. #702 (holotype: B; other Isotypes: FI, FR, H, IBF, LI, M, PRA, W -1444, hb. Dunkel, hb. Gottschl.-64487, hb. Schabelreit.).
Isotype: PAL-Gr 125292—Italy, “Friaul-Julisch Venetien, Prov. Udine, Paularo, Ligusullo → Rif. Valdajer, 1179 m s.m., oberhalb Straßenmauer unter Fichtenwald, MTB 9444/3, 46°32′34″ N, 13°04′50″ E,” 26 Jul 2013, Gottschlich 60728 in Hieracia Europ. Sel. #708 (holotype: FI; other isotypes: B, FR, H, IBF, LI, M, PRA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 125291—Italy, “Friaul-Julisch Venetien, Prov. Udine, Cividale, Riekatal: E Clodig, 319 m s.m., Straßenböschung. MTB 9847/2, 46°9′25″ N, 13°36′4″ E,” 30 May 2012, Gottschlich 58379 in Hieracia Europ. Sel. #707 (holotype: FI; other isotypes: B, FR, H, IBF, LI, M, PRA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
  • Hieracium diaphanoides subsp. forstense Gottschl. in Mitt. Pollichia 98: 76. 2017 [file:///C:/Users/wg/Dropbox/PC/Downloads/Mitt-Pollichia_98_0075-0078(4).pdf.
Isotype: PAL-Gr 123321—Germany, “Rheinland-Pfalz, Landkreis Bad Dürkheim/Weinstraße, Forst: “Grüne Bank” westlich oberhalb Forst, 390 m s.m., Eichen-Kiefern-Wald mit Kastanien. MTB 6514/42, 49°25′33″ N, 8°9′44″ E,” 28 May 2015 Gottschlich 63987 in Hieracia Europ. Sel. #651 (holotype: B; other isotypes: FI, FR, H, IBF, LI, M, POLL, PRA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
  • Hieracium diaphanoides subsp. volakasense Gottschl. and Dunkel in Stapfia 111: 18. 2019 [file:///C:/Users/wg/Dropbox/PC/Downloads/STAPFIA_0111_0005-0032.pdf.
Isotype: PAL-Gr 129010—Greece, “Ostmazedonien, Nom. Drama, M Falakro: Straße zum Skicenter Falakro, 1050 m s. m., Kalkfels unter Buchenwald. 41°17′46″ N, 24°0′11″ E” 21 Jun 2019, Gottschlich 72493 and Dunkel in Hieracia Europ. Sel. #805 (holotype: B; isotypes: FI, FR, H, IBF, LI, M, MSTR, UPA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45180—Italy, “Südtirol, Prov. Bozen, Schnalstal: Straße entlang des Vernagt-Stausees, 200 m E Finailbach, 1750 m s.m., felsiger Steilhang mit Lärchen. MTB: 9230/4, 46°44′ N, 10°49′ E,” 22 Jul 2000, Gottschlich 41544 in Hieracia Europ. Sel. #149 (holotype: LI, other isotypes: FI, G, IBF, W, hb. Gottschl.).
Isotype: PAL-Gr 45124—Italy, “Südtirol, Prov. Trento: Campitello di Fassa, Valle di Duron, E Rif. Micheluzzi, 1840–1880 m s.m., Schneise im Bergfichten-Wald auf Basalt. MTB 9536/1, 46°29′ N, 11°42′ E,” 2 Jul 2005, Gottschlich 44748, Dunkel, Prosser, Pujatti, and Zidorn in Hieracia Europ. Sel. #93 (holotype: FI; other isotypes: ROV, hb. Gottschl.).
  • Hieracium balearicum Arv.-Touv., Hier. Gall. Hisp. Cat.: 157. 1913) [8536] (≡Hieracium elisaeanum subsp. balearicum (Arv.-Touv.) Greuter 2007).
Isotype: PAL-Gr 8536—Spain, Balearic Islands: “Balearium insula Majore in rupestribus mis Puig mayor de Torella fal. calcar., 1000–1300 m s.m.,” 8 Jun 1885, Porta & Rigo in Iter Hisp. ‚608 (holotype [specified here]: GRM; other isotypes: many).
Isotype: PAL-Gr 121156—Italy, “Toskana, Prov. Lucca, Alpi Apuane, Val Serenaia, Umgebung Rifugio Val Serenaia, 1040–1060 m s,m., Schuttflächen, Abbruchkanten, Bachbett. MTB 1827/3, 44°8′9″ N, 10°11′49″ E” 17 Jun 2014, Gottschlich 62662 in Hieracia Europ. Sel. #612 (holotype: FI; other isotypes: B, FR, H, IBF, LI, M, PRA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45264—Italy, “Abruzzen, Prov. L’Áquila, Gran Sasso, Calascio: S “Piano Buto,” 1050 m s.m., Kalkfelsböschung. MTB 3648/3, 42°18′ N, 13°40′ E” 3 Jun 2004, Gottschlich 48826 in Hieracia Europ. Sel. #235 (holotype [to be specified]: FI; other isotypes: B, FI, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
  • Hieracium faucis-jovis (‘faucisjovis’) Gottschl. in Webbia 64: 182. 2009 [48447].
Isotype: PAL-Gr 48447—Italy, “Toskana, Lucca, Bagni di Lucca: S unterhalb Foce a Giovo, 1650 m s.m., Felsheiden, Wiesen, Sandstein. MTB 1829/4, 44°7′ N, 10°36′ E,” 1 Jul 2005, Gottschlich 50227 in Hieracia Europ. Sel. #283 (holotype: FI; other isotypes: B, GOET, IBF, H, LI, M, PRA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 52611—Italy, “Piemont, Prov. Torino, Valle di Susa, zwischen Salbertrand und Pramand, S Galleria dei Sarazeni, 2170 m s.m., feuchte Balmen. MTB 0609/2, 45°4′33″ N, 6°49′33″ E,” 10 Aug 2010, Gottschlich 56082 in Hieracia Europ. Sel. #358 (holotype: FI; other isotypes: B, GOET, H, IBF, LI, M, PRC, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45246—Italy, “Abruzzen, Prov. L’Áquila, Roccaraso: an der Straße zum Wintersportgebiet “Aremogna,” oberhalb der großen Hotels, Kalkschutthang, 1530 m s.m., MTB 4150/1, 41°50′ N, 14°3′ E,” 1 Jun 2001, Gottschlich 43900 in Hieracia Europ. Sel. #217 (holotype: FI [to be specified]; other isotypes: B, FI, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45230—Italy, “Abruzzen, Prov. Téramo, Montagna dei Fiori: zwischen San Giacomo und Ripe: SW Abzweigung Richtung Cerqueto, 890 m s.m., Kiefernforst. MTB 3247/2, 42°46′ N, 13°37′ E,” 30 May 2004, Gottschlich 48696 in Hieracia Europ. Sel. #201 (holotype: FI [to be specified]; other isotypes: B, FI, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 121148—Italy, “Piemont, Prov. Torino, Canavese, Valperga: Auffahrt Sacro Monte di Belmonte 626 m s.m., Granitfels. MTB 0611/2, 45°21′57″ N, 7°37′24″ E,” 25 May 2013, Gottschlich 59937 in Hieracia Europ. Sel. #604 (holotype: FI; other isotypes: B, FR, H, IBF, LI, M, PR, TO, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 121154—Italy, “Toskana, Prov. Lucca, Alpi Apuane, Val Serenaia, sent. 179 unterhalb “Cresta Garnerone,” 1475 m s.m., Marmorfelsrasen. MTB 1827/3, 44°7′30″ N, 10°11′33″ E,” 11 Aug 2014, Gottschlich 63270, Soldano and Soldano in Hieracia Europ. Sel. #610 (holotype: FI; other isotypes: B, FR, H, IBF, LI, M, PRA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 48589—France [Pyrénées-orientales], “Cerdagne: Gorges de Llo [Sègre], éboulis herbeux, 1520 m. 30 Jul 1923, Sennen in Pl. Espagne #4768. Holotype [specified here]: BC; other isotypes: BM, MPU, PH [2×], RNG).
Isotype: PAL-Gr 45125—Austria, “Kärnten, Ankogelgruppe: Kölnbreinspeicher, Nordufer des Stausees, Wanderweg 02, zwischen Abzweigung ins Kleinelendtal und Osnabrückerhütte, ca. 500 m SSW P. 1916 (sphalm. “2916“), 1910–1920 m s.m., Grünerlengebüsch mit Calluna und Hochstaudenfragmenten, Silikat. MTB 8945/2, 47° 4′ N, 13°18′ E,” 13 Aug 2005, Dunkel 13747 in Hieracia Europ. Sel. #94 (holotype: W, other isotypes: KL, hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45272—Italy, “Abruzzen, Prov. Téramo, Gran Sasso: Vado di Sole → Castelli: ESE Mte. Vito, 1350 m s.m., Buchenwaldrand. MTB 3548/4, 42°24′ N, 13°47′ E,” 30 Jul 2002, Gottschlich 46235 in Hieracia Europ. Sel. #243 (holotype [to be specified]: FI; other isotypes: B, FI, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45269—Italy, “Abruzzen, Prov. L’Áquila, Carsóli: oberhalb Tufo, 900 m s.m., Buchenwald mit Castanea sativa. MTB 3844/4, 42°9′ N, 13°6′ E,” 29 May 2004, Gottschlich 48660 in Hieracia Europ. Sel. #240 (holotype [to be specified]: FI; other isotypes: B, FI, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 129011—Greece, “Nom. Kilkis, M. Paiko, Straße S Livadia, beim Aussichtspunkt, 1110 m s.m., felsige Straßenböschung, Kalk. 40°59′22″ N, 22°20′48″ E,” 19 Jun 2019, Gottschlich 72391 and Dunkel in Hieracia Europ. Sel. #806 (holotype: B; other isotypes: FI, FR, H, IBF, LI, M, MSTR, UPA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 121154—Italy, “Toskana, Prov. Massa-Carrara, Alpi Apuane, sent. 172: Foce di Pianza → M. Sagro, 1300–1500 m s.m., Marmorfelsspalten und Felsrasen. MTB 1826/4, 44°6′ N, 10°9′ E,” 2 Jul 2005, Gottschlich 50251 and Marchetti in Hieracia Europ. Sel. #605 (holotype: FI; other isotypes: B, FR, H, IBF, LI, M, PRA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45068—Italy, “Südtirol, Prov. Bozen: Vinschgau, Straße Kastellbell Richtung St. Martim im Kofel, Kehren unterhalb Koben, 1500 m s.m., lichter Kiefernwald, Orthogneis/Glimmerschiefer. MTB: 9331/3, 46°38′ N, 10°51′ E,” 19 Jun 1999, Gottschlich 38147 and Dunkel in Hieracia Europ. Sel. #37 (holotype: LI; other isotypes: IBF, hb. Dunkel, hb. Gottschl., hb. Naturkundemus. Bozen).
Isotype: PAL-Gr 121144—Italy, “Piemont, Prov. Biella, Rosazza, T. Pragnetta:. Parkplatz/Einstieg sent. E 30, Parkplatzrand, Alluvionen, 949 m s.m. MTB 0313/2, 45°40′26″ N, 7°58′22″ E,” 6 Aug 2014, Gottschlich-63201 in Hieracia Europ. Sel. #601 (holotype: FI: other isotypes: B, FR, H, IBF, LI, M, PRA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
  • Hieracium lanudae Gottschl. in Webbia 64: 179. 2009 [48446].
Isotype: PAL-Gr 48446—Italy, “Emilia-Romagna, Prov. Reggio, Passo del Cerreto, M. La Nuda: Bergstation Sessellift bis Ende Schlepplift, 1500–1700 m ss.m., Silikat-Felsschutt. MTB 1727/1, 44°16′ N, 10°14′ E,” 3 Jul 2005. Gottschlich 50262 in Hieracia Europ. Sel. #282 (holotype: FI; other isotypes: B, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 63660—Italy, “Piemont, Prov. Cuneo, Valle Máira, Vallone di Elva, 1213–1250 m s.m., Kalkfels an der Straße. MTB 1408/4, 44°31′3″ N, 7°5′24″ E,” 8 Jul 2013, Gottschlich-60566 and Dunkel in Hieracia Europ. Sel. #551 (holotype: FI; other isotypes: B, FR, H, IBF, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
  • Hieracium marchesonii Gottschl. in Webbia 66: 204. 2011 [52605].
Isotype: PAL-Gr 52605—Italy, “Marche, Prov. Macerata, Monti Sibillini, Ussita, Frontignano, S-Hsng M. Bicco oberh. Sent. 158, 1820–1880 m s.m., Kalkschutt. MTB 3045/3, 42°54′52″ N, 13°11′7″ E,” 12 Jul 2009, Gottschlich 54829 in Hieracia Europ. Sel. #352 (holotype: FI; other isotypes: B, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 125293—Italy, “Friaul-Julisch Venetien, Prov. Udine, Ravascletto, Sträßchen zum Pizzo di Mede, 1302 m s.m., Calamagrostis-Bestand am Fichtenwaldrand. MTB 9443/4, 46°31′58″ N, 12°56′7″ E,” 7 Aug 2011, Gottschlich-57551 in Hieracia Europ. Sel. #709 (holotype: FI; other isotypes: B, FR, H, IBF, LI, M, PRA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 121163—Italy, “Toskana, Prov. Massa-Carrara, Alpi Apuane, “Dogna della Techhia” NE Castelpoggio, 900 m s.m., Gebüsche auf Kalkfeinschutt. MTB 1826/4, 44°7′ N, 10°5′ E,” 26 May 2004, Gottschlich-48586 in Hieracia Europ. Sel. #619 (holotype: FI; Isotypi: Hier. Eur. Sel. Nr. 619 (B, FR, H, IBF, LI, M, PAL, PRA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45112—Austria, “Niederösterreich, Östliche Niederösterreichische Voralpen, Mitterbach, Gemeindealpe, Umgebung Gipfelkreuz, 1600–1626 m s.m., Weiden, Latschengebüsche. MTB 8157/3, 47°48′ N, 15°14′ E,” 17 Jul 2005, Gottschlich 50396 in Hieracia Europ. Sel. #81 (holotype: W #2007–20446, other isotypes: hb. Gottschlich, etc.).
  • Hieracium murorum subsp. nivimarginatum Gottschl. in Webbia 66: 224. 2011 [54848].
Isotype: PAL-Gr 54848—Italy, “Marche, Prov. Macerata, M. Sibillini. Ussita: Frontignano, Abstieg neben der Seilbahn, 1400–1700 m, Kalkschutt u. Kalkrasen mit Bucheninseln. MTB 3045/3, 42°54′59″ N, 13°10′45″ E,” 12 Jul 2009, Gottschlich 54833 (holotype: FI; other isotypes: B, M, hb. Dunkel, hb. Gottschl.).
  • Hieracium necopinum Buttler in Strid, Mount. Fl. Greece 2: 639. 1991 [28275].
Holotype: PAL-Gr 28275—Greece, “Epirus (UTM: DK-4; Distr. No. 514), montes Timfi: in ascensu a lacuna media (infra refugium E.O.S. sita) ad lacunam Dhrakolimni, praesertim in latere boreali monis Ploskos, alt. 1750–2000 m. 23 Jul 1977, Greuter 15015 & al. p.p. (two plants) (isotypes: none).
Isotype: PAL-Gr 45038—Italy, “Südtirol, Prov. Trento, Brentonico: M. Baldo, M. Altissimo, “Pra delle Varsive” N Rif. Graziani, 1650- 1800 m s.m., alpine Matten. MTB: 0131/3, 49°48′ N, 10°53′ E,” 15 Jun 1984, Gottschlich 28238, Festi, Prosser & Pujatti in Hieracia Europ. Sel. #7 (holotype: LI; other isotypes: ROV, hb. Gottschl.).
Isotype: PAL-Gr 52610—Italy, “Südtirol, Prov. Bozen, Vinschgau, Schnalstal: Waalweg 3 zwischen Schloß Juval und Altratteis, 800–900 m s.m., Laubmischwald. MTB 9331/2, 46°39′ N, 10°57′ E,” 5 Sep 1999, Gottschlich-39050 in Hieracia Europ. Sel. #112 (holotype: FI, other isotypes: B, GOET, H, IBF, LI, M, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45081—Italy, “Friaul-Julisch Venetien, Prov. Udine, Forni Avoltri: zwischen Frasseneto und Sigilleto, 1115 m s.m., Schieferböschung. MTB: 9442/2, 46°34′ N, 12°48′ E,” 10 Aug 2000, Gottschlich 41738 in Hieracia Europ. Sel. #50 (holotype: LI; other isotypes: FI, MFU, W, hb. Gottschl.).
Isotype: PAL-Gr 48435—Italy, “Abruzzen, Prov. Téramo, Gran Sasso: Kammweg Vado di Corno → M. Brancastello, N-Seite, 1950–2000 m s.m., Kalkfelsrasen, Kalkfelstreppen. MTB 3547/2, 42°27′ N, 13°36′ E,” 30 Jul 2005, Gottschlich 50521 in Hieracia Europ. Sel. #271 (holotype [to be specified]: FI; other isotypes: B, FI, GOET, IBF, H, LI, M, PR, W, hb. Brandstätter, hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45200—Italy, “Abruzzen, Prov. L’Áquila, Campo di Giove, M. Porrara: N-Ende der “Paradina,” Kammjoch, 1750 m s.m., zerklüfteter Kalkfels. MTB 4050/2, 41°59′ N, 14°5′ E,” 29 Jul 2004, Gottschlich 48951 in Hieracia Europ. Sel. #169 (holotype: FI [to be specified]; other isotypes: B, FI, GOET, IBF, H, LI, M, PR, W, hb. Brandstätter, hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 121153—Italy, “Toskana, Prov. Lucca, Alpi Apuane, Val Serenaia, sent. 179, unterhalb “Cresta Garnerone,” 1476 m s.m., Marmorfelsspalten. MTB 1827/3, 44°7′41″ N, 10°11′21″ E,” 11 Aug 2014, Gottschlich 63271, Soldano and Soldano in Hieracia Europ. Sel. #609 (holotype: FI; other isotypes: B, FR, H, IBF, LI, M, PRA, W, hb. Brandstätter, hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 52613—Italy, “Piemont, Prov. Torino, Valle di Susa, Straße zwischen Susa und Colle di Finestre bei “Piano del Tiraculo,” 2165 m s.m., Zwergstrauchheiden. MTB 0908/1, 45°4′21″ N, 7°3′13″ E,” 8 Aug 2010, Gottschlich 55947 in Hieracia Europ. Sel. #360 (holotype: FI; other isotypes: B, GOET, H, IBF, LI, M, PRC, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
  • Hieracium permaculatum subsp. cacuminicola Gottschl. in Webbia 66: 226. 2011 [52607].
Isotype: PAL-Gr 52607—Italy, “marche. Prov. Pésaro-Urbino, Nordseite M. Nerone, 1470 m s.m., Kalkfelsstufen, Kalkschutt. MTB 2441/1, 43°33′30″ N, 12°31′3″ E,” 10 Jul 2010, Gottschlich 55485 in Hieracia Europ. Sel. #354 (holotype: FI; other isotypes: B, GOET, IBF, H, LI, M, PESA, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isoneotype: PAL-Gr 45239—Italy, “Abruzzo, Prov. Áquila, Gran Sasso M. Corvo: Trockental “Solagne” SE Stazzo di Solagne, 1700–1900 m s.m., lichter Buchenwald, Kalkschutt. MTB 3546/2, 42°28′ N, 13°27′ E,” 27 Jul 2004, Gottschlich 48919 in Hieracia Europ. Sel. #210 (neotype [Gottschlich in Stapfia 89: 160. 2009: [https://www.zobodat.at/pdf/STAPFIA_0089_0001-0328.pdf; to be specified]): FI; other isoneotypes: B, FI, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
  • Hieracium pintodasilvae de Retz in Agron. Lusit. 35: 307. 1974 (≡Pilosella pintodasilvae (de Retz) Mateo, Cat. Fl. Prov. Teruel: 143. 1990) [9226].
Isotype: PAL-Gr 9226—Portugal, “Bragança (Lusitania, Trás-os-Montes e Alto Douro), pr. Carragosa, 850 m s.m., in pratis,” 12 Jul 1971, Pinto da Silva, Teles and Martins 8849 in Soc. Échange Pl. Vasc. Europe Occid. Bassin Médit. #7027 (holotype: LISE #75233; other isotypes: many).
Isotype: PAL-Gr 45275—Italy, “Abruzzen, Prov. L’Áquila, Monte Simbruini: Capistrello → Valico Serra S. Antonio, 1150 m s.m., Straßenböschung. MTB 4046/1, 41°57′ N, 13°21′ E,” 5 Jun 2006, Gottschlich 51143 in Hieracia Europ. Sel. #246 (holotype [to be specified]: FI; other isotypes: B, FI, G, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45276—Italy, “Abruzzen, Prov. L’Áquila, Gran Sasso, Campo Imperatore: Trockental W Canali di Marcegli, 1600 m s.m., Kalkfels, Felsrasen, Schutt. MTB 3648/1, 42°23′ N, 13°42′ E,” 25 Jul 2004, Gottschlich 48863 in Hieracia Europ. Sel. #247 (holotype [to be specified]: FI; other isotypes: B, FI, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 129007—Greece, “Thessalien, Nom. Magnísia, Volos, SW Chanía, 750 m NW Agriolefkes, Mt. Pilio Ski Resort, 1220 m s.m., Straßenböschung unter Buchenwald. 39°23′28″ N, 23°4′35″ E,” 5 Jul 2013, Dunkel 30124 in Hieracia Europ. Sel. #802 (holotype: B; other isotypes: FI, FR, H, IBF, LI, M, UPA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45261—Italy, “Abruzzen, Prov. Téramo, Monti della Laga: Ceppo, 1340 m s.m., Buchen-Bergahorn-Wald. MTB 3346/2, 42°40′ N, 13°27′ E,” 26 Jul 2002, Gottschlich 46119 in Hieracia Europ. Sel. #232 (holotype: FI; other isotypes: B, FI, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 125287—Italy, “Friaul-Julisch Venetien, Prov. Udine, Ragogna, Monte di Ragogna: “tornante panoramico,” 358 m s.m., schattige Straßenböschung mit Salix-Gebüsch. MTB 9843/2, 46°11′13″ N, 12°57′58″ E,” 27 Jul 2012, Gottschlich 59008 in Hieracia Europ. Sel. #704 (holotype: FI; other isotypes: B, FR, H, IBF, LI, M, PRA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 129013—Greece, “Ostmakedonien, Nom. Xanthi, Lykodromio, Ag. Georgios → M. Erymanthos, 1350 m s. m. 41°18′12″ N, 24°43′42″ E,” 24 Jun 2019, Gottschlich 72671 and Dunkel, in Hieracia Europ. Sel. #808 (holotype: B; other isotypes: FI, FR, H, IBF, LI, M, MSTR, UPA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45161—Italy, “Piemont, Prov. Vercelli, Varallo, Gambararo, Nordportal des südlichen Eisenbahntunnels, 440 m s.m, Mauerfugen. MTB 0115/4, 45°48′ N, 8°16 E,” 30 May 2007, Gottschlich 40509, Soldano and Rotti in Hieracia Europ. Sel. #130 (holotype: FI, other isotypes: B, GOET, H, IBF, LI, M, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
  • Hieracium schmidtii subsp. cochleariforme Gottschl. in Webbia 66: 228. 2011 [52604].
Isotype: PAL-Gr 52604—Italy, “Umbrien, Prov. Perúgia, Monti Sibillini, Forca Campanina, Straße Richtung Nórcia, NW Costa Jovine, 1245 m s.m., Straßenrand. MTB3244/2, 42°46′2″ N, 13°9′37″ E,” 28 May 2009, Gottschlich 54606 in Hieracia Europ. Sel. #351 (holotype: FI; other isotypes: B, GOET, H, IBF, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45227—Italy, “Abruzzen, Prov. L’Áquila, L’Áquila, Camarda Richtung Aragno: W Autobahn, beim Wasserwerk, 1010 m s.m., Eichen-Mischwald. MTB 3646/2, 42°23′ N, 13°27′ E,” 29 May 2006, Gottschlich 50994 in Hieracia Europ. Sel. #198 (holotype [to be specified]: FI; other isotypes: B, FI, GOET, IBF, H, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45207—Italy, “Abruzzen, Prov. Chieti, Majella, Abstieg M. Cavallo Richtung Selvaromana, Valle del Infierno,” 1200–1400 m s.m., artenreiche Hochstauden. MTB 3850/4, 42°0′ N, 14°9′ E,” 1Aug 2002, Gottschlich 46300 and Conti in Hieracia Europ. Sel. #178 (holotype [to be specified]: FI; other isotypes: B, FI, GOET, H, IBF, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype:PAL-Gr 52614—Italy, “Piemont, Prov. Torino, Valle di Susa, S unterhalb Colle di Finestre, 1995 m s.m., im Schatten von Lärchen und Wacholderbüschen. MTB: 0908/1, 45°4′44″ N, 7°3′8″ E,” 9 Aug 2010, Gottschlich 55979 in Hieracia Europ. Sel. #361 (holotype: FI; other isotypes: B, GOET, H, IBF, LI, M, PRC, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
  • Hieracium semipallescens Gottschl. in Webbia 66: 210. 2011 [52609].
Isotype: PAL-Gr 52609—Italy, “Marche, Prov. Pésaro-Urbino: Nordseite M. Nerone, 1470 m s.m., Kalkfelsstufen, Kalkschutt. MTB 2441/1, 43°33′30″ N, 12°31′3″ E,” 10 Jul 2010, Gottschlich 55844 in Hieracia Europ. Sel. #356 (holotype: FI; other isotypes: B, GOET, H, IBF, LI, M. PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 121159—Italy, “Toskana, Prov. Lucca, Alpi Apuane, Arni, Le Gobbie, Passo degli Uncini, 1370–1400 m s.m., aufgelichteter Buchenwald. MTB 1927/1, 44°3′8″ N, 10°13′35″ E,” 1 Jul 2015, Gottschlich-64318 in Hieracia Europ. Sel. #615 (holotype: FI; other isotypes: B, FR, H, IBF, LI, M, PRA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 45130—Austria, “Kämten, Karnische Alpen, Stranig: Fahrweg zur Staniger Alm oberhalb des Stranig-Baches 50 m unterhalb der Gedenktafel für Georg Hohenwarter, 1350 m s.m., durch Wegebau angerissene Steilböschung (paläozoische Schiefer) mit Grünerlen, darüber Buchen-Fichten-Wald. MTB 9344/4, 46°36′ N, 13°8′ E,” 11 Aug 1993, Gottschlich 22471 in Hieracia Europ. Sel. #99 (holotype: LI; other isotypes: KL, M, hb. Gottschl.).
Isotype: PAL-Gr 45256—Italy, “Abruzzen, Prov. L’Áquila, Roccaraso, an der Straße zum Wintersportgebiet “Aremogna,” oberhalb der großen Hotels, 1530 m s.m., Kalkschutthang. MTB 4150/1, 41°50′ N, 14°3′ E,” 1 Jun 2001, Gottschlich 43903 in Hieracia Europ. Sel. #227 (holotype [to be specified]: FI; other isotypes: B, FI, GOET, H, IBF, LI, M, PR, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 129008—Greece, “Zentralmakedonien, Nom. Thessaloniki, Mount Chortiatis: Kisson, nahe dem Sperrgebiet der Sendemastanlage, 934 m s.m., Eichenwald. 40°35′51″ N, 23°6′18″E,” 11 Jul 2018, Gottschlich 70162 in Hieracia Europ. Sel. #803 (holotype: B; other isotypes: FI, FR, H, IBF, LI, M, MSTR, UPA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Lectotype [designated here]: PAL-Gr 66707—Montenegro, “in einer Doline bei Njegus. Die zahlreichen Drüssen von stupposum, Habitus undBlattform von racemosum, Behaarung und kleine Köpfe von stepposum. Zahn,” Jul 1904, Sagorski (isolectotypes: none known).
  • Hieracium torrigliense Gottschl. in Webbia 64: 184. 2009 [49584].
Isotype: PAL-Gr 49584—Italy, “Ligurien, Prov Génova, Propata → Capp.la Tre Croci, 1195 m s.m., grasige Straßenböschung unter Buchenwald. MTB 1421/3, 44°34′32″ N, 9°11′45″ E,” 27 Jun 2008, Gottschlich 53414 in Hieracia Europ. Sel. #301 (holotype: FI; other isotypes: B, GOET, H, IBF, LI, M, PRA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 125289—Italy, “Friaul-Julisch Venetien, Prov. Udine, Paluzza, Sutrio, Fahrweg zum Hotel “Alle Trote,” 521 m ü.m., Wegrand und Gebüsch unter Kiefern-Fichten-Wäldchen. MTB 9443/4, 46°30′5″ N, 12°59′59″ E,” 27 Jul 2015, Gottschlich 64604 in Hieracia Europ. Sel. #705 (holotype: FI; other isotypes: B, FR, H, IBF, LI, M, PRA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl.).
Isotype: PAL-Gr 129009—Greece, “Zentralmakedonien, Nom. Iamthia, M. Vermio: Sraße Richtung Kato Vermio, 1390 m s.m., Kalkfelsböschung. 40°33′57″ N, 22°1′36″ E,” 17 Jun 2019, Gottschlich 72322 and Dunkel in Hieracia Europ. Sel. #804 (holotype: B #101042256; other isotypes: FI, FR, H, IBF, LI, M, MSTR, UPA, W, hb. Brand, Hb. Gottschl.).
Isotype: PAL-Gr 45131—Austria, “Tirol, Verwallgruppe, Galtür, Kopsstraße bei Überquerng durch denSchilift, 1700 m s.m., Schipisten mit Erlengebüschen. MTB 9026/2, 46°58′ N, 10°8′ E,” 22 Aug 1999, Gottschlich 38955 in Hieracia Europ. Sel. #100 (holotype: B; other isotypes: B, H, IBF, LI, M. PRA, W, hb. Brandstätt., hb. Dunkel, hb. Gottschl., etc.).
  • Holosteum breistrofferi Greuter and Charpin in Exsicc. Genav. Conserv. Bot. Distrib. Fasc. 2: 23. 1971 [=Holosteum umbellatum subsp. hirsutum (Mutel) Breistr. 1970] [6635].
Isotype: PAL-Gr 6635—France, “prope jugum ‘pas de l’Echelle’ ad septemtrionem pagi ‘Saint-Geniez’ provinciae Alpium inferiorum, 1200 supra maris aequor. In petrosis calcareis ad pedes rupium praeruptarum ad meridiem spectantium, uno in loco abundans in consortio Veronicae hederifoliae subsp. trilobae, Anthrisci caucalidis, etc., 12 May 1970, Charpin, Greuter and Monthoux in Exsicc. Genav. #57 (holotype: G; other isotypes: GB, GZU, LD, NO, OSC, RSA, SP).
Isotypes: PAL-Gr 134863 and 134864—Greece, Crete, “Ep. Milopotamos, Idhi Oros, Doline N des Berges Piperos an der Straße zwischen Kirche Ajios Fanourios uns Berg Skinakas (35°12′30″ N; 24°52′30″ E); lehmig-kiesiger Dolinengrund und Felswand, ca 1450 m; 7 Sep 1895, Risse 1938 (holotype: B; other isotypes: none).
Isotype: PAL-Gr 39666—Australia, [“between Hutt and Murchison R, W. Australia”] (from holotype label)], [Drummond VI] 115 (holotype [specified here]: TCD #9741; other isotypes: K [2×], MEL [2×]).
Isotype: PAL-Gr 44388—Cuba, “Prov. Matanzas: Municipio Martí, Ciénaga del Majaguillar al NO de Martí, Ciénaga de Gonzalito cerca del Canal de Blanquizal, alt. 5 m, 22°58′39″ N, 80°58′06″ W. Herbazales de ciénaga,” 28 Feb 2009, Greuter 27013 & al. (holotype: HAJB #505; other isotypes: B, JE, NY).
  • Hypericum hircinum var. albimontanum Greuter, Colloque OPTIMA Crète Guide Excurs.: 25. 1975 (≡Hypericum hircinum subsp. albimontanum (Greuter) N. Robson) [13837].
Holotype: PAL-Gr 13837—Greece, Crete, “Ep. Sfakia: b. Aj. Rùmeli, 100 m ü.M. Schattige Grasborde längs Wassergräben, unter Gebüsch stellenw. hfg.; abweichend durch breitere, am Grund herzförmige, am Rand knorpelig gezähnelte Blätter,” 10 Jul 1961, Greuter 3837 (isotypes: G, W, Z).
  • Hypericum jovis Greuter, Colloque OPTIMA Crète Guide Excurs.: 25. 1975 [14571].
Holotype: PAL-Gr 14571—Greece, Crete, “Ep. Monofatsi: N-Hang des Berges Kòfinas, 1100 m.ü.M., Kalk. Hfg. in schattigen Ritzen der N-exp. Felswände; die erheblich abweichende mittelkretische Felsrasse, durch vermutlich hybride Zwischenformen mit dem Typus verbunden,” 27 May 1962, Greuter 4571 (isotypes: G, LJU, M, UPA, W, Z).
Holotype: PAL-Gr 126167—Greece, “Nomos Kiklades, Paros, Stroumboulas SSW von Lefkes, S-Seite, SE-exponierter Hang, 695 m; (37°2′6″ N, 25°11′26″ E). Kalk, Steinfluren in der Pistacia lentiscus-Sarcopoterium-Phrygana. 4 Jun 2017, Karl (isotypes: B, C, LD, M, MSB, W).
Isolectotype: PAL-Gr 66357—Bosnia and Herzegovina, “auf Steingeröll neben der Eisenbahn unterhalb des Hmubergs, bei Mostar, ca. 75 m, Jul 1901, Sagorski (lectotype [designated here]: JE # 6943; other isolectotypes: B, HBG [3×], JE #6944].
Isotype: PAL-Gr 66296—Portugal, “Alentejo litoral. S. Tiago do Cacém in arenaceis maritimis promontorii Cabo de Sines, 50 m s.m., 8 Sep 1938, Rothmaler 14196 (holotype [specified here]: JE #2232; other isotypes: B, JE #2233, S).
Isotype: PAL-Gr 66721—Italy, “Basilicata, Pollino National Park, Mt Alpi (Latronico, Potenza), 40°7′9.65″ N, 15°59′16.95″ E, carbonate stony slopes. Alt. 1750 m,” 11 Jul 2014, Scafidi and Di Gristina (holotype: PAL #102699; other isotype: FI).
Holotype: PAL-Gr 49913—Greece, “Nom. Arkadia, Ep. Mantinia: 6 km NNE of Tripolis, Tripolis-Pirgos road, by the branching of the road to Nestani, alt. 650 m, 37°33′30″ N, 22°24′10″ E. Cultivated fields, wetland patches, roadsides,” 31 May 1995, Kamari & al. in OPTIMA Iter Medit. 7 #322 (Isotypes: B, BEO, BRNM, MA, SALA, UPA, W).
Isotype: PAL-Gr 39677—Australia, [Western Australia, “Swan River, 1854” (from holotype label)], [Drummond VI] 11 (holotype [specified here]: G #388717; other isotypes: BM, G #388716, K, LD, MEL, NSW, NY, P).
Isotype: PAL-Gr 39678—Australia, [Western Australia, “Swan River, 1854”] (from lectotype label), [Drummond VI] 15 (holotype [specified here]: G #388691; other isotypes: BM, G #388690, E, K, MEL, NSW, NY, P, PERTH, TCD).
Isotype:PAL-Gr 39676—Australia, [Western Australia, “Swan River, 1854”] (from holotype label), [Drummond VI] 12 (holotype [specified here]: G #388683; other isotypes: BM, G #388682, MEL [2×], NSW, P, PERTH).
Isotype: PAL-Gr 39675—Australia, [Western Australia, “Swan River, 1854”] (from holotype label)], [Drummond VI] 13 (holotype [specified here]: G #388685; other isotypes: BM, G #388684, K, MEL, NY, P [2×], TCD).
Holotype: PAL-Gr 51901—Greece, “Nom. Lakonia, Ep. Epidhavros Limiras: Mt. Korakia N of the village Richea, alt. 500–700 m, 36°51′35″ N, 23°0′0″ E, 8 Jun 1995, Iatrou & al. In OPTIMA Iter Medit. 7, #1901 (isotypes: B, BRNM, SALA, UPA).
Holotype: PAL-Gr 28414—Greece, “Macedonia occidentalis (UTM: DK-3; Distr. No. 340), a pago Eptahori 2 km occidentem versus, alt. 700–750 m. In clivis sterilibus detritu schistoso margaceo obtectis,” 24 Jul 1977, Greuter15152, Charpin, Bernardi & al. (isotypes: ATH, B, BM, C, E, ERE, G, M, TAD, UPA).
Isotype: PAL-Gr 126211—Cuba, “Prov. Guantánamo. Mun. San Antonio del Sur, Baitiquirí, camino a la Mina de Yeso, montes secos, caliza, 200–400 m,” 13 May 1980, Álvarez de Zayas & al. In HFC 43147 (holotype [specified here]: HAJB #1461; other isotypes: B, HAJB #1460, JE [2×]).
Isotype: PAL-Gr 129315—Italy, Sardinia, “südöstlich von Cágliari, am Rand der Staubstraße zum Capo S. Elia, auf Kalksteinfelsen und sandigen Flächen, 15–25 m; 39°11′5″ N, 9°8′45″ E,” 16 Nov 2000, Erben E1145 (holotype: MSB; other isotype: FI).
  • Limonium cephalonicum R. Artelari, Biosust. Melet. Gen. Limonium: 57. 1984 [38814].
Isotype: PAL-Gr 38814—Greece, “Ins. Kephallinia: ad castrum pagi Assos; in saxosis calcareis maritimis,” 9 Oct 1982, Artelari 376 (holotype: UPA; other isotypes: B [2×]).
Isotype: PAL-Gr 129640—Spain, Balearic Islands: “Mallorca: Alcudia, nordöstlich von Alcudia, Felsküste bei El Mal Pas, 2–8 m (2n = 27),” 10 Oct 1989, Erben E766, Morales Valverde and Rosselló (holotype: M; other isotype: MA).
  • Limonium coronense R. Artelari, Biosust. Melet. Gen. Limonium: 71. 1984 [38812].
Isotype: PAL-Gr 38812—Greece, “Prov. Messinia: prope Castrum oppidi Coroni, in saxosis calcareis maritimis,” 28 Aug 1982, Artelari 356 (holotype: UPA; other isotypes: none known).
  • Limonium himariense F. K. Mey. in Haussknechtia Beih., 15: 113. 2011 [52733].
Isotype: PAL-Gr 52733—Albania, “Himara, Dhermi, Strand, an Felsen im flachen Meer,” 7 Nov 1961, Meyer 5964 (holotpe [specified here]: JE #16701; other isotype: JE #16702).
Isotype: PAL-Gr 129193—Italy, “Prov. Savona: Ventimiglia, Punta delle Roccia, lehmige Felshänge am Meer,” 27 Sep 1977, Erben E229 (holotype: M; other isotype: G).
Isotype: PAL-Gr 129736—Italy, Sardinia, “Prov. Sassari. Ca. 1 km westlich von Porto Cervo, feucht-sandige Flächen an der Mündung eines kleinen Flusses ins Meer,” 25 Aug 1973, Erben E67 (holotype: MSB; other isotypes: FI, M).
Isotype: PAL-Gr 129712—Greece, “Karpathos, Damatria,” 30 Jun 2002, Brullo and Giusso (MSB; other isotypes: CAT, FI).
Isotype: PAL-Gr 38815—Greece, “Ins. Zakynthos: ad pomontorium Skinari, prope pagum Korithi: in saxosis calcareis maritimis,” 19 Feb 1982, Phitos and Kamari 18978 (holotype: UPA; other isotype: B).
  • Limonium saracinatum R. Artelari, Biosust. Melet. Gen. Limonium: 42. 1984 [38819].
Isotype: PAL-Gr 38819—Greece, “Ins. Kephallinia: inter promontorium Sarakinato et pagum Poros, in saxosis calcareis maritimis,” 27 Aug 1976, Artelari 93 (holotype: UPA; other isotypes: none traced).
Holotype: PAL-Gr 53056—Greece, “Nom. Messinia, Ep. Kalamata: Taijetos Pass between Tripi and Artemisio, alt. 1200–1350 m, 37°4′0″ N, 22°16′0″ E, 10 Jun 1995, Kamari & al. In OPTIMA Iter Medit. 7: #2056 (isotypes: B, UPA).
Isotype: PAL-Gr 66705—Greece, “Attica: Laurion,” 1885, Haussknecht (holotype [specified here]: JE #5696–5697 [2 sheets]; other isotypes: JE #6132–6133 [2 sheets], LD).
Isolectotype: PAL-Gr 66696—Greece, “Laurion,” 1885, Haussknecht (lectotype [designated here]: JE #7424¸other isolectotypes: JE ##7420, 7421, 7422, 7423).
Isotype: PAL-Gr 66698—Greece, “Korinth; am Meeresufer,” 28 Apr 1885, Haussknecht (holotype [specified here]: JE #1663¸other isotypes: JE ##1661–1662, PI).
  • Maytenus revoluta subsp. bissei Mory in Feddes Repert. 96: 549. 1985 [48553].
Isotypes: PAL-Gr 48553 and 128076—Cuba, “Prov. Holguín, Mun. Moa. La Veguita, monte la Breña, alrededores del campamento Los Carboneros, pluviosilva destruída, cerca 400–500 m s.m,” 14 Apr 1981, Bisse & al. in HFC 44129 (holotype [specified here]: HAJB #676; other isotypes: B [2×], HAJB, JE, NY).
Holotype: PAL-Gr 15346—Greece, “Dodecanesus, ins. Kárpathos: in latere austro-orientali et ad cacumen montis Kollas, 750 et 900 m supra maris libram. In herbidis inter saxa, solo calcareo, perrara (duo specimina tantum detexi). 17 May 1963, Greuter 5346 (isotypes: none).
  • Medicago strasseri Greuter, Matthäs, and Risse in Willdenowia 12: 201. 1982 (≡Medicago arborea subsp. strasseri (Greuter, Matthäs, and Risse) Sobr.-Vesp. and Ceresuela, 2000) [https://www.jstor.org/stable/3995928 (accessed on 6 March 2024)].
Isotype: PAL-Gr 34773—Greece, Crete, “Eparchie Rethimni: Petres-Schlucht N von Karoti, 30°20′30″ N, 24°21‘30″ E. Kalkfelswände, 30 m. 26 May 1982, Greuter 19632, Matthäs and Risse (holotype: B; other isotypes: LD).
  • Merendera greuteri Gabrieljan in Fl. Rastitel’nost’ Rast. Res. Arm. 12: 15. 1999 (≡Colchicum greuteri (Gabrieljan) K. Perss. 2007) [38798].
Isotype: PAL-Gr 38798—Armenia, “Shirak distr., in vicinitate pag. Areg, m. Arteni in steppis tragacanthaceis, 1500–1700 m s.m.,” 9 Apr 1998, Gabrielian in ERE #147519 (holotype: ERE #661; other isotypes: B, ERE #662, 663).
Isotypes (2): PAL-Gr 45665 and 38805—Greece, “Nomos Evros, Eparhia Soufli, 1 km S of Dhadhia, 100 m, 41°7′20″ N, 26°13′ E. Grassland on micaschist hill,” 13 Jun 1992, Greuter 23299 (holotype: UPA; other isotypes B [3×], C, G, LD).
Isotype: PAL-Gr 32305—Greece, “Thraki, eparhia Soufli, montes Boukate dag, a pago derelicto Pessani 6 km septemtriones versus, in colle ad occidentem stationis televisoriae, alt. 400 m. In cacumine rupestri granitico e sylva Quercus frainetto proeminente. In fissuris,” 15 Jul 1978, Greuter 15988 (holotype: UPA; other isotypes B, C).
Holotype:PAL-Gr 51721—Greece, “Nom. Lakonia, Ep. Epidhavros Limiras: Between Metamorfosi and Richea, summit area of Mt. Koulochera, alt. 800–950 m, 36°49′40″ N, 22°58′50″ E, 6 Jun 1995, Kamari & al. In OPTIMA Iter Medit. 7: #1721 (isotypes: B, BRNM, MA, SALA, UPA).
Isotype (by conservation): PAL-Gr 62559—USA, “Grown in S. Renner’s garden in St. Louis from the seed of watermelon ‘Crimson Sweet’ bought in a store in St. Louis,” Renner 2816 (holotype: M; other isotypes: B, BM, K, L, LE, MO, P).
Holotype: PAL-Gr 14855—Greece, Crete “Ep. Sfakia: b. der Quelle Dzaràni am W-Abfall des Berges Dzaranokefàla, 1850 m.ü.M., Kalk. Schattige Höhlung der Felswand mit lehmiger Komposterde (Ziegenläger), lokal reichl.; durch grössere Zartheit u. schwächere Behaarung aller Teile, längere, weniger gekrümmte Fr-Stiele abweichend,” 4 Jul 1962, Greuter 4855 (isotypes: G, LD, M, W, Z).
Holotype: PAL-Gr 1844—Greece, Crete, “Lefka Ori N., Hagios Pneuma, Falaise calcaire à 1500 m, exposition NE,” 20 Jul 1964, Zaffran 5907 (isotype: hb. Zaffran).
Isotype: PAL-Gr 129938—Greece, Cuba, “Prov. Holguín, Mpio. Mayarí: charrascales de La Caridad cerca del río Naranjo. Alt. 300–450 m s.m., 20.474592° N, 75.740322° W,” 13 Jun 2018, Acosta & Gómez in HFC 42165 (holotype: HAJB #1255; other isotypes: B, FTG, HAC, HAJB #1254, HAJU, herb. Jard. Bot. Holguín).
Isotype: PAL-Gr 58192—Greece, Cuba, “Prov. Sancti Spíritus. Trinidad. Topes de Collantes. Mogote mi Retiro,” Olmstead & al. in HFC 72727 (holotype [specimen shown in Willdenowia 33: 177, in Fig. 3]: HAJB #711; other isotypes: B, HAJB ##710, 712 [2×], JE).
  • Narcissus tortifolius Fern. Casas in Saussurea 8: 43. 1977 [26657].
Isotype: PAL-Gr 26657—Spain, “Almería, Sorbas: Venta de los Castaños, 2oS WG 8511, ad 400 m, in gypsaceis siccis,” 23 Mar 1977, Fernández Casas 1484 and Pueche (holotype [specified here]: MA #227840; other isotypes: BCN, C, MA #208486.
Holotype: PAL-Gr 53511—Greece, “Nom. Messinia, Ep. Kalamata: 6–8 km NE of Ano Amfia along the road to Poliana, alt. 600–800 m. 37°8′20″ N, 22°7′30″ E. Dry river bed and rocky limestone slopes with maquis,” 14 Jun 1995, Kamari & al. In OPTIMA Iter Medit. 7: #2511 (isotypes: B, BEO, BRNM, MA, SALA, UPA, W, etc.).
  • Nigella carpatha Strid in Opera Bot. 28: 49. 1970 [15134].
Holotype: PAL-Gr 15134—Greece, “Regio aegaea austr., Insula Karpathos: ad boreo-occidentem porti Finiki versus promontorium, alt. 8–15 m. In petroso-argillosis plani sublitoralis. Ad pedes muriculi, uno in loco copiose,” 11 May 1963, Greuter 5134 (isotypes: G, LD, HUJ, M, W, Z).
Holotype: PAL-Gr 49916—Greece, “Nom. Arkadia, Ep. Mantinia: 6 km NNE of Tripolis, Tripolis-Pirgos road, by the branching of the road to Nestani, alt. 650 m, 37°33′30″ N, 22°24′10″ E,” 31 May 1995, Kamari & al. in OPTIMA Iter Medit. 7 #325 (isotypes: B, UPA, SALA, BRNM, MA, BEO, W, etc.).
  • Onobrychis aragatzi Arevsch. in Novosti Sist. Vysš. Rast. 41: 93. 2010 [48158].
Isotype: PAL-Gr 48158—Armenia, “Aragatz mountain, South macroslope, meadow-steppe, varied grasses, 1800 m a.s.l.,” 19 Jul 2007, Arevschatian 2009 (holotype [specified here]: ERE #531; other isotypes: ERE ##532, 533, LE).
Holotype: PAL-Gr 14638—Greece, Crete, “Ep. Sfakia: S-Abfall des Grates swl. des Xilòskala-Passes, 1400–1500 m.ü.M., Plattenkalk. Verbr. u. hfg. in Ritzen sonniger Felswände, in grossen Büschen; schöne neue Art aus der sect. Eubrychis subsect. Albae ser. Verae,” 2 Jun 1962, Greuter 4638 (isotypes: ATH, G, HUJ, LD, LJU, M, UPA, W, Z, hb. Creutzburg, Hb. Zaffran).
  • Onosma albanica (‘albanicum’) Dörfl. and Ronniger in Anz. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 55: 283. 1918 (≡Onosma pseudoarenaria subsp. albanica (Dörfl. and Ronniger) Rauschert 1976) [=Onosma pseudoarenaria subsp. fallax (Borbás) Rauschert 1976 ≡ Onosma helvetica subsp. fallax (Borbás) Teppner 1971] [https://www.zobodat.at/pdf/AAWW_55_0001-0404.pdf (accessed on 6 March 2024)].
Isotype: PAL-Gr 5314—Albania, “Distr. Luma. Zwischen Eichengebüsch auf Weiden am Wege von Kula Luma nach Göstil,” [25 Jun 1916/1918, Dörfler 687] (Holotype (specified by Vogt & al. in Willdenowie 48: 78. 2018): B #9006177; other isotypes: WU, Z).
  • Onosma psammophila (‘psammophilum’) Rech. f. and Riedl in Bot. Not. 124: 79. 1971 [30626].
Isotype: PAL-Gr 30626—Greece, “Macedonia orientalis: in arenosis maritimis ad sinum Orphani ab ostiis fluvii Strymon c. 10 km occidentem versus,” 21 Jul 1970, Rechinger 39000 (holotype: W; other isotypes: B, G).
Isolectotypes (2): PAL-Gr 6445 and 8516—Greece, “bei Argos, Tiryns-Ruinen,” 17 Apr 1911, Müllner (lectotype [designated here]: WU [Halácsy, Greek hb.]; other isolectotypes: GB?, WU?).
  • Ophrys candica Greuter, Matthäs and Risse in Willdenowia 15: 53. 1985 [=Ophrys holosericea subsp. candica (E. Nelson ex Soó) Renz and Taubenheim [https://www.jstor.org/stable/3996541?seq=31 (accessed on 6 March 2024)].
Holotype: PAL-Gr 12518—Greece, Crete, “Ep. Kidonia: b. Dorf Prasses, 500 m.ü.M. Raine und Wegborde der Kulturzone,” 24 Apr 1960, Greuter 2518 (isotypes: none).
Holotype: PAL-Gr 7093—Greece, “Attika, Mt. Parnis. Parasitisch auf Pterocephalus perennis,” end May 1974, Baumann and Baumann (isotype: B).
  • Papaver gabrielianae M. V. Agab. and Fragman in Fl. Rastitel’nost’ Rastitel’nye Resursy Armenii 16: 27. 2007 [44990].
Isotype: PAL-Gr 44990—Armenia, “Sjunik, m. Mets Ishkhanasar, road from lake Sevlich, 2680 m, 39°49′ N, 44°45′ E,” 6.Jul 2006, Aghababian, Gabrielian, Nersessian, and Sarkissian (holotype: [specified here]: ERE #635; other isotypes: ERE #636–638 [3×]).
Isotype: PAL-Gr 62627—Armenia, “Ararat distr., Gorovan, sandy soils, 39°53′40.31″ N, 44°43′59.89″ E, 940 m,” 21 May 2010, Gabrielian and Aghababyan (holotype[specified here]: ERE #639; other isotypes: B, ERE ##640–642 [3×]).
Isotype: PAL-Gr 52145—Cyprus, “Division 2 sensu Meikle 1977/1985, Kykko, Vrysi tou Klamenou (‘Klamenon’), road towards Kykko monastery, screes, alt. 900 m,” 14 Apr 2010, Christodoulou (holotype [specified here]: B #100413396; other isotypes: B #100413395, CYP, ERE).
  • Papaver roseolum M. V. Agab. And Fragman in Fl. Rastitel’nost’ Rastitel’nye Resursy Armenii 16: 27. 2007 [44991].
Isotype: PAL-Gr 44991—Armenia, “Avan, dry stony slopes,” 25 May 2006, Gambarian (holotype [specified here]: ERE #597; other isotypes: ERE ##598–602 [5×], HUJ).
  • Parietaria cardiostegia Greuter in Fl. Australia 3: 190. 1989 [33354].
Isotype: PAL-Gr 33354—Australia, New South Wales, “‘Mesa’ hills c. 12 km NW of Fowlers Gap beyond NW domain border, alt. 250–300 m, 31°1′ S, 141°39′ E. Hillsides in open semi-desert area,” 4 Sep 1981, Greuter 18387 (holotype: B; other isotypes: K, NSW).
Isotype: PAL-Gr 23946—Greece, “Laconia: Peninsua Malea, in arenosis N Neapolis,” 8 May 1964, Rechinger 24505 (holotype: W; another isotype: B, S).
Isotype: PAL-Gr 30640—Greece, “Insula Kythera: Kapsali, substr. Calc., 50–200 m,” 5 May 1964, Rechinger 24363 (‘24505’) (holotype: W #1981–10835; other isotypes: B, E, GZU, MA, MO, S, WU).
Holotype (2 sheets): PAL-Gr 17650/1 and 2—Greece, Crete: “Ep. Sitia: prope Vái, 5 m. In plano arenoso vel argilloso juxta mare lucum formans. Arbor dioica, truncis pluribus fasciculato-aggregatis. Baccae cum axibus fructiferis vivide miniatae,” 2 Oct 1966, Greuter 7650 (isotypes: ATH, B, E, G, GB, K, LD, M, UPA, W, hb. Zaffran).
Isotype: PAL-Gr 126168—Cuba: “Prov. Holguín. Rafael Freyre: Loma El Templo, al oeste de la Bahía Naranjo. Bosque semideciduo microfilo. 70–80 m s.m.,” 23 May 2017, González & al. In UCLV 12355 (holotype: ULV [specimen shown in Willdenowia 49: 188, Fig. 1]; other isotypes: B, HAC, HAJB, ULV).
Isotype: PAL-Gr 5298—Greece, Crete: “Distr. Sphakia. Insel Gaudos, Im Dünensande der Nordküste. (Neu für Europa!),” 20 Mar 1904, Dörfler in Iter Cret. #1148 (holotype [specified here: WU #99192; other isotypes: B).
Holotype [specified here]: PAL-Gr 16568—Greece, “Aegäis, Nom. Attikì, Ep. Kìthira: Hügel südl. Kàlamas, 100 m.ü.M., Neogen. Mehrfach unter Sklerophyllen-Gebüsch,” 12 May 1964, Greuter 6568 (isotypes: none).
  • Polygala negevensis Danin in Israel J. Bot. 36: 67. 1987 [123562].
Isotype: PAL-Gr 123562—Israel: “S. Negev: 22 km NNW of Elat, crevices of hard limestone in a wadi,” 1 Jul 1986, Danin and Chouate (holotype: HUJ; other isotypes: B, G, K, MSB, P, W [2×]).
Isotype: PAL-Gr 6018—Greece, “Karpathos, ad rupes calcareas maritimas litor. Bor. Or. Insulae, prope promont Grea,” 25 May 1886, Major119 (holotype: G #301451); other isotypes: K, LE).
Isotype: PAL-Gr 6131—Turkey, “Pentes schisteuses bordant le Guzel-Déré, en amont de Sédichig, à 4 lieues au NO. De Mersina,” 5 Jun 1855, Balansa in Pl. d’Orient #640 (holotype [specified here]: G #301447 [2 sheets]; other isotypes: B, BM, C, FI-W, G #301448, G-BOIS, GOET, JE, LAU, LE, MANCH, MPU, P, TL, W [2×], WAG).
Isotype (2 sheets): PAL-Gr 44578/1 and 2—Italy, Sicilia “Prov. Trapani, loco classico et unico. In dumetis et ad rupes calcareas clivium asperorum sylvestrium septemtriones spectantium,” 4 Jun 2005, Raimondo, Greuter 26423 and Aghababyan (holotype: PAL; other isotypes: B, FI).
  • Ptilotrichum cyclocarpum subsp. pindicum Hartvig in Strid, Mountain Fl. Greece 1: 305. 1986 (≡Phyllolepidium cyclocarpum subsp. Pindicum (Hartvig) Cecchi) [28326].
Isotype: PAL-Gr 28326—Greece, “Epirus (UTM: DK-4; Distr. No. 514), montes Timfi: In latere orientali montis Astraka, alt. 2000–2100 m. In clivis asperis herbosis petroso-glareosis calcareis,” 23 Jul 1977, Greuter 15066 & al. (holotype: C #10008875; other isotypes: ATH, G).
Isotype: PAL-Gr 39660—Australia, [Western Australia, “Swan River”] (from holotype label), [Drummond VI] 220, [1854]” (holotype [specified here]: KW #1001667; other isotypes: A, BM, CGE, E, G, K, LD, MEL, W).
Isotype: PAL-Gr 35216—Greece, Crete, “Prov. Hania. Distr. Sfakion. Lefka Ori, Mt. Svouritchi [Svourihti], summit area, c 2 km NE of the summit of Mt. Pachnes. Alt: 2100–2300 m. N-NE facing slopes with limestone cliffs or E facing scree slopes. Petals yellow,” 12 Jul 1980, Baden and Franzen 537 (holotype: C; other isotypes: none).
Isotype: PAL-Gr 129151—Cyprus, “Division 7 (sensu Miekle 1985), Ypsarovounos forest, c. 2.5 km southwest of Mandres Ammochostou, on somewhat vertical or steep, almost bare and soft gypsum faces of few square metres, with flattish patches on or at their base, alt. c. 280 m, UTM 36 N 572184 E, 3909366 N,” 5 Apr 2018, Hadjikyriakou 7600 (CYP; other isotypes: ARI, B, G, STU, hb. Hadjikyr.).
Holotype [specified here]: PAL-Gr 66351 p.p., fertile branches only—France, “Arnas (Rhône),” 11 Apr 1874 (isotype: LY).
  • Salsola carpatha P. H. Davis in Notes Roy. Bot. Gard. Edinburgh 21: 139. 1953 (≡Caroxylon carpathum (P. H. Davis) Akhani and Roalson 2007) [1108].
Isotype (holotype fragment): PAL-Gr 1108—Greece, “Karpathos, Vurgunda (N.W. of Olymbos), 5–20 m. Calc. sea rocks with Galium canum,” 24 Jul 1950, Davis 18025 (holotype [specified here]: K #899553; other isotypes: B, E, K #899552, S, W).
Isolectotype: PAL-Gr 66367—Greece, “Pindus Tymphaeus: in summo montis Zygos (Lakmon veter.) supra Metzovo, alt 4500–5000′, substratu silico-serpentino,” Jul 1885, Haussknecht (lectotype [designated here]: JE #16780; other isolectotypes: JE #16781, K).
Isotype: PAL-Gr 38806—Greece, “Nomos Lakonias, Ep. Epidavrou-Limiras, Ins. Elafonisos: in arenosis littoreis ad occidentem insulae spectantibus. 36°29′ N, 22°56′ E, 24 Mar 1993, Phitos, Kamari, and Christodoulakis 23209 (holotype: UPA; other isotype: B).
Isotype: PAL-Gr 66359—Spain, “Sierra de Loxa,” Jun [1849], del Campo in Hohenacker, Pl. Hispan. #41 (lectotype [Webb ms. in schedis, designated here]: G #388986; other isolectotypes: BM; G [2×], JE ([6×], K [2×], MICH, WAG).
Holotype: PAL-Gr 17713—Greece, Crete, “Ep. Sfakia: supra hiatum Kakískala propr fontrm Linoseli, 1450 m. Ad rupes praeruptas calcareas sole illustres frequens,” 11 Oct 1966, Greuter 7713 (isotypes: ATH, B, E, G #388987, GB, K, LD, M, UPA, W).
Holotype: PAL-Gr 14574 –Greece, Crete, “Ep. Monofatsi: Gipfel des Berges Kòfinas, 1200 m.ü.M., Kalk. S-exp. Felswände in grossen polsterförmigen Büschen, 27 May 1962, Greuter 4574 (isotypes: G, HUJ, LD, W, Z).
Isotype: PAL-Gr 122002—Cuba: “Provincia de Holguín. Cerro Verde, km 10, carretera Holguín-Gibara. Colinas al este de la carretera. 80°57′80″ N, 76°15′93″ W. 180 m s.n.m. Matorral xeromorfo espinoso sobre serpentina degradado. Forma parte del estrato herbáceo que rodea los arbustos,” 21 Oct 2016, Méndez Santos and Hernández Peña in HPC 12029 (holotype: HIPC [specimen shown in Rodriguésia 70(e02982017): 3, Fig. 1]; other isotypes: B, HAC, HAJB, JE, ULV).
Isotype: PAL-Gr 14720—Greece, Crete, “Ep. Lassithi: Eingang der diktäischen Zeusgrotte ob Psichrò, 1000 m.ü.M., Kalk. Häufig an feucht-schattigen, bemoosten Felsen,” 13 Jun 1962, Greuter 4720 (holotype: LD; other isotypes: G, W, Z).
Isotype: PAL-Gr 121142—Greece, “Nomos Dhodhekanisos, Eparhia Rodhos: Chalki, Klisoures, crevices of vertical N-facing limestone cliffs at Klisoura, 200–400 m, 36°13′58.3″ N, 27°32′34.28″ E,” 2 Aug 2016, Cattaneo CK35 (holotype C; other isotypes ATH, herb. Cattaneo).
Holotype: PAL-Gr 25109—Greece, “Macedonia occ. (distr. Almopia/Peonia): montes Paiko (Meniki), in colle Pirgos ad sept. Pagi Livadhia, alt. 1400 m. In pascuis petrosis calcareis zonae silvticae Fagi,” 29 Jul 1976, Greuter 13993 (isotypes: G, LJU).
Isotype: PAL-Gr 35207—Greece, “(Phthiotis): in collibus denudatis an fontem 4–6 km a Domokos meridiem versus, ca. 550 m, substr. serpent.,” 6 May 1961, Rechinger 22877 (holotype: B; other isotype: G).
Holotype: PAL-Gr 15928/1 and 2 (2 sheets):—Greece, “Sterea, prov. Viotia, distr. Levadhia: supra pagum Arahova secus viam ad Kalivia ducentem, alt. 1050–2000 m. In rupestribus apricis clivorum asperorum meridiem spectantium,” 14 Jun 1963, Greuter 5928 (isotypes: B, C, LD, UPA).
  • Silene greuteri Phitos in Bot. Chron. 2: 53. 1983 (≡Silene integrifolia subsp. greuteri (Phitos) Akeroyd in Bot. J. Linn. Soc. 97: 341. 1988 [35227].
Isotype: PAL-Gr 35227—Greece, Crete, “prov. Rethimno: prope pagum Mariou in faucibus Kourtaliotiko, alt. c. 350 m, in glareosis,” 13 Apr 1982, Tsanoudakis 7067 (holotype: UPA; other isotypes: none traced).
  • Silene intonsa Melzh. and Greuter in Willdenowia 12: 29. 1982 (≡Heliosperma intonsum (Greuter and Melzh.) Niketić and Stevan. 2007) [https://www.jstor.org/stable/3996065 (accessed on 6 March 2024)].
Isotype: PAL-Gr 29746—Greece, “Ipiros: prov. Ioannina, distr. Konitsa: Konitsa. Valley of the river Aoos. On both sides of the river and E. of the bridge. Alt. 460–500 m. Stony, rocky slopes with scattered shrubs. In clefts of moist rocks. Flowers white,” 22 May 1973, Stamatiadou 17079 (holotype: ATH; other isotype: B).
Isotype: PAL-Gr 29638—Greece, “Nom. Messinias, Ep. Kalamon: Mt. Taygetos, S part, NE of Makrovouna, above a place called Ag. Dimitrios, along the valley leading to the summit ridge, 1600–1950 m. Chiefly limestone but some outcrops of schist. Timberline at c. 1900 m formed by Pinus nigra and Abies cephalonica. Rocky places, 1800 m, limest. Petals white above, pinkish beneath,” 30 Jun 1979, Strid and Papanicolaou 15236 (holotype: B; other isotypes: C, G).
Holotype: PAL-Gr 30997—Greece, “Peloponnes, Messenien, Ep. Pilia: Bucht Vodhokoilia zwischen Petrohori und Paleokastro Navarinou. Ölhaine; 0–5 m. Blüten weiß,” 17 Apr 1979, Greuter 17203 and Merxmüller (isotype: B).
Isotype: PAL-Gr 31507—Greece, “Macedonia or., prov. Dhrama: mons Orvilos, in crista meridionali verticis Trisla supra refugium veterum Preseki, alt. 1450 m (41°20′30″ N, 23°38′ E), In clivis graminosis, solo calcareo,” 21 Aug 1978, Greuter 16619 (holotype: B; other isotypes: AAU, C, G, M, UPA).
Isotype: PAL-Gr 31764—Greece, “Macedonia occ., inter prov. Kozani et Grevena: montes Vourinos, in summo vertice Vourinos ejusque crista septemtrionali, alt. 1600–1850 m (40°10′30″ N, 21°40′ E). In fissuris et scansilibus rupium serpentinicarum septemtriones spectamntium,” 26 Jul 1978, Greuter 16349 (holotype: B; other isotypes: AAU, C, G, UPA, hb. Düll).
Holotype: PAL-Gr 20697—Greece, “Epirus, prov. Ioannina, distr. Dhodhoni: montes Timfi, in latere boreo-occidentali montis Ploskos, alt. 1800–1900 m. Ad rupes calcareas umbrosas. Flores albi,” 20 Aug 1974, Charpin 11294, Dittrich, Greuter 12466 and von Auw (isotypes: ATH, G, UPA).
Isotype:PAL-Gr 17514—Greece, Crete, “Ep. Temenos: ad cacumen montis Júktas, 700 m. In praeruptis rupestribus calcareis ad occidentem spectantibus haud raro. Plane suffruticosa, ex axillis foliorum emortuorum fasciculos foliorum juvenilium proferens,” 14 Sep 1966, Greuter 7514 (holotype: G; other isotypes: B, E, LD, W).
Isotype: PAL-Gr 31777—Greece, “Macedonia occ., inter prov. Kozani et Grevena: montes Vourinos, in summo vertice Vourinos ejusque crista septemtrionali, alt. 1600–1850 m (40°10′30″ N, 21°40′ E). In pratis et pascuis rupestribus, solo ophiolithico,” 26 Jul 1978, Greuter 16333 (holotype: B; other isotypes: none).
Isolectotypes (2): PAL-Gr 8365 and 8366—Greece, Crete, “Amalos,” 12 Jun 1884, Reverchon [Pl. Cret. #249 &] in Baenitz, Herb. Europ. [#5239] (lectotype [specified here]: WU #76665; other isotypes: WU #76664, etc.).
  • Solenopsis minuta subsp. annua Greuter, Matthäs, and Risse in Willdenowia 14: 30. 1984 (≡Solenopsis annua (Greuter, Matthäs, and Risse) Hand and Christodoulou 2020) [https://www.jstor.org/stable/3995693?seq=4 (accessed on 6 March 2024)].
Isotype: PAL-Gr 34297—Greece, Crete, “Eparchie Rethimni: Tal des Sfakoriako S der Prasiano-Schlucht, 35°18′30″ N, 24°33′ E. Grasflächen und Pterideten auf Schiefer, 100–120 m,” 24 May 1982, Greuter 19301, Matthäs and Risse (holotype: B; other isotypes: none known).
Isotype: PAL-Gr 39674—Australia, [Western Australia, “among the Eucalypti thickets on the Gardner and Fitzgerald rivers”] (from holotype label), [Drummond VI] 19 (holotype [specified here]: BM #550676; other isotypes: G, LD, K, MEL, NWS, NY, P).
Isolectotype (lectotype fragment): PAL-Gr 6005—[Greece, Crete], no label data, H[erbarium] G[undelsheimer]]; (lectotype [designated here]: M #30481).
Isolectotype: PAL-Gr 5268– Greece, “Insula Denusa,” 10 Jun 1898, Leonis in Dörfler, Fl. Aegaea #172 (lectotype [designated by Brullo and Erben in Phytotaxa 240: 177. 2016]: M #173917; other isolectotypes: B [2×], FI [3×], FR, MPU).
Isotype: PAL-Gr 7758—Serbia, Kosovo, “SW-Serbia (Kosmet): Miruša sub monte Koznik, in declivibus lapidosis glareosis, solo serpent., cca. 550 m s.m.,” 30 May 1968, Mayer (holotype: LJU #63171; other isotype: BM).
Isotype: PAL-Gr 39664—Australia, [Western Australia, “Between Moore and Murchison Rivers” (from holotype label)], [Drummond VI] 120 (holotype [specified here]: MEL #75843; other isotypes: BM, L, NY, PERTH).
  • Tabebuia triorbicularis var. obovata Borhidi in Acta Bot. Hung. 26: 18 1980. [=Tabebuia calcicola Britton] [http://real-j.mtak.hu/3636/ (accessed on 6 March 2024)].
Isotype: PAL-Gr 123178—Cuba, “Prov. Sancti Spíritus. Mun. Trinidad: Trinidad, Loma del Mirador, en la carretera a Topes de Collantes, 600 m,” 3 May 1977, Bisse & al. in HFC 34683 (holotype [specified here]: HAJB #882; other isotypes: B, BP, HAC, HAJB, JE).
  • Taraxacum basalticum Soest in Bull. Soc. Echange Pl. Vasc. Eur. Occid. Bassin Médit. 15: 111. 1974 [9270].
Isotype: PAL-Gr 9270—France, “Saint-Georges-sur-Allier (France, Puy-de-Dôme), alt. 600 m, pelouse sur basalte,” 26 Apr 1973, Billy in Soc. Echange Pl. Vasc. Eur. #7090 (holotype: LG other isotypes: GENT, LD, MA [2×], MSB, P, SEV).
  • Taraxacum caudatuliforme Soest in Proc. Kon. Ned. Akad. Wetensch., Ser. C, Biol. Med. Sci. 69: 467. 1966 [30620].
Isotype: PAL-Gr 30620—Bulgaria, “ad versuras, 13 km E Plovdiv (Philippopel),” 18 Apr 1961, Rechinger 21758 (holotype [specified here]: W #1981–10461; other isotypes: B, G, L, hb. Soest).
  • Taraxacum euranum Soest in Bull. Soc. Echange Pl. Vasc. Eur. Occid. Bassin Médit. 16: 130. 1976 [23785].
Isotype: PAL-Gr 23785—France, “Jaudrais (France, Eure-et-Loire), friche argileuse humide le long de la route de Senonches (D 140),” 23 Apr 1975, de Retz 70916 in Soc. Echange Pl. Vasc. Eur. #8011 (holotype: LG; other isotypes: C, G, GENT, M, MA [2×], MSB, P [2×], SEV).
  • Taraxacum exiguum Soest in Bull. Soc. Echange Pl. Vasc. Eur. Occid. Bassin Médit. 16: 130. 1976 [23786].
Isotype: PAL-Gr 23786—France, “Villliers-en-Arthies (France, Val-d’Oise), prairie rase et clairière du bois de la Bucaille,” 25 Apr 1975, de Retz 70935 in Soc. Echange Pl. Vasc. Eur. #8012 (holotype: LG; other isotypes: C, G, GENT, MA [2×], MSB, P, SEV).
  • Taraxacum graniticum Soest in Bull. Soc. Echange Pl. Vasc. Eur. Occid. Bassin Médit. 16: 131. 1976 [23787].
Isotype: PAL-Gr 23787—France, “Gelles (France, Puy-de-Dôme), près de Say-Soubre, alt. 750 m, pré fumé sur granite,” 4 May 1975, Billy in Soc. Echange Pl. Vasc. Eur. #8013 (holotype: LG; other isotypes: C, G, GENT, LD, M, MA, MSB, P, SEV).
  • Taraxacum inconspicuum Soest in Bull. Soc. Echange Pl. Vasc. Eur. Occid. Bassin Médit. 15: 111. 1974 [9266].
Isotype: PAL-Gr 9266—France, “Saint-Genés-Champanelle (France, Puy-de-Dôme), alt. 850 m, pré sur granite, à Thèdes,” 13 May 1973, Billy in Soc. Echange Pl. Vasc. Eur. #7094 (holotype: LG: other isotypes: G, GENT, L, LD, MA [2×], MSB, P [2×], SEV).
  • Taraxacum retortum Soest in Bull. Soc. Echange Pl. Vasc. Eur. Occid. Bassin Médit. 16: 131. 1976 [23797].
Isotype: PAL-Gr 23797—France, “Urdos (France, Pyrénées-Atlantiques), le long de la route du Col du Somport, 200 m avant la frontière avec l’Espagne, alt. 1600 m,” 13 Jun 1975, de Retz 71454 in Soc. Echange Pl. Vasc. Eur. #8023 (holotype: LG; other isotypes: C, G, GENT, L, LD, MA [2×], MSB, P [3×], SEV).
  • Taraxacum roseolepis in Bull. Soc. Echange Pl. Vasc. Eur. Occid. Bassin Médit. 16: 132. 1976 [23799].
Isotype: PAL-Gr 23799—France, “Villiers-en-Arthies (France, Val-d’Oise), prairie arbustive le long de la route de Villarceaux (D 142),” 24 Apr 1975, de Retz 70938 in Soc. Echange Pl. Vasc. Eur. #8025 (holotype: LG; other isotypes: C, G, GENT, LD, M, MA [2×], MSB, P, SEV).
Holotype: PAL-Gr 13683—Greece, Crete, “Ep. Lassithi: N-Hang des Berges Spathì von der Alp Lekanìda bis zum Gipfel, 1650–2150 m.ü.M., Kalk. Auf lockerem u. verfestigtem Schutt verbreitet u. meist hfg.,” 23 Jun 1961, Greuter 3683 (isotypes: G, LD, W, Z).
  • Tetrazygia decorticans Bécquer in Willdenowia 37: 314. 2007. (≡Miconia decorticans (Bécquer) Bécquer and Majure) [https://doi.org/10.3372/wi.37.37120 (accessed on 6 March 2024)].
Isotype: PAL-Gr 121242—Cuba, prov. Sancti Spíritus “Reserva Ecológica Alturas de Banao, base de la Teta de Juana, orillas del arroyo del tunel de Caja de Agua,” 640 m, 21°51’50.1″ N, 79°35’59.7″ W (holotype [specified by Bécquer in Revista Jard. Bot. Nac. Univ. Habana 44: 189. 2023]: HAJB #897; other isotypes: B, FLAS, GH, HAC, HAJB ##898–899 [2×], JE [2×], K, MO, NY, US).
Isolectotype: PAL-Gr 5249—Greece, “m. Malevo Laconiae,” Jun 1896, [Leonis] in Heldreich, Pl. Exsicc. Fl. Hellen. (lectotype [designated here]: WU (Halácsy, Greek hb.); other isolectotypes: none traced).
Isotype: PAL-Gr 39366—Israel, “Jerusalem,” May 1846, Boissier (holotype [specified here]: G-BOIS #G404534; other isotypes: JE [2×], K, MEL).
Holotype: PAL-Gr 53188—Greece, “Nom. Arkadia, Ep. Megalopolis: NE foothills of the Taijetos range, c. 0.5 km W of Neochori, alt. c. 1100 m, 37°11′0″ N, 22°13′45″ E. Mixed fir, pine and chestnut wood by a stream, on mixed substrate,” 11 Jun 1995, Kamari & al. in OPTIMA Iter Medit. 7 #2188 (isotypes: B, UPA).
Holotype: PAL-Gr 40262Cuba, “Prov. Camagüey, km 3 de la carretera a Lesca al nordeste de Camagüey, alt. 120 m, 21°27′10″ N, 77°50′00″ W. Matorral xeromorfo espinoso degradado en serpentina,” 2 Mar 2000, Greuter 25137 & al. (isotypes: B, HAJB, HIPC, US).
  • Ventenata subenervis Balansa in sched. Pl. d’Orient, 1854: #7. 1854 [8494].
Isotype: PAL-Gr 8494—Turkey, “Smyrne, sur les collines incultes,” 8 May 1854, Balansa in Pl. d’Orient, 1854 #7 (holotype [specified here]: P #3645076; other isotypes: B, BM, BR [2×], FI, FR, G [2×], GOET, K [3×], KFTA, MPU, P [15×], US, W [3×], WAG).
  • Veronica stamatiadae M. A. Fisch. and Greuter in Pl. Syst. Evol. 125: 249. 1976 [27056].
Isotype: PAL-Gr 27056—Greece, “Insula Kastellorizo (Megisti) Dodekanesi, Insula Ro: ad orientem sinûs Ajios Jeorjios, alt. 10–30 m. In rupestribus calcareis dumulosis. Ad rupes,” 12 Apr 1974, Greuter 11808 (holotype [specified here]: WU #70317; other isotypes: ATH, C, ERE, G, H, WU #70316).
Holotype: PAL-Gr 27107—Greece, “Insula Kastellorizo (Megisti) Dodekanesi, in scopulo Psomi, alt. c. 15 m. In rupestribus calcareis. Flores coeruleo-violascentes,” 17 Apr 1974, Greuter 11938 (isotypes: ATH, G).
Isolectotype: PAL-Gr 66704—Syria “borealis. Aleppo. Très remarquable par ses feuilles courtes, son calyce très-obliquement [tronqué] et son fruit longuement stipité dans le calyce. A placer [près] du V. sericocarpa! [manu Boissier],” 1865, Haussknecht (lectotype [designated here]: JE #6825; other isolectotypes: G-BOIS, JE # 6827).
Isotype: PAL-Gr 66362—Greece, “Agrapha (Dolopia veterum): in reg. super. Pindi summi montis Karáva alt 5500–6500′ substratu schistoso,” 1–3 Jul 1885, Haussknecht (holotype [specified here]: JE #20586–20587 [2 sheets]).
  • Zanthoxylum rolandii Beurton in Feddes Repert. 97: 35. 1986 [126212].
Isotype: PAL-Gr 126212—Cuba, “Prov. Pinar del Río, al pie oeste de la Loma Peluda (Preluda) de Cajálbana, cuabales, serpentina,” 21 Jan 1981, Berazaín & al. in HFC 43476 (holotype [specified here]: HAJB ##958; other isotypes: B [2×], HAJB #959–961 [3×], JE [2×]).

5. Conclusions

The type herbarium in PAL-Gr is composed of four packages with 338 sheets, corresponding to 335 specimens, three of them of two sheets each (Code, Art. 8.3), and pertaining to 327 different names. Grouped by type categories, there are 61 holotypes (+2 fragments), 6 second-step holotypes, 236 isotypes, 3 lectotypes (+1 fragment), 17 isolectotypes, 1 neotype, 2 isoneotypes, and 1 isoepitype. Of these, 82 are here newly designated (or, in 67 cases, specified), but 22 remain unspecified first-step holotypes, as the information necessary for the specification of a second-step type is lacking.
A grouping of the type specimens (not counting duplicates) by country/region of origin reveals the large predominance of Mediterranean, principally Greek material:
Albania7
Armenia5
Bosnia and Herzegovina1
Bulgaria1
Cyprus1
Greece150
France13
Israel3
Italy70
Libya2
Montenegro3
Morocco1
Portugal2
Serbia3
Spain9
Syria1
Turkey2
Σ Mediterranean274
Austria4
Germany4
Σ Central Europe8
Australia13
Cuba31
USA1
Σ Extra-European45
Σ Total327

Funding

No external funding of this research was received.

Data Availability Statement

Data are contained within the article.

Acknowledgments

Rosa Rankin Rodríguez, Palermo, helped with the protologue scans and the scans of Cuban type-specimens. Natale Surano, Palermo, took charge of uploading the digital images onto the Internet. The Foundation Herbarium Greuter bore the publication costs.

Conflicts of Interest

The author declares no conflicts of interest. The funders had no role in the design of the study, in the collection, analysis, or interpretation of data, in the writing of the manuscript, or in the decision to publish the results.

Reference

  1. Turland, N.J.; Wiersema, J.H.; Barrie, F.R.; Greuter, W.; Hawksworth, D.L.; Herendeen, P.S.; Knapp, S.; Kusber, W.-H.; Li, D.-Z.; Marhold, K.; et al. (Eds.) International Code of Nomenclature for Algae, Fungi, and Plants (Shenzhen Code), Adopted by the Nineteenth International Botanical Congress, Shenzhen, China, July 2017; Koeltz Botanical Books: Glashütten, Germany, 2018; [Regnum Veg. 159.]. [Google Scholar]
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Greuter, W. The Greuter Herbarium in Palermo: An Inventory of Its Type Specimens Available Online, with Some Thoughts on Type Terminology (Occasional Papers from the Herbarium Greuter, N° 5). Plants 2024, 13, 1086. https://doi.org/10.3390/plants13081086

AMA Style

Greuter W. The Greuter Herbarium in Palermo: An Inventory of Its Type Specimens Available Online, with Some Thoughts on Type Terminology (Occasional Papers from the Herbarium Greuter, N° 5). Plants. 2024; 13(8):1086. https://doi.org/10.3390/plants13081086

Chicago/Turabian Style

Greuter, Werner. 2024. "The Greuter Herbarium in Palermo: An Inventory of Its Type Specimens Available Online, with Some Thoughts on Type Terminology (Occasional Papers from the Herbarium Greuter, N° 5)" Plants 13, no. 8: 1086. https://doi.org/10.3390/plants13081086

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