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1 September 2017

16 Pages

Near-Term Pathways for Achieving Forest and Agricultural Greenhouse Gas Mitigation in the U.S.

,
and
1
Department of Public Administration, North Carolina State University, Raleigh, NC 27695, USA
2
Energy Initiative and Nicholas Institute for Environmental Policy Solutions, Duke University, Durham, NC 27708, USA
*
Author to whom correspondence should be addressed.

Abstract

U.S. forests and agriculture present unique opportunities to mitigate greenhouse gas (GHG) emissions. U.S. forests currently remove a large amount of carbon dioxide from the atmosphere each year and store it as a terrestrial carbon sink, a trend that is projected to continue, although at a decreasing rate over time. Agriculture is and will continue to be a net source of GHGs. To encourage additional mitigation, analyses suggest addressing forest loss, forest aging, wildfire, and encouraging greater forest growth. In agriculture, analyses suggest addressing animal operation methane emissions and nitrous oxide from fertilizer use. Absent new targeted policies to encourage mitigation practices such as these, existing programs may need to be better leveraged for GHG mitigation, even if that is not their explicit objective. Leveraging existing programs requires coordinated outreach efforts to ensure that practices are not cross-purposed. Development of standards and verification practices is also necessary to ensure desirable outcomes. Finally, greater mitigation may be possible by maximizing the effectiveness of voluntary efforts from private and non-governmental organizations, and not necessarily the implementation of new policies. This conclusion represents a departure from traditional commentary on the subject, but arguably represents a more realistic path forward to achieving climate mitigation objectives in the near-term.
For decades, research has sought to estimate both the consequences of global climate change on agriculture and forestry, as well the potential greenhouse gas (GHG) mitigation available from the two sectors [1,2]. More recent research has likewise sought to describe the policy and market means by which to achieve these contributions [3,4,5,6]. In practice, however, policies and incentives have yet to emerge to support forest and agricultural GHG mitigation efforts on a broad scale. Questions thus remain as to how agricultural and forest mitigation can be achieved in the near-term, defined here as the next 10–15 years, absent policies specifically targeted to addressing climate change.
The assessment below begins with a review of estimates of GHG mitigation potential from U.S. farms and forests. We discuss mitigation potential as identified in the literature and by public entities, providing estimates of both technical and economic mitigation potential where available. We then review the policy or market instruments capable of facilitating mitigation in the agriculture and forest sectors. Next, we briefly discuss the likelihood of new policy or market development in the near future, concluding with a brief discussion on the most likely path or paths for achieving GHG mitigation in U.S. forests and farms in the coming decade.

2. Policy Options to Achieve GHG Mitigation in the Forest and Agricultural Sectors

The previous section considers baseline levels of GHG emissions and carbon sequestration in forestry and agriculture as well as levels that could be achieved by enacting policies aimed at mitigating GHGs. Such policies could, in principle, fall into one of four categories.

2.1. Direct Government Action

Governments can act on regulatory authority they have over land use to require certain activities to mitigate GHGs on private lands. As a general matter, private land-use restrictions are more of a state and local matter than a federal one in the United States, though they may be in response to a federal statute (e.g., the Clean Water Act). While states, counties and municipalities do enact land-use ordinances, they are seldom if ever targeted at GHG mitigation. For instance, there may be state forest practices acts that limit or exclude timber harvesting in certain areas, but these are not typically aimed at forest carbon sequestration as a primary goal.
A number of resource agencies at the federal and state level are also responsible for managing publicly owned lands and must decide how to manage them. For instance about 250 million acres of forestland are managed by the federal government [7], primarily through the U.S. Forest Service and the Bureau of Land Management. The management plans governing these agency actions go through extensive planning and review processes and could well be developed to take forest carbon sequestration or other GHG mitigation benefits into consideration [30].
Governments at different levels can also mandate—or, in some cases, establish non-mandatory goals—that a certain amount of energy is produced by renewable sources. Currently, about 30 U.S. states have renewable portfolio standards that mandate minimum percentages of electric power from renewable sources, including biomass [31]. These standards create an economic incentive for the production and use of biomass as an energy feedstock.

2.2. “Carbon” Price

A carbon price has long been considered a possible option for mitigating GHGs from forestry and agriculture [15]. This type of policy mechanism creates a price per unit of GHG (usually per ton of CO2e) that rewards reductions in GHG emissions. For agriculture and forestry, this could be reductions of CO2, methane (converted to CO2e for pricing purposes), or nitrous oxide and/or increases in CO2 removed from the atmosphere and sequestered in forest pools or soil.
The carbon price approach follows from consideration of various “cap-and-trade” proposals at the U.S. federal level starting midway through the first decade of the 21st century. The policy momentum for carbon pricing at the federal level started in earnest with the introduction of the bipartisan McCain-Lieberman Climate Stewardship Act (S 139) in the Senate in 2003. While the bill did not pass, it received more votes than anticipated and set the stage for successor bills in both chambers of Congress, culminating in the Waxman-Markey American Clean Energy and Security Act (HR 2454), which passed the House of Representatives in 2009, but failed to advance in the Senate. While Waxman-Markey covered about 85 percent of the country’s emissions, forests and agricultural sources themselves were not subject to the cap. However, reductions and sequestration in those sectors were eligible to generate emissions offset credits, which could be sold at the carbon price to firms in the regulated sectors that did have compliance obligations.
There has been little momentum to establish a national carbon market since July of 2010, when the U.S. Senate announced it would not consider the companion bill to HR2454. California did enact a cap-and-trade program that was similar in scope, meaning it does have offsets for some forest and agricultural activity. So far, however, California forest and agriculture offsets have been limited in scale—far less than the mitigation potential estimates discussed in the previous section. Nine northeastern U.S. states instituted the Regional Greenhouse Gas Initiative in 2009, a cap-and-trade program for power plant CO2 emissions, with forest and agriculture offsets eligible, but program prices have been too low to induce any offsets to market.
While the California and RGGI programs create a carbon price through market-based regulatory action, there is also a carbon price through markets driven by voluntary demand for carbon offsets (e.g., the American Carbon Registry (ACR), Climate Action Reserve (CAR), and Verified Carbon Standard (VCS)). These programs cover a larger portfolio of agriculture and forest activities than do the California and RGGI regulatory programs. In addition to the agriculture categories in California and RGGI, the voluntary market creates offset demand for soil carbon sequestration from agricultural practices, nitrous oxide reductions from fertilizer management, grassland management, livestock management (including enteric fermentation), avoided conversion of grasslands, and wetlands restoration. These activities cover a majority of the full agricultural emission reduction potential referenced in the previous sections. Yet, these voluntary carbon market activities have not had a high rate of adoption, either in terms of the number of projects undertaken or the percentage of credits generated [32]. Low adoption rates may be due to a general lack of demand for voluntary offsets, the economic particulars of agricultural and forest mitigation activities, or both.
While there has been limited momentum for the U.S. to incorporate forestry or agriculture into regulatory compliance-driven carbon pricing systems, it is illustrative to look at countries that have explored this option. One country—New Zealand—has a regulatory system that includes these sectors in some fashion. Forestry is part of New Zealand’s economy-wide emissions trading system [33]; land that was in forest before 1990 must submit New Zealand Units (NZUs) to cover emissions that occur from deforestation. If the land is harvested and replanted, there is no emissions compliance obligation. Land established as forest after 1989 can opt in to the Emission Trading Scheme (ETS) to earn “removal” credits for the amount of carbon removed from the atmosphere and stored as carbon stocks, which they can sell to other parties who have to meet emissions obligations. Though some exemptions exist, New Zealand forest owners generally do face a carbon price and thereby have incentives to keep carbon stored in forests. In contrast, New Zealand’s agriculture sector, which accounts for 48 percent of New Zealand emissions [34], is not subject to compliance obligations to cover its emissions by submitting NZUs, though there is mandatory reporting of emissions from the sector [35].
Australia’s Carbon Farming Initiative (CFI) incentivizes farmers to mitigate GHGs from agricultural and forest land practices by generating emission offset credits for those actions on a per tonne CO2 equivalent basis. Farmers can sell these credits through a government-funded Emissions Reduction Fund or to private party buyers in the voluntary offsets market [36]. The CFI at one time had been seen as a core component of Australia’s economy-wide cap-and-trade emissions trading system, but that system was repealed in 2014.
Common themes emerge from the carbon pricing experiences in the U.S., New Zealand and Australia. One is that the sectors have remained largely out of emissions compliance requirements (with the limited exception of New Zealand forests) and have been primarily included in voluntary offset credit generation efforts, which limits the strength of incentives for mitigation. Accordingly, the uptake by farmers has been low. Substantial work has been aimed at developing market-based incentives for farmers and forest owners, however, as crediting protocols have been developed and outreach efforts have been expanded.

2.3. Government-Financed Incentives

Another way to induce private landowners to undertake mitigation activity voluntarily is for the government to pay them to do so. There is a long history of these types of programs in the U.S., for instance the Conservation Reserve Program (CRP) and the Wetlands Reserve Program (WRP), administered by the U.S. Department of Agriculture (USDA). These payments are not specifically targeted for carbon or GHGs, though they could be modified to be. Whether or not such a modification would occur, the actions that are paid for (e.g., tree-planting, wetlands restoration, fallowing, adoption of no-till agriculture) generally provide some measure of GHG mitigation; the 2016 U.S. Biennial report, for example, estimates that the CRP and activities administered under the Natural Resources Conservation Service alone contribute approximately 50–60 TgCO2e/yr of GHG mitigation [14].
The USDA also issues Conservation Innovation Grants (CIGs) to “stimulate development and encourage adoption of innovative conservation approaches and technologies.” Some of these grants have been specifically targeted for GHG mitigation activity. Though relatively small in scale, the idea behind CIGs is to develop innovative programs that are further disseminated to ideally enhance broader adoption.
The government can likewise incentivize land-based GHG mitigation activity through tax incentives, although this is often an indirect incentive for farmers. For instance, the federal tax code has had various subsidies for fuel producers to produce biofuel blends, which creates a demand for biofuel feedstocks that incentivizes farmers to produce those feedstocks [37]. These credits are in various states of play, with some recently extended and others recently terminated but subject to industry efforts to reinitiate [38].

2.4. Information Programs and Technical Assistance

The USDA, in cooperation with land-grant universities throughout the United States, has provided technical assistance to farmers and forest owners for over 100 years through agricultural and forest extension programs established by the Smith-Lever Act of 1914. These extension programs, which aim to anchor their technical assistance in scientific evidence-based practice, cover the full range of agricultural and forest management activity. In the last decade, this has grown to include technical guidance on practices that mitigate GHGs (see, e.g., [39]). This technical assistance allows farmers to consider whether and how to engage in the agricultural offsets referenced in the carbon pricing discussion above, or perhaps the private voluntary actions as discussed below.
The EPA’s AgSTAR program is specifically aimed at providing information and data on biogas recovery in the agricultural sector, primarily from animal manure management. The underlying notion is that biogas (essentially methane) can be generated by anaerobic digestion and captured rather than be emitted directly into the atmosphere. The captured biogas can also be used as a productive energy input if attached to an electricity generating device or conditioned and injected into a natural gas pipeline [40]. By encouraging the destruction of methane that would otherwise be emitted to the atmosphere, these programs can produce GHG benefits, especially if the captured methane is used as a substitute for fossil fuels. Possible revenue streams for farmers adopting these technologies include sales (or cost displacement) of electricity and natural gas, as well as sales of renewable energy credits to meet renewable portfolio standards and GHG emission offset credits as referenced above [41].

3. Near-Term Opportunities and Incentive Streams

Since taking office in January of 2017, the Trump Administration has indicated that the use of federal policy to reduce national GHG emissions will be a much lower priority than it was in the previous Administration [42]. There nonetheless remains interest across various stakeholders about what actions could be taken to motivate GHG mitigation in the near-term. We consider three general streams of policies here for agriculture and forestry. The first is the suite of existing policies and programs already in place at federal, state, and local levels. The second consists of new policies or programs that could be included in the upcoming federal Farm Bill reauthorization legislative process or developed under existing authorities at the USDA or other federal agencies. The third includes regulation, funding, or direct-action strategies implemented by non-federal actors, private finance, or non-governmental organizations. Each is further discussed below.

3.1. Continued Use of Existing Policies and Programs

There are a variety of policies and programs already being implemented by federal and state entities with the potential to encourage greater GHG mitigation in forestry and agriculture. The U.S. Biennial report referenced above notes several in its review of current policies and measures, including the RFS, conservation programs like the CRP, stewardship and management programs operated by the Natural Resource Conservation Service, biogas reduction operations under the AgSTAR program, and hazardous fuels reduction and forest restoration programs [14]. GHG mitigation estimates are not available for each and every activity listed, but could collectively yield well over 200 TgCO2e/yr in annual reductions by 2020.
Recent analysis highlights the role that existing efforts have in driving GHG emissions and sequestration in the U.S. land sector. In a review of current policies with the potential to affect GHG mitigation in forests, agriculture, and other, non-working lands, McGlynn et al. consider the types of policy actions referenced in the previous section (e.g., regulations, government incentives, data and technical assistance, etc.) [4]. Across these categories of activities, the review identifies 70 authorizing or facilitative policies or programs in operation at the time of the report. Arraying the budgets for these practices (a rough proxy of programmatic reach or capacity) by the area of effect (a rough proxy of the GHG sink or flux they could influence) creates a visual indication of both gaps and opportunities present in current policy (Figure 2).
Figure 2. Policy scope and area reach of selected U.S. policies with the potential to influence GHG mitigation in forests and agriculture. Green circles represent net GHG reductions, yellow circles represent uncertain mitigation benefits, and red circles represent net increases in GHG emissions. Source: [4].
Apart from program summaries and analyses, efforts have been made in recent years to better coordinate existing programs and authorities to achieve greater GHG mitigation in practice, such as the aforementioned USDA Building Blocks initiative. Even in the absence of new authorities, efforts to better coordinate or communicate the existence of tax and other incentives can help to facilitate preservation or sustainable management activities [43,44]. If mitigation is to be achieved or facilitated through existing programs, many of which may not be specifically targeted to carbon storage or GHG emissions reduction, it is necessary to first identify relevant programs and then determine how they can be best leveraged to achieve climate benefits [4]. Such an exercise is well beyond the scope of this short review, but a few examples are nonetheless worthwhile to provide. For example, Clean Water Act incentives for increased nutrient management by Confined Animal Farming Operations (CAFOs) could have ancillary GHG benefits by addressing ammonia and other pollutants [26]. Habitat mitigation requirements as driven by the Endangered Species Act or wetland mitigation requirements motivated by the Clean Water Act and its implementing regulations could also serve as a vehicle to also emphasize GHG mitigation [4].
Opportunities also exist in state and local-level programs to encourage GHG mitigation in forest and agricultural sectors. To date, California and the northeastern states in the RGGI program plan to continue and even strengthen their cap-and-trade policies into the future and other states have pledged to pursue climate action following the decision of President Trump to withdraw the United States from the Paris Agreement [45]. These efforts are unlikely to fully compensate for a near-term absence of climate policy at the federal level, and the effect that expansion of state programs would have on incentives for land-based GHG mitigation is far from clear, but they do underscore that actions on climate change emerge from a dynamic involving many different levels of government and a diverse set of constituencies.
There likewise exists countless non-GHG policies enacted at the state and local level with the potential to influence mitigation efforts in the forest and agriculture sectors. “Smart growth” strategies implemented by state and local governments can help to reduce working land competition with urban development [19]. Habitat management and protection provisions meanwhile can help to encourage the maintenance of vegetation (and by extension, carbon) on those sites subject to development [46]. The GHG mitigation achieved through these and similar programs will be a function of the practices encouraged (e.g., fertilizer management, forest retention, soil carbon management), thus the wide array of program designs and conditions under which they are implemented makes estimation of net GHG reductions incredibly difficult.

3.2. New Farm Bill or Agency Initiatives

In their review of the literature of forest and agricultural mitigation potential, McGlynn et al. detail over two dozen new policies or practices that could be implemented to increase GHG mitigation [4]. Many of these programs and practices require new authorities, however, limiting their near-term potential. At the same time, negotiations around the primary vehicle for U.S. federal agricultural policy—the Farm Bill—have begun in earnest. The Farm Bill operates on approximately a five-year schedule, and provides both authorization and appropriations for a wide array of policies and programs with the potential to affect the way forests and farms are managed.
Though is unlikely that new policies or practices to specifically address GHG mitigation will find their way into the latest iteration of the Farm Bill, it is possible that new programs unrelated to GHG mitigation could still bring additional carbon storage or GHG emission reduction as a co-benefit. One example worth noting is the Soil Health and Income Protection Program (SHIPP) proposed by Sen. John Thune (R-SD) [47]. SHIPP is a short-term (3–5 years) conservation program for low-productivity acres that provides rental payment and crop insurance assistance. The program, requires a “low-cost perennial” to be planted on enrolled acres, with possible harvest after nesting or brooding season. Such a policy could generate GHG reductions, for example, should it be effective in stemming the reversion of currently-uncultivated lands into crop production [48,49].
More broadly, support for conservation programs remains high [50]. Though programs such as the CRP have seen decreased enrollment in times of high commodity prices, a recent run of low prices has led to a call to increase the cap on CRP acres [51]. Current discussions suggest an increase to 30 million acres, a 6 million acre increase over the current cap as set by the 2014 Farm Bill [51]. Depending on the conservation practices implemented, increases of this magnitude hold significant GHG mitigation potential. Recall that the USDA Building Blocks aim to increase GHG mitigation by 0.8 TgCO2e/yr by reforesting an additional 400,000 acres of CRP land [20]. Elsewhere, Wear and Coulston consider a hypothetical, CRP-like afforestation policy across approximately 20 million acres of land, nationwide, yielding an increase of 689 TgCO2e/yr between 2013 and 2025 [52].
Changes have also been suggested for the CRP and other easement-based programs to allow increased flexibility in management of enrolled lands. Sen. Thune proposed allowing enrolled lands “to be mitigated for land of equal or greater conservation and wildlife benefit” [51]. Important details are missing on how such a program would operate, but the general concept suggests the possibility of using federal conservation programs as a stepping-stone to increased environmental benefits, potentially including GHG mitigation. There remains an open question as to whether carbon can be included in the calculation to meet the threshold of “equal or greater ecological and economic values compared to the acreage that is mitigated” [51].
In addition to support for conservation programs, multiple stakeholders have identified farm safety net programs as an important concern going into the next Farm Bill, particularly those programs that guard against aforementioned long-term price declines. While there is increased interest in these programs, there is also a general sense that there will be less money to go around [53]. Past studies have assessed opportunities to restructure support programs to facilitate carbon storage and cost savings, showing the potential to increase carbon sequestration by more than 366 TgCO2e/yr at a cost savings of over $6 Billion [54], but a policy shift of this magnitude is presently unlikely.
The concept of using crop insurance to encourage climate adaptation and mitigation behaviors has also emerged in recent years [19], though specific provisions encouraging this have yet to be seen in early Farm Bill discussions. The same is true for increased use of wood products, such as deployment of tall wood buildings [19,20]. In the case of both crop insurance reform and the creation of new markets for forest products, GHG mitigation can be yielded through increasing the incentive to invest in higher carbon land uses and reducing the incentive to convert to lower carbon ones [22,55]. The use of wood products to substitute for other building materials also has potential GHG benefits owing to the greater amount of emissions associated with the production of steel or cement alternatives [56].
It is equally possible that existing programs that currently achieve some measure of GHG mitigation could be modified or removed completely, reducing the amount of mitigation achieved by farms and forests. For example, Monke (2017) reports that there are several dozen Farm Bill programs that operate without a budget baseline, making them more vulnerable to reduced or eliminated budgets in future Farm Bill reauthorizations [57]. While most of these programs are largely irrelevant to agricultural and forest mitigation, multiple bioenergy programs are potentially vulnerable, including the Biomass Crop Assistance Program (BCAP) and the biorefinery assistance program. It is difficult to approximate the marginal contribution each of these programs contribute to current patterns of GHG emissions and carbon sequestration, though one analysis suggests that BCAP operating alongside the RFS reduces net GHG emissions by 300 TgCO2e/yr over the course of a decade relative to the RFS operating by itself [58].

3.3. Private Sector and NGO Interventions

With uncertainty about the scale and scope of government policy efforts to reduce GHGs, voluntary mitigation efforts in agriculture and forestry may be an important path for the next several years in the United States [59]. For example, Lewandrowski and Zook (2015) identify a host of other vehicles by which farmers could engage more in agricultural mitigation [60], including private sector supply chain initiatives and joint public-private partnerships focused on voluntary GHG mitigation. Facing pressure from investors and consumers alike, corporations have demonstrated an ability to address environmental concerns in the agricultural and forest sectors. The world’s largest retailer Walmart has long been an impetus for these efforts and in early 2017 pledged to reduce one gigaton of GHG emissions from its supply chain, including products originating in the agriculture and forestry sectors [61]. Other prominent examples include Smithfield Foods, Cargill, Monsanto, and General Mills [62,63]. Taken both individually and as a whole, these private sector and NGO activities have the potential to expand land-based mitigation, but their scale will depend on sustained public and private sector budget commitments. In recognition of this potential gap in either capacity or motivation, recent years have seen an expansion of private conservation finance, a prominent example being NatureVest, a joint initiative between The Nature Conservancy and JP Morgan Chase & Company [64].

4. Conclusions, Challenges, and Opportunities

So what can be said of near-term opportunities for GHG mitigation in U.S. agricultural and forest sectors? If we are to begin by addressing the greatest emission sources and sinks, then available data suggest that early efforts in the agricultural sector should focus on addressing methane emissions from animal operations and nitrous oxide emissions from fertilizer use. In forestry, the data suggest an emphasis on addressing forest loss, forest aging, and wildfire, while encouraging greater forest growth. Apart from the magnitude of mitigation potential, other considerations are likewise relevant, such as cost and the timing of mitigation (Table 2). There are trade-offs to consider, as well. For example, some activities are more expensive in absolute terms but result in greater mitigation (afforestation, for example), yielding a lower per-ton cost of mitigation. Some activities like afforestation may also take long periods of time to yield significant mitigation, whereas others may yield GHG emission reductions on much shorter timescales (e.g., fertilizer management). Finally, there is a need to carefully assess the broader implications of any strategy to increase GHG mitigation. An important example in this regard is any effort to address a decline in forest sequestration over time, as well as the difficulties associated with appropriately accounting for sequestration changes given the complexities of changing climate and associated disturbances. For example, replacing older, slower growing trees with younger ones may lead to an increase in emissions, as might any effort to reduce long-term emissions from catastrophic wildfire [65,66].
Table 2. Relative magnitude, cost, and timing of select practices to increase GHG mitigation in the U.S. forest and agricultural sectors. The number of symbols in each column provides a relative ranking of practice performance, ranging from low to high (for magnitude and cost) and short to long (for timing). “Varies” indicates substantial range for a practice depending on the details of a particular application.
As the review above indicates, there are already a large number of programs with the potential to achieve varying amounts of GHG mitigation across forest and agricultural sector activities. Greater use of existing programs is one alternative to the creation of new ones, implying that outreach and engagement efforts are warranted to ensure that available resources are being leveraged as well as possible. The USDA Building Blocks represent but one approach to use existing programs to generate additional GHG mitigation, but countless other opportunities exist across local, state, and federal governments and in the private sector.
One concern with consciously leveraging a variety of individual programs to achieve GHG mitigation is coordination, or ensuring that the programs are not cross-purposed or that they do not result in negative co-effects (e.g., conversion of native habitat in favor of high-production monocultures). A wide array of programs not specifically designed to achieve GHG mitigation may be increasingly relied upon to achieve just that. As a result, greater emphasis on the development of standards and installation of appropriate verification practices is necessary to ward off unintended negative consequences. For example, research has documented the potential of indirect market effects to negatively affect GHG mitigation efforts by inducing the increased use, production, or consumption elsewhere, be it in the case of biofuel production [23], timber set-asides [67], or expanded use of wood products [56]. Expanded use of existing programs likewise requires careful attention to setting an appropriate baseline so as to judge additional contributions. Failing to distinguish between what would have been done anyway under existing authorities and what is the product of new outreach initiatives, expanded program budgets, or expanded program offerings threatens to inflate reported gains regardless of actual mitigation achieved.
Despite a robust and established literature on GHG mitigation opportunities in the forest and agricultural sectors, actions to realize those opportunities have been limited. Though one may ask whether agricultural and forest mitigation is better suited to a carbon price approach, as discussed above, or as part of complementary policies and private sector initiatives (e.g., [60]), the argument is somewhat moot at the present point in U.S. history. Practitioners must pursue the options available to them. Recent statements and committed practices by private and non-federal government stakeholders suggests that there is a willingness to pursue GHG mitigation, so the most important question is how to tap that willingness with the policies and mechanisms in place, rather than how to design the optimal policy framework.

Acknowledgments

The authors gratefully acknowledge research assistance provided by Emery Kiefer and Nicholas Riggs. The views, findings, conclusions, and recommendations expressed in this article are those of the authors. Partial funding was provided under USDA Cooperative Agreement number 58-0111-16-005.

Author Contributions

C.G. and B.M. jointly conceived the analysis, C.G., B.M. and M.P. wrote the paper.

Conflicts of Interest

The authors declare no conflict of interest.

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