3.1. Birds of Prey
Our evaluations show that birds of prey are detected in archaeological assemblages with less constancy than the mammalian species discussed below. The constancy with which even the most frequent raptor species are found at archaeological sites of a particular epoch is often only 1–2%. White-tailed eagles (Haliaeetus albicilla) are usually the most abundantly recorded raptor species in all three study regions and in all periods investigated. This huge bird, which lives near all large bodies of water, was an attractive hunting bird of prey. However, the results of our study show striking developments in space and time concerning the constancy of white-tailed eagles and other birds of prey.
At Mesolithic forager stations, bones of white-tailed eagles are found with high constancy throughout the study area. In western CE and eastern CE, they are recorded at every fourth to fifth site (a constancy of 23% and 20%, respectively), which is an enormously high value for birds of prey (
Table 4 and
Table 5). White-tailed eagles are also the raptor species with the highest constancy in FS, although the value here is significantly lower, at 5% (
Table 6). Overall, raptor bones are much rarer at Stone Age sites in FS than in western CE. The data basis for eastern CE is considerably smaller, but in eastern CE there are more similarities to western CE than to FS.
Records of birds of prey in burial contexts are rare. Only at the cemetery island of Oleniy Ostrov, located on Lake Onega, Karelia, is it evident that the osprey (
Pandion haliaetus) played a special role, at least in the regional ideology, in about 6400 BCE [
23,
24]. In one case, two legs of an osprey were deposited by the body of an elderly man, and, in another case, parts of a wing were deposited by the body of a child. In the context of our study, it is particularly noteworthy that, in addition to the remains of this bird of prey, the same graves also contained tooth pendants made of elk and beaver incisors. Bones from the white-tailed eagle were found at Oleniy, too, but none of these could be related to a particular grave [
23]. Additionally, at the Neolithic burial site of Tamula (Estonia), in about 4200 BCE, an adult male was buried together with parts of a golden eagle [
23,
25]. All the other Stone Age remains of raptor species from the study area derive from settlement contexts.
It is noticeable in all study regions that the evidence of birds of prey becomes rarer with the beginning of the Neolithic age. In FS, this trend continues into the Iron Age, in eastern CE even to Roman Period, when birds of prey can only be detected sporadically. The few records from the Roman Period come almost exclusively from eagles—white-tailed as well as golden eagles (Aquila chrysaetos). The development in western CE is somewhat different. Here, the numbers and frequency of birds of prey in the archaeological material increase again in the Iron Age. The constancy values are still much lower than for bear, elk or beaver, but at least an increase in the relative number of records can be observed for white-tailed eagle, golden eagle, goshawk (Accipiter gentilis), and red kite (Milvus milvus), as compared to earlier epochs.
From the Iron Age to the late medieval period, the relative number of records increase slowly but steadily for many raptor species in western CE, albeit at a low level. This applies, in particular, to the white-tailed eagle, goshawk, sparrow hawk (Accipiter nisus), common kestrel (Falco tinnunculus), peregrine falcon (Falco peregrinus), red kite, and common buzzard (Buteo buteo). In the Roman period and early medieval, the white-tailed eagle—mostly, again, the raptor species with the highest consistency—can be found at every 20th archaeological site, and even at the late medieval sites at every eleventh site. In the late medieval sites, goshawks are found even more regularly than white-tailed eagles (a constancy of 11%).
In eastern CE, the development is very similar. In no time slice is the number of recorded species of birds of prey higher (this is, of course, also due to the significantly larger number of sites from the late medieval period, so it is also a statistical effect). From early medieval times, goshawks replaced white-tailed eagles as the most frequently detected species and can be found at one in seven sites (a constancy of 14%)—this is a very high value for birds of prey. Other species such as the sparrow hawk, common kestrel, and common buzzard now also reach peak values.
The development in FS was noticeably different to the other regions, at least on a completely different level. Birds of prey also played a minor role here until the Roman Period, but this changed suddenly and completely in the early medieval period. Bones of hawks can now suddenly be found at almost three-quarters of all sites (a constancy of 72%). This is the highest value of all species discussed in this paper. Sparrow hawks and peregrine falcons also reach stable constancy values of over 10%. In late medieval times, the hawk’s constancy declined to a—still high—25%. Presently, the white-tailed eagle has reached a maximum value with evidence at 27% of the archaeological sites. As we will see, they are found in the late medieval period with a much higher degree of continuity than, for example, bears and beavers.
In order to classify these developments, the archaeological background of the finds must be taken into account. For earlier periods, raptors have come to light only sporadically in burials (for Sweden [
26]) but, for the middle and later part of the first millennium AD, mostly the goshawk but also the sparrow-hawk, peregrine falcon, and, rarely, gyrfalcon (
Falco rusticolus) are now found in dozens of burials [
27,
28]. This is the case for parts of FE (Norway and Sweden) and north-western CE (northern Germany), whereas no such finds have come to light from Denmark, England, and the entire Slavic area east of the Elbe River. As we will discuss later, Lithuania is somehow special due to its records.
Sweden is outstanding with c. 45 graves, mostly cremation, which were directly covered by mounds after the funeral pyre had burnt down [
27,
29]. The carefully excavated burial at Rickeby, to the north of Stockholm, demonstrates that entire animals were deposited: one horse, several dogs, several raptors, and other birds. The respective burials in Sweden are usually above average in social standing, and a substantial number of these are men with weapons and helmets, which could be interpreted as members of retinues.
There are also instances beyond Sweden that deserve a mention. Goshawk bones have come to light during the re-analysis of animal remains found in the southeast Norwegian ‘royal’ Viking Age ship burial at Gokstad, dated to around 900 CE, and is the only Norwegian find to that effect so far [
28,
30]. When it comes to western CE, only a handful of inhumation graves with goshawks are known, the oldest of which is that of a wealthy woman in Quedlinburg (c. 500 CE), alongside likewise wealthy men with weapons in Eschwege-Niederhone and Alach [
31,
32]. The latter three are the so-called ‘Gründergräber’, the earliest and at the same time richest in the given cemeteries, which is a hallmark of a new leading family.
The somewhat enigmatic late Vendel Age ship finds at Salme, outer Estonia (c. 700 CE), cannot be elaborated upon here [
33]. The slain warriors in the vessels originate from Sweden, but why did they bring a number of raptors with them? It is also worth a mention that, according to Wigand von Marburg, a scribe of the Teutonic Order, a Duke of the Grand Duchy of Lithuania received many animals for his burial in late 1300 CE, including raptors [
34]. No such thing has been found in burials so far.
Parallel to the appearance of particular raptors in burials, the same bird species also came from settlement contexts in remarkable numbers, although, in this case, skeletons of birds carry more evidential value than single bones (see [
35], as an introduction). Remarkably, the number of Swedish sites with such bones amounts to 40 in the period from c. 600 to 1500 CE and, notably, these are not rural but ‘special sites’: royal seats, trading places, and cities [
27]. Also, one Norwegian royal seat, from early post-1000 CE, in the very south-east of the country (present-day Kungahälla, situated in Bohuslän along the west coast of Sweden) has yielded a number of raptor bones, again mainly goshawks [
36]. The same pattern can be observed in Sweden; the bones of goshawk (but also the sparrow-hawk and peregrine falcon) from special sites can be encountered for in western CE, mostly for fortifications as seats of noble persons in the period post-1000 CE [
37]. As regards raptors from trading sites, such as Groß Strömkendorf (Reric) and also Birka in Sweden, one may argue that these birds were trading goods, as is known from younger periods, whereas for finds in (pre)urban contexts, a close look on the area of provenance is needed; do they originate from ordinary living quarters or other parts of the city [
35]?
Denmark and the west Slavic area in the north of western CE become visible not by raptor species in graves but again by finds from settlement contexts. For instance, partial raptor skeletons originate from Schleswig (layers from the 11th to 14th century CE), with the Danish royal seat nearby. The houses themselves at Schleswig are ordinary in size; however, they stand out in two respects, namely silk finds and actual bird skeletons, which are a rare find [
38]. Totally unique with its minimum number of 41 raptors from several species, including a number of partial skeletons, is the West-Slavic seat of power in Starigard-Oldenburg, with its period of use in late pre-1000 CE [
39,
40]. The situation is more complex later on with a repeated shift in political control between West-Slavic rulers and church representatives (bishops), which has to be omitted here. Raptor bones were also recorded in Mikulčice, the seat of power of the short-lived Great Moravia, mainly in the ninth century CE [
41,
42].
Apart from bones, there is a second archaeological source material, the visual one, that needs a mention; this is represented by riders with raptors on their fist. Recently, a fibula with such an image has come to light in the burial of a woman, dated to the seventh century CE and above average in furnishings, in the cemetery in Münstermaifeld, western Germany [
43]. This find is more or less contemporary with the aforementioned burials from Quedlinburg, Alach, and Eschwege-Niederhone (all western CE). Another similar record derives from a wealthy burial in Stare Mesto, one more seat of power of Great Moravia [
44]. Depictions of riders with raptors on their fist are also known from FS, for the Viking Age and the following centuries [
45,
46]. Outside our main area of concern, but important, such imagery is encountered in mosaics for areas that saw a Germanic intrusion during the Migration Period (these mosaics can probably be ascribed to the Vandals in Tunisia and the Westgoths on the Iberian Peninsula, whereas authors from Greek and Roman antiquity did not know about this kind of hunting [
47]).
There is yet another, third, source material that comes to mind in the present context. Very rarely, birds, bones, or skeletons have been found together with either small metal bells or gloves and hoods of organic materials. Of particular importance is a complete skeleton of a gyrfalcon (here, an exotic species; cf. [
48]), found together with a bell at the medieval French royal seat, which was unearthed beneath the present Louvre [
49]. But, notably, these finds do not speak for themselves, they can only be understood in a broader context.
At the same time, eagle remains were sometimes found in large quantities in settlement contexts, e.g., in the aforementioned Hedeby (western CE; 9th–11th century CE [
50]) or Klaipėda in Lithuania (eastern CE; 13th–14th century CE [
51]. In both cases, the eagle bones were found in common household rubbish. Smaller numbers of such finds are common throughout the medieval period.
3.2. Brown Bear
The constancy of bear remains at archaeological sites is the greatest in eastern CE during almost all periods (
Figure 1a). Two trends can be observed there: After the Mesolithic age, the consistency of bear remains decreases slowly but continuously from 40% to 10%, with its lowest value during the Roman period. Afterwards, in medieval times, there is a renewed, sharp increase to values that are still above those of the Mesolithic age. Exactly opposite is the development in FS. Coming from relatively low values of a maximum of 17%, the constancy of bear remains here reaches high values in the Iron Age (31%) and then an extraordinarily high value of 82% during the Roman period; afterwards, the values drop back to the earlier, relatively low level. In western CE, in contrast, there are no significant variations in the constancy of bear findings over the investigated millennia. The values fluctuated by 16% over the entire period of the study.
The proportion of bears in the NISP of wild animals is always low and mostly between 1 and 2% (
Figure 1b); however, it shows maximum values during the Roman period in all three regions studied. While this peak is hardly pronounced in western CE, it is very clear in eastern CE and FS. Here, 4% (eastern CE) and 6% (FS) of the wild animal bones originate from bears at this time. After the Roman period, the proportions return to the earlier level.
As was the case with the aforementioned raptors, bear remains used to be a rare grave find for earlier times (see [
52] for Central Europe), but they became very numerous in parts of the first millennium AD, again with a particular emphasis on Sweden [
53]. Burials with claws, and rarely with teeth or actual skins, are recorded as a particular burial rite for substantial parts of the Iron Age in both FS (foremost in Sweden and less so in Norway) and northern western CE (Denmark and Germany, but hardly at all in the Slavic region east of the Elbe river [
54,
55,
56,
57,
58]. Two focal ranges can be noticed; one being the transition from the late pre-Roman to the Roman Iron Age and thus c. 2000 years ago, and the other for the middle of the first millennium CE. As a major rule, it seems that bear claws were added to the funeral pyre and later placed in the urns of the deceased. The main part of the burials has been granted one to five claws, which, by the mere number, may indicate a paw rather than a skin. Generally, men and women received such claws in cremation burials, which range from poor to wealthy but, as an exception from the rule, a continental group, again around 2000 years of age, consists of cremated men that are often richly furnished [
59]. Yet another exception are cremation burials in Funen, Denmark, which date to first century CE and have yielded one to several perforated claws [
60].
More exceptions can be found in the burial rite itself; it relates to (mostly) inhumation burials with a deposition of unusual bear-parts. Firstly, this relates to skin remains or groups of claws in all corners of a burial that are indicative of decayed skins, as known from little more than a dozen, mainly in Migration Period, burials in Norway and Sweden [
58]. All these burials, except one, are above average by their other-than-skin furnishings; however, as a matter of fact, five belong to the richest of their time: four men (three with splendid weapons; graves in Snartemo and Evebø, both Norway, and mound two in Högom in Sweden) and one woman (the so-called ‘petty queen’ from Krosshaug in south-western Norway). Most recently, skins have also been proven by microscopic analysis in a few burials, some wealthy, in Finland, with a dating from 9th to 17th century CE. Secondly, a type of perforated bear-tooth or bear-claw, worn on long straps at the hip or thigh, has come to light in around 40 middle class burials of women and sometimes children from the late fifth to the middle of seventh century CE in ‘linear cemeteries’ (Reihengräberfelder), mostly in the Alamannic area in present-day south-western Germany [
55].
The most remarkable finds of bears have not been mentioned so far. These are burials of actual bears known from the northern part of Scandinavia, with a time-depth that covers almost the past two millennia in Norway but only the period post-1000 CE in Sweden [
61,
62]. A few burials in Sweden are well known and well documented, such as that in Sörviken, Stensele Parish, Lapland, excavated in the 1950s. This burial also yielded, as a rare find, an unfired lead bullet, which belonged to a so-called muzzle loader, a weapon that became more frequent in northern Sweden in the second half of 17th century CE.
Unique for the north are the findings from Frösö, Jämtland, in northern Sweden [
63]. Excavations beneath the choir of a medieval stone church brought to light the bones of wild animals, the foremost being bear, together with the stump of a birch tree, surrounded by a dark layer with bones and fire-cracked stones, radiocarbon-dated to the Viking Age. This site has seen intense natural-scientific research and wide-ranging interpretations.
3.3. Beaver
Beaver remains are found regularly and in high constancy in most investigated periods of all three study regions (
Table 1,
Table 2 and
Table 3). In FS, the development of the constancy of their remains at archaeological sites is similar to that of the bear. Similarly, the constancy of beaver remains falls off continuously and sharply from the Stone Age (approx. 35%) to Roman times (1%). In the medieval period, beaver remains are found in higher constancy again, but the values no longer reach the level of pre-Christian periods (
Figure 2a). The development in eastern CE is the opposite (
Figure 2); here, the constancy of beaver remains is generally very high and even tends to increase slightly from the Stone Age (c. 40%) to medieval times (c. 50%). Only during the Bronze Age can a temporary decrease in the constancy of beaver remains be observed.
In western CE, the constancy of the species is very stable from the Neolithic age to the late Middle Ages, with values around 17%. Only in the Mesolithic age are the values higher (28%). Regarding the proportion of beaver in the NISP of remains from wild mammal species, we found striking differences between the three study areas (
Figure 2). While the species in western CE never reaches values higher than 6% (at the maximum in the Neolithic age), the values in eastern CE are consistently high, although with a decreasing tendency. While in the Mesolithic age 33% of all wild mammal bones originate from beavers, the proportion decreases to about 12% in later epochs. In the late medieval period, the value even drops to 5%.
The constancy with which beavers can be detected at archaeological sites in eastern CE is even higher. Except for the Bronze Age, it is always over 40% and even increases continuously over time. In the Middle Ages, beavers can be found at every second archaeological site. In FS, the proportion of beaver in the NISP of wild animals was high in the Neolithic age at 21%, but afterwards the value decreases to a constant level, which is only about 2%.
At a few Stone Age sites distributed over all three investigated regions, beaver dominates the refuse fauna, for example, Kaulenkalns (eastern CE, Latvia; Paaver 1965, cited in [
64]), Dąbki (eastern CE, Poland [
64]), sites in the Dalarna district (FS, Sweden [
65]), several locations in south-eastern Finland (in particular, Outokumpu Sätös [
66]), and even more in the north [
67], or the German sites of Heidmoor (western CE [
68]) and Hüde I (western CE [
69]).
In all three areas investigated, almost all beaver remains originate from settlement contexts, regardless of the period from which the finds come. Only 4% (western CE), 3% (FS), and less than 1% (eastern CE) of the beaver records derive from graves. However, the large Mesolithic cemetery of Oleniy Ostrov in Karelia mentioned above provides 1201 beaver tooth pendants as grave goods [
70]. At that site, beaver teeth were associated more commonly with female graves.
3.4. Elk
The archaeozoological record shows that during the Mesolithic age, but also in later periods, elks were a key game species for humans in FS, eastern CE, and also in north-western CE. In eastern CE, elk achieve almost universally high levels of constancy (
Table 2;
Figure 3a). They are highest in the Mesolithic at 60%, then drop to a low of 14% in the Bronze Age, before rising again almost continuously. At sites from the Middle Ages, elk remains are found again with a constancy of 50–60%. FS follows this development with a time lag. Here, the constancy is also very high in the Stone Age but reaches its maximum value (52%) only in the Bronze Age. Additionally, almost half of the mammal bones found at Mesolithic sites in FS and eastern CE derive from elk (
Table 1 and
Table 2;
Figure 3b). This period also includes special depositions of parts of elk bodies in the shallow water of small lakes after the animals had been slaughtered. Such depositions are known from several sites in Denmark [
71,
72,
73,
74] and Sweden [
75,
76,
77]. At the same time, elks are frequently depicted in Scandinavian rock art—often it is the dominant motif—and objects of fine art all around the Baltic Sea [
78,
79,
80,
81]. The spread of the elk motif in prehistoric art also reached eastern CE and the north of western CE [
82].
In FS, this ‘era of the elk’ is followed by a massive decline in constancy values to a minimum value of only 1% in the centuries around the birth of Christ. From the Iron Age and the Roman period, there were nearly no records of elk at archaeological sites in FS: in the Iron Age, the species was found in one grave and two settlement contexts only—all three in Finland. In the late Middle Ages, elk remains in FS are again found much more constantly (31%). The relative frequency of elk remains in FS in comparison to other wild mammal species is similar to the constancy (
Figure 3b). The only significant deviation is the low proportions of elk in the NISP. In eastern CE, the proportion of elk remains in the archaeological record decreases steadily from the Mesolithic (49%) to the Roman period (2%). In the Middle Ages, the values rise again to the level of the Iron Age (by 11%); however, later the size of the population declines in eastern CE. In late medieval times, the distribution of the species was restricted to Latvia and more eastern and northern regions. The situation changed after the beginning of 19th century CE and, by 1810, the distribution of elk in Europe increased greatly and encompasses today’s total FS and the complete area from the Gulf of Finland to the Oder River [
83].
In western CE, we see a completely different situation. Here, the constancy with which elk remains are recorded at archaeological sites is low since the Neolithic age never reaches 10%, and is only less than a third (27%) during the Mesolithic age (
Table 3;
Figure 3a). The proportions of elk remains are even lower and never reach over 3% (
Table 3;
Figure 3b). An earlier study has shown that elks in Western Germany up to the river Elbe have only been occurring in very low population densities since the Neolithic age [
84]. From the Bronze Age to medieval times, we see here two distinct populations, the first in the triangle of the Harz mountains in the north, the Thuringian Forest in the southwest, and the Ore mountains in the southeast, and the second being a population along the Elbe River, which likely had a fairly high population density. Both populations became extinct at 1200 CE at the latest. Following the archaeological records, in the eastern part of western CE, the elk population was much larger; here, the species is more often identified at archaeological sites from the Stone Age up to early medieval times. But again, also in this area, the youngest record dates from approximately 1200 CE. In the north of western CE, at Mesolithic sites, elks can be recorded with high constancy, but after the ‘neolithisation’ about 4000 BCE, records become very rare [
84]. Only in the hinterland of the southern coast of the Baltic Sea, do the species survive much longer. However, as in the other parts of western CE, it was last mentioned in 13th century CE [
85].