Pattern of Diversity and Prediction of Suitable Areas of Grasshoppers from the Qinghai–Tibet Plateau in China (Orthoptera: Acridoidea)
Key Laboratory of Zoological Systematics and Application of Hebei Province, College of Life Sciences, Hebei University, Baoding 071002, China
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Authors to whom correspondence should be addressed.
†
These authors contributed equally to this work.
Insects 2025, 16(2), 191; https://doi.org/10.3390/insects16020191
Submission received: 13 January 2025
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Revised: 31 January 2025
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Accepted: 3 February 2025
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Published: 10 February 2025
(This article belongs to the Section Insect Ecology, Diversity and Conservation)
Simple Summary
This study explores the diversity and distribution of different grasshopper species across the Qinghai–Tibet Plateau, a region renowned for its rich diversity. Through the compilation and analysis of grasshopper specimens, the literature, and geographic distribution data, we have created a species database that details the diversity and distribution of 390 species. The research highlights that the Tibetan Plateau’s endemic species are predominantly found in the Bayan Har Mountains and the Hengduan Mountains. We also predict changes in suitable habitats for grasshoppers under future climate scenarios, providing insights for pest management in agricultural landscapes and forests, as well as for biodiversity conservation.
Abstract
The Qinghai–Tibet Plateau is recognized as a biodiversity hotspot, with a wide variety of grasshopper species, including several endemic to the region, which play significant roles in both agricultural and forestry ecosystems. The purpose of this study was to analyze the species diversity and distribution pattern of grasshoppers on the Qinghai–Tibet Plateau. A comprehensive database comprising 390 grasshopper species was established through specimen collection, a literature review, and a geographical distribution data analysis. Diversity analysis showed that the diversity of species under the five vegetation types was relatively average. However, the alpine cold vegetation of Qinghai–Tibet and subtropical evergreen broad-leaved forest still showed a relatively high Shannon index and Simpson index. Grasshopper species are mainly concentrated in the eastern and southern parts of the Qinghai–Tibet Plateau. The richness pattern showed that grasshopper species diversity was particularly high in certain mountain areas, with Bayankala Mountain and Hengduan Mountain being endemic hotspots. The MaxEnt models were used to assess the potential habitats for four dominant genera of grasshoppers under projected climate change scenarios for 2050 and 2070. Altitude was the factor affecting the distribution of Locusta, Chorthippus, and Kingdonella, while precipitation and temperature were the factors affecting the distribution of Leuconemacris. These findings improve our understanding of the distribution patterns of different grasshopper species across various habitat types on the Qinghai–Tibet Plateau and provide valuable insights for developing targeted ecological protection strategies in response to environmental changes.
1. Introduction
The Qinghai–Tibet Plateau, renowned as the ‘roof of the world’ and the ‘third pole’ is a highly biodiverse region with significant scientific research value. It also serves as an early warning zone and a sensitive area for global climate change. Issues such as glacier retreat, permafrost degradation, and a sharp decline in biodiversity have become increasingly prominent. These phenomena are warning us that human communities and natural ecosystems may face additional significant risks [1,2,3,4,5]. With a total area of 2,581,300 km2, the Qinghai–Tibet Plateau in China stands among the world’s biodiversity hotspots [6]. Due to the high altitude, diverse climatic conditions, and complex topography, the Qinghai–Tibet plateau provides a diverse range of habitats for insects. Many rare and primitive grasshopper species are exclusively distributed on the Qinghai–Tibet Plateau [7]. Grasshoppers refer to Orthoptera Acridoidea insects that exhibit rich species diversity worldwide—approximately 8270 species and 1244 subspecies [8]—occupying various niches within nature. They are more sensitive to changes in their habitats and play crucial roles as indicators for fluctuations in natural landscapes or habitats [9]. Currently, studies on grasshoppers within the Qinghai–Tibet Plateau primarily focus on compiling species lists and controlling a few harmful species, while investigations into their diversity and geographical distribution patterns remain scarce.
Grassland is the main ecological type on the Qinghai–Tibet Plateau, covering over 60% of its total area [10]. Grasshoppers are the typical phytophagous insects that harm grassland, and their diversity and distribution are closely related to the health and sustainable development of the ecosystem [11]. Relevant studies have shown that grasshopper outbreaks are closely related to climatic conditions [12]. Global warming increases the frequency of grasshopper outbreaks, leading to detrimental effects on agricultural production [13,14]. For example, warming oceans bring more cyclones that create more favorable breeding conditions for grasshoppers, leading to a 2020 plague on the East African continent not seen in decades [15]. Agricultural planting zones on the Qinghai–Tibet Plateau are mostly located in river valleys, which are the main zones for grasshopper breeding and activities [16]. The maximum entropy model (MaxEnt) was first introduced into the prediction of species distribution area by Steven et al. in 2004 [17]. With its strong prediction ability, MaxEnt is one of the commonly used prediction models for potential geographic distribution [18]. According to the Fifth Assessment Report of the United Nations Intergovernmental Panel on Climate Change, the global climate will continue to warm, and, compared with 1986–2005, by 2100, the global climate will increase by 0.3 °C to 4.5 °C [19]. Global warming may lead to changes in the habitat areas of species. In the current context, some researchers have used this model to investigate the potential spatial distribution pattern of grasshoppers in various regions [20,21,22], but no studies have been conducted to predict the suitable grasshopper areas in the Qinghai–Tibet Plateau under climate change scenarios based on the MaxEnt model.
The Tibetan Plateau is a biodiversity hotspot of global significance, home to a wealth of grasshopper species. In this study, the species diversity, species richness pattern, and endemic distribution pattern of grasshopper in the Qinghai–Tibet Plateau were analyzed by using the information on grasshopper species distribution recorded in specimens and the literature. Then, we extracted two widely distributed genera from the Qinghai–Tibet Plateau, i.e., Locusta and Chorthippus, and two endemic genera, i.e., Leuconemacris and Kingdonella, and predicted their current and future suitable habitats (2050, 2070). The aim of our study was as follows: (1) to reveal the species diversity and distribution pattern of grasshoppers on the Qinghai–Tibet Plateau; (2) to identify the endemic distribution area of grasshoppers on the Qinghai–Tibet Plateau; and (3) to predict the habitat of grasshoppers on the Qinghai–Tibet Plateau. In addition, to assess the potential impacts of future climate change on this habitat, we conducted an analysis of the biogeographical characteristics of grasshoppers and their interactions with the environment in this region. The analysis above provides a theoretical foundation and practical guidance for developing effective grasshopper management strategies and conserving biodiversity.
2. Materials and Methods
2.1. Determination of the Research Area
Located in South–Central Asia, the Qinghai–Tibet Plateau is the largest plateau in China and the highest plateau in the world. It covers six provinces in southwest China: Tibet, Qinghai, Xinjiang, Gansu, Sichuan, and Yunnan. Lóczy [23] described the boundary of the Qinghai–Tibet Plateau, but the extent and specific boundary of the Tibetan Plateau have been the focus of controversy among many scholars [24]. The research area of this study was based on the 2021 Tibetan Plateau Range dataset, the results of the second comprehensive Scientific expedition to the Tibetan Plateau [25], which was downloaded from the Data Center for Resources and Environmental Sciences, Chinese Academy of Sciences (http://www.resdc.cn, accessed on 20 August 2024). The clipping function of ArcGIS 10.8 was used to convert the dataset into the Qinghai–Tibet Plateau region within China (Figure 1).
2.2. Data Collection and Organization
The inspected specimen data collected in the field were used for species diversity analysis under different habitat types. The data of species geographic distribution were used to analyze the distribution pattern of abundance and endemic species and to predict the suitable areas (Table 1, Appendix F). The geographical distribution data were partly obtained from global biodiversity information websites (https://www.gbif.org/, accessed on 20 August 2024) and Orthoptera Species File (Version 5.0/5.0) records (http://orthoptera.archive.speciesfile.org/Common/editTaxon/SearchForTaxon.aspx, accessed on 20 August 2024), and partly from specimens stored in scientific research institutions in China. The samples are collected in the field from the Laboratory of Systematics and Molecular Adaptation Evolution of Hebei University, the Museum of Hebei University, and the Northwest Plateau Ecology Research Institute. The species information and geographical distribution information were verified and corrected, and the distribution information containing only the name of the collection site was also included in the geographical distribution database of Acridoidea on the Qinghai–Tibet Plateau after calibration. Data on environmental variables for projections of suitable areas are available at 30 arc seconds (1 km2) from the World Climate website (http://www.worldclim.org, accessed on 20 August 2024). The vegetation normalization index was downloaded from the National Data Center for Ecological Sciences (http://www.nesdc.org.cn/, accessed on 20 August 2024) [26], with a resolution of 30 m. Potential evaporation and actual evaporation are downloaded at 30 arc seconds (1 km2) from the CGIAR website (http://www.cgiar.org/, accessed on 20 August 2024).
2.3. Species Diversity Analysis
The iNEXT program package in R 4.2.3 [27] was used to construct rarefaction curves under different habitat types, aiming to assess the adequacy of specimen collection. Vegetation type data were obtained from the Data Center for Resources and Environmental Sciences, Chinese Academy of Sciences (http://www.resdc.cn, accessed on 20 August 2024). According to the Vegetation Regionalization System of China, the Qinghai–Tibet Plateau was categorized into six habitat types: warm temperate evergreen broad-leaved forest, subtropical evergreen broad-leaved forest, tropical monsoon rainforest area, temperate grassland area, temperate desert area, and alpine vegetation area of Qinghai–Tibet Plateau [28]. The following diversity indices were selected for analysis using Past 4.1.3 software: Shannon–Wiener Index (H), Pielou Evenness Index (J), Margalef Richness Index (R), Simpson Index (D), and Berger–Parker Dominance Index (W). The Shannon Index is commonly employed to measure species diversity within an ecosystem by considering both species richness and evenness. The Pielou Evenness Index is a variant of the Shannon Index that quantifies the uniformity degree of species in a community. The Margalef Richness Index solely considers species count as a measure of species richness in a community. Like the Shannon Index, which measures species diversity within ecosystems, Simpson’s Diversity Index places greater emphasis on the abundance of common species. Lastly, the Berger–Parker Dominance Index measures the relative importance of the most dominant species in a community [29].
2.4. Analysis of Species Richness and Endemic Distribution Pattern
The entire Qinghai–Tibet Plateau region was divided into 0.5° × 0.5° grid geographic operational units. Following the principle of retaining only one piece of geographic distribution information for each species within a unit, the geographic distribution data in the database were manually sorted and screened. By using the ArcGIS 10.8 geographic operating system, we acquired patterns of species richness and endemic distribution for grasshoppers to determine their endemic distribution center on the Qinghai–Tibet Plateau. Two methods were used to identify endemic distribution areas at a 1° × 1° grid resolution: the unweighted average method and the endemic reductive analysis method. When using the unweighted mean group method to identify these areas based on data from endemic geographical records, a data matrix was established with geographic operational units as columns and species as rows. A value of “1” indicated presence while “0” indicated absence of a particular species in each unit. This matrix was then imported into Past 4.1.3 for cluster analysis [30] using the Jaccard similarity coefficient [31,32], with Bootstrap resampled 1000 times for testing purposes. According to Morrone’s definition [33] of an endemic distribution area, regions containing two or more identical endemic species in consecutive branches were considered. Only those with Bootstrap values greater than 60 were identified as endemic distribution areas.
To prepare a matrix for endemic reduction analysis, we selected a geographical operating unit with no species distribution to serve as an outgroup and named it “A0”. The characteristic reduction analysis was carried out in TNT1.1 software. Use new technical algorithms that have significant advantages in handling complex and large amounts of data [34]. The number of maximum trees was set to 1000, initial addseqs was set to 3, and the rest of the parameters were set to default. The optimal tree was searched using sectorial search and tree fusing tools. All generated parsimony trees were combined into a strict consensus tree. Branches with a bootstrap value of 50 or greater and that are geographically contiguous were selected as candidate branches for endemic distribution areas [35]. When two or more identical endemic species are present within these branches, the corresponding units are identified as endemic distribution areas.
2.5. Prediction of Suitable Areas
The geographical distribution data of the most widely distributed and most harmful grasshopper Genera Locusta and Chorthippus, as well as the endemic grasshopper genera Leuconemacris and Kingdonella, were extracted from the previously established geographical distribution database of grasshoppers on the Qinghai–Tibet Plateau. To reduce the distribution superposition and sampling bias, the overlapping data were discarded, and the systematic sampling method was applied to random sampling of species distribution information [36]. The environmental variables were selected as modern bioclimate factors, wind speed, solar radiation, actual evaporation, potential evaporation, vegetation normalization index, and altitude. Bioclimatic factors in the future period reduce the uncertainty between models by taking the mean value of MIROC-ESM and CCSM4 models [37]. Referring to previous studies, this paper selected bioclimatic factors under the scenario with a representative concentration path of 8.5 [38].
The software ENMTools v.1.3 was used to perform a Pearson correlation test to assess collinearity among all environmental variables [39]. When the correlation between two or more variables exceeded 0.85 (|r| > 0.85) [40,41,42], environmental factors with low contribution rates were eliminated using the Jackknife method. Relevant environmental variables with a cumulative contribution rate of 90% were then considered as the final set of environmental variables [43].
The selected species distribution data and modern environmental variable data were imported into MaxEnt version 3.4.3 software. Seventy-five percent of the distribution data were used as the training set, and 25% were used as the test set to construct the MaxEnt model for predicting species distributions. To prevent overfitting of the model, the combination of linear, quadratic, and hinge function feature types is selected [44,45,46]. The same parameter Settings were used to predict grasshopper habitats in 2050 and 2070, respectively. The prediction results were reclassified by ArcGIS 10.8, and the suitable areas were divided into four categories according to the probability range of species existence: non-suitable areas (0–0.2), low suitable areas (0.2–0.4), medium suitable areas (0.4–0.6), and high suitable areas (0.6–1) [2].
3. Results
3.1. Analysis of Species Composition
In this study, we compiled a species list of Acridoidea in the Qinghai–Tibet Plateau by consulting specimen records, records from the literature, and database records. The list includes 390 species from 121 genera and 4 families (see Appendix F). A total of 10,255 grasshopper specimens belonging to 4 families, 71 genera, and 168 species were included for the analysis to reveal the diversity structure of grasshopper communities in the region (see Appendix A).
The findings indicated that Acrididae accounted for a significant proportion (91.54%) of grasshoppers, highlighting its dominant position within the grasshopper community on the Qinghai–Tibet Plateau (Table 2). In terms of genus hierarchy, Chorthippus (Acrididae) emerged as an extremely dominant genus while Indopodisma (Acrididae), Kingdonella (Acrididae), and Aeropus (Acrididae) were identified as common genera. No specific dominant species were observed. Instead, there was a substantial presence of both common and rare species. This suggests high species diversity. It also indicates a relatively even distribution within the grasshopper community on the Qinghai–Tibet Plateau. Common species such as Indopodisma kingdoni (Acrididae), Kingdonella bicollina (Acrididae), and Locusta migratoria migratorioides (Acrididae) played crucial ecological roles within this community on the Tibetan Plateau, whereas rare species like Catantops pinguis (Acrididae), Promesosternus vittatus (Acrididae), and Assamacris curticerca (Acrididae) added complexity to enhance overall diversity within grasshopper communities on this plateau (Table 2).
3.2. Analysis of Species Diversity in Different Habitat Types
The results of the species sampling adequacy test under different habitat types indicated that samples from all habitat types except for the warm temperate evergreen broad-leaved forest were deemed adequate (Figure 2). Therefore, the warm temperate evergreen broad-leaved forest habitat type was excluded, and the species diversity of grasshoppers in the remaining five habitat types was subsequently analyzed.
The diversity analysis results showed that the species and individual number were the most in the subtropical evergreen broad-leaved forest and the alpine vegetation area of the Qinghai–Tibet Plateau, while the species and individual number were the least in the temperate desert area (Figure 3a,b). The species richness index of subtropical evergreen broad-leaved forests was the highest (Figure 3c). The evenness index was the highest in temperate grassland areas, and the distribution of grasshopper species in this habitat was relatively well distributed, with no obvious dominant species (Figure 3f). The Shannon Index and the Simpson Index were the highest in the alpine vegetation area of the Qinghai–Tibet Plateau (Figure 3d,e). The Shannon Index considered species richness and evenness, and the Simpson Index considered species richness and relative abundance, indicating that the overall grasshopper diversity in this region was high. In summary, the alpine vegetation area and the subtropical evergreen broad-leaved forest on the Qinghai–Tibet Plateau show significantly high species diversity. In contrast, temperate desert areas have the lowest species diversity. The spatial distribution of grasshopper species was the most uniform in temperate grassland areas.
3.3. Distribution Pattern Analysis
Based on the species distribution data of Acridoidea on the Qinghai–Tibet Plateau, a species distribution database of Acridoidea on the Qinghai–Tibet Plateau was constructed (see Appendix A for details). A total of 1540 distribution records of 390 species of 4 families were recorded, including 661 distribution records of 227 endemic species on the Qinghai–Tibet Plateau. We constructed the species richness pattern of Acidoidea on a 0.5° grid scale. The distribution of grasshopper species is mainly concentrated in the eastern and southern parts of the Qinghai–Tibet Plateau (94–104° E, 26–34° N), with the highest species richness in the eastern part of Qinghai Province, the Hengduan Mountains region, and the Metuo and Chaiyu regions of Tibet (Figure 4).
The distribution information of 227 endemic species and 661 geographic coordinates on the Tibetan Plateau was imported into ArcGIS 10.8 reoperating system to generate the distribution points of endemic species on the Tibetan Plateau, and the number of unique species in each 1° unit was counted to generate the distribution pattern of endemic species on the Tibetan Plateau (Figure 5). Based on the unweighted average method, five endemic areas were identified by consensus: the Himalayas (F8, F9), the Bayan Kala Mountains (D12, D13), the Jinsha River coast (E14, F14), the Western Sichuan (E15, E16), and the Hengduan Mountains (G14, G15) (Figure 6). Based on endemic reduction analysis, three endemic distribution areas were identified by consensus, namely, the Jinsha River coast (E14, F14), Hengduan Mountain Range (G14, G15), and Hengduan Mountain Range region in Western Sichuan (F15, E15, E16). The identified endemic species are shown in the attached table (see Appendix B and Appendix C).
3.4. Prediction of Suitable Areas
In predicting the potential suitable habitats of four genera in the Qinghai–Tibet Plateau, relevant environmental factors were selected (see Appendix D). The MaxEnt model was employed to predict the potential distribution areas of four genera of Acrididae on the Qinghai–Tibet Plateau under three scenarios: current conditions (a), the RCP8.5 scenario for 2050 (b), and the RCP8.5 scenario for 2070 (c) (Figure 7). The AUC values exceeded 0.95, indicating a high degree of reliability in predictive performance. The results demonstrated varying degrees of influence from different environmental factors on the distribution patterns of each grasshopper genera. Notably, elevation was found to significantly impact the distribution of the Locusta. In contrast, annual precipitation, precipitation during the driest month, and seasonal temperature emerged as determinants affecting the distribution pattern of Leuconemacris. The distribution of Kingdonella was influenced by both elevation and wind speed, while that of Chorthippus exhibited a correlation with elevation and actual evaporation.
A comparative analysis between current and future suitable distribution areas revealed that all types within the Locusta displayed varying degrees of increase in suitable habitats alongside a trend toward expansion into lower elevations (see Appendix E). Although suitable regions for Leuconemacris within the Qinghai–Tibet Plateau are limited without an evident outbreak risk, Western Sichuan remains an area warranting close monitoring. The anticipated future suitable range for Kingdonella is expected to align closely with its current habitat preferences primarily located south of Bayanhar Mountains. Furthermore, evidence suggests a slight expansion in overall suitable areas for Chorthippus.
4. Discussion
This study provides a comprehensive analysis of the species composition of grasshoppers in the Qinghai–Tibet Plateau, revealing the diverse structure of grasshopper communities in this region. The findings reveal that, at the family level, Acrididae constitutes 95.84% of the individual grasshoppers. This finding aligns with previous observations regarding the dominant status of Acrididae in studies conducted on grasshopper fauna in Qinghai, China [47]. The large proportion of Acrididae in the Qinghai–Tibet Plateau can be attributed to the good adaptability of this family to high altitude, drought, and low temperature [48]. In contrast, Pamphagidae, Pyrgomorphidae, and Dericorythidae account for less than 5% of the species. In this study, Chorthippus was identified as an extremely dominant genus. Chorthippus species showed a good survival advantage in the study area, which was related to its excellent reproductive ability [49,50]. Genera such as Indopodisma, Kingdonella, and Aeropus are recognized as common genera. It must be pointed out here that we did not find dominant species in the Tibetan Plateau, which is different from what other scholars have found in investigating other ecosystem types [51,52,53]. Grasshoppers on the Qinghai–Tibet Plateau are mainly composed of many common and rare species. Rare species include species such as Catantops pinguis, Assamacris curticerca, and Promesosternus vittatus.
The formation of species diversity must be the result of the interaction of many environmental factors [54], and factors affecting species diversity vary in different regions [55]. This study reveals differences in the distribution of Acridoidea diversity in different habitats on the Qinghai–Tibet Plateau. The number of species and individuals in subtropical evergreen broad-leaved forests and alpine vegetation on the Qinghai–Tibet Plateau is the largest, which is related to the complexity of regional habitats and the richness of resources. The subtropical evergreen broad-leaved forest area provides diverse microhabitats and abundant food resources, creating favorable conditions for the survival and reproduction of Acridoidea [56,57]. Although the alpine vegetation area of the Qinghai–Tibet Plateau is restricted by temperature factors, it may promote the maintenance and enrichment of species diversity because it is less subjected to human disturbance [58,59]. Arid, barren, and highly fluctuating temperatures restrict the reproduction and survival of the Acridoidea species, which is the reason why temperate desert areas show the lowest species diversity, with the lowest number of species and individuals among all vegetation types [60].
The results of the distribution pattern indicate that the eastern and southern regions of the Qinghai–Tibet Plateau, particularly in eastern Qinghai Province, along the Hengduan Mountains, and in the Medog and Zayü areas of Tibet, are identified as hotspots for grasshopper species richness. This finding aligns with previous research on biodiversity within the Qinghai–Tibet Plateau [61]. Through unweighted average grouping methods and endemicity analysis, five endemic distribution areas have been identified that may represent significant geographical isolation zones crucial for species formation and divergence. Geographical isolation is recognized as a vital driver for speciation and evolution, especially under complex topographic features and varied climate conditions characteristic of the Qinghai–Tibet Plateau [62]. The endemic distribution areas located in the Himalayas, Bayanhar Mountain Range, Jinsha Riverbanks, Western Sichuan Province, and along the Hengduan Mountains facilitated both the formation and evolution of endemic species due to prolonged geographical isolation coupled with environmental heterogeneity [63,64].
For most environmental change scenarios, the negative impact on species richness in small hotspot areas is greater than the national average [64,65]. The Qinghai–Tibetan Plateau features a complex geological structure and exhibits sensitivity to climate change and human activities. Significant alterations in the fragile ecological balance of this region underscore the urgency of understanding and protecting its ecosystem quality [1]. Grasshoppers, as primary consumers within ecosystems, are widely utilized as biological indicators for studying the effects of global changes on these systems [66]. The MaxEnt model was used to predict potential distribution areas for four genera of grasshoppers in the Qinghai–Tibetan Plateau region for present conditions, as well as projections for 2050 and 2070. Results indicate that altitude and annual precipitation play critical roles in elucidating the relationship between grasshoppers and environmental factors. Similar findings were reported by Zhang (2024) in research conducted on grasshoppers in Ningxia Province, China [67]. Furthermore, previous large-scale studies have assessed multiple factors contributing to grasshopper population dynamics. These investigations revealed that grasshopper density is influenced by complex interactions among plant functional groups (PFG), nutrient litter (VT), and soil types (ST) [68]. Future research could enhance predictions regarding potential distribution areas of grasshoppers by incorporating additional environmental variables.
There are some limitations in this study, such as the temporal and spatial limitations in the sampling process, which failed to comprehensively observe the seasonal changes and spatial distribution patterns of grasshoppers. In the future, we need to further explore the dynamic relationship between grasshopper communities and environmental factors such as climate change, vegetation types, and human activities. Future research should combine long-term monitoring with multi-factor analysis to offer a more comprehensive assessment of grasshopper community conditions and species diversity on the Qinghai–Tibet Plateau.
5. Conclusions
The unique ecology of the Qinghai–Tibet Plateau promotes a diverse population of grasshoppers, and climate change may facilitate the expansion of their habitat. These findings underscore the imperative for targeted conservation initiatives in the identified priority areas, such as the Bayan Har Mountains and the Hengduan Mountains, to protect the region’s unique biodiversity and ecosystem services. The relative stability of the grasshopper community contrasts with the rapidly changing context of global biodiversity, highlighting the need to deepen our ecological understanding of the Qinghai–Tibet Plateau to predict and mitigate future changes to its unique ecosystems.
Author Contributions
Conceptualization, X.W. and B.B.; methodology, B.B., X.W. and Z.P.; software, B.B. and Q.Z.; validation, B.B.; formal analysis, B.B. and X.W; investigation, Z.P., Q.Z., B.B., X.W. and X.L.; resources, X.L. and D.Z.; data curation, B.B.; writing—original draft preparation, B.B. and X.L.; writing—review and editing, B.B., X.L., Q.Z. and Z.P.; visualization, B.B.; supervision, D.Z.; project administration, X.L.; and funding acquisition, X.L. All authors have read and agreed to the published version of the manuscript.
Funding
This research was funded by the Hebei Provincial Innovation Capacity Enhancement Program-Special Project for High-level Talent Team Building (No. 225A2904D) and the National Natural Science Foundation of China (No. 32070473).
Data Availability Statement
All the quantitative data analyzed are included in the Appendix A, Appendix B, Appendix C, Appendix D, Appendix E and Appendix F.
Acknowledgments
We would like to acknowledge the constructive feedback provided by the anonymous reviewers, which significantly improved the quality of this manuscript. We also wish to thank all the volunteers who participated in the study, without whom this research would not have been possible.
Conflicts of Interest
The authors declare no conflicts of interest.
Appendix A. Species List of Acridoidea in Qinghai-Tibetan Plateau
| Family | Genus | Species Name |
| Pamphagidae | Filchnerella | Filchnerella sunanensis Liu, 1982 |
| Pamphagidae | Filchnerella zhengi Huo, 1994 | |
| Pamphagidae | Filchnerella rufitibia Yin, 1984 | |
| Pamphagidae | Filchnerella kukunoris Bei-Bienko, 1984 | |
| Pamphagidae | Filchnerella beicki Ramme, 1931 | |
| Pamphagidae | Eoeotmethis | Eoeotmethis longipennis Zheng, 1985 |
| Pamphagidae | Beybienkia | Beybienkia amicus (Bey-Bienko, 1959) |
| Pyrgomorphidae | Aularches | Aularches miliaris (Linnaeus, 1958) |
| Pyrgomorphidae | Mekongiella | Mekongiella kingdoni (Uvarov, 1937) |
| Pyrgomorphidae | Mekongiella pleurodilata Yin, 1984 | |
| Pyrgomorphidae | Mekongiella rufitibia Yin, 1984 | |
| Pyrgomorphidae | Mekongiella xizangensis Yin, 1984 | |
| Pyrgomorphidae | Mekongiella wardi (Uvarov, 1937) | |
| Pyrgomorphidae | Paramekongiella | Paramekongiella zhongdianensis Huang, 1990 |
| Pyrgomorphidae | Mekongiana | Mekongiana gregoryi (Uvarov, 1925) |
| Pyrgomorphidae | Mekongiana xiangchengensis Zheng, Huang&Zhou, 2008 | |
| Pyrgomorphidae | Yunnanites | Yunnanites coriacea Uvarov, 1925 |
| Pyrgomorphidae | Yunnanites zhengi Mao&Yang, 2003 | |
| Pyrgomorphidae | Yunnanites albomargina Mao&Zheng, 1999 | |
| Pyrgomorphidae | Tagasta | Tagasta indica Bolívar, 1905 |
| Pyrgomorphidae | Atractomorpha | Atractomorpha melanostriga Bi, 1981 |
| Pyrgomorphidae | Atractomorpha himalayica Bolívar, 1905 | |
| Pyrgomorphidae | Atractomorpha sinensis Bolívar, 1905 | |
| Catantopidae | Spathosternum | Spathosternum xizangensis Yin, 1982 |
| Acrididae | Peripolus | Peripolus nepalensis Uvarov, 1942 |
| Acrididae | Calliptamus | Calliptamus italicus (Linnaeus,1758) |
| Acrididae | Calliptamus barbarous (Costa, 1836) | |
| Acrididae | Calliptamus abbreviates Ikonnikov, 1913 | |
| Acrididae | Xenocatantops | Xenocatantops brachycerus (Willemse, 1932) |
| Acrididae | Xenocatantops acanthraus Zheng, Li&Wang, 2004 | |
| Acrididae | Xenocatantops sauteri (Ramme, 1941) | |
| Acrididae | Stenocatantops | Stenocatantops splendens (Thunberg, 1815) |
| Acrididae | Stenocatantops mistshenkoi Willemse, 1968 | |
| Acrididae | Catantops | Catantops simlae Dirsh, 1956 |
| Acrididae | Catantops splendena (Thunb) | |
| Acrididae | Catantops pinguis pinguis (Stål, 1860) | |
| Acrididae | Schistocerca | Schistocerca gregaria (Forskal, 1775) |
| Acrididae | Patanga | Patanga japonica (Bolívar, 1898) |
| Acrididae | Patanga humilis Bi, 1986 | |
| Acrididae | Chondracris | Chondracris rosea brunneri Uvarov, 1924 |
| Acrididae | Choroedocus | Choroedocus robusia (Servile, 1839) |
| Acrididae | Shirakiacris | Shirakiacris yunkweiensis (Chang, 1937) |
| Acrididae | Shirakiacris brachyptera Zheng, 1983 | |
| Acrididae | Eucoptacra | Eucoptacra moiuoensis Yin, 1984 |
| Acrididae | Apalacris | Apalacris varicornis Walker, 1870 |
| Acrididae | Apalacris xizangensis Bi, 1984 | |
| Acrididae | Ecphanthacris | Ecphanthacris mirabilis Tinkham, 1940 |
| Acrididae | Assamacris | Assamacris longicercus (Huang, 1981) |
| Acrididae | Assamacris curticerca (Huang, 1981) | |
| Acrididae | Promesosternus | Promesosternus himalayicus Yin, 1984 |
| Acrididae | Promesosternus vittatus Yin, 1984 | |
| Acrididae | Habrocnemis | Habrocnemis sinensis Uvarov, 1930 |
| Acrididae | Oxya | Oxya agavisa agavisa Tsai, 1931 |
| Acrididae | Oxya chinensis (Thunberg, 1815) | |
| Acrididae | Oxya intricata (Stål, 1861) | |
| Acrididae | Oxya adentata Willemse, 1925 | |
| Acrididae | Oxya brachyptera Zheng&Huo, 1992 | |
| Acrididae | Oxya velox (Fabricius, 1787) | |
| Acrididae | Oxya japonica (Thunberg, 1824) | |
| Acrididae | Caryanda | Caryanda gyirongensis Huang, 1981 |
| Acrididae | Caryanda bidentata Zheng&Liang, 1985 | |
| Acrididae | Parapodisma | Parapodisma astris Huang, 2006 |
| Dericorythidae | Conophymacris | Conophymacris xianggelilaensis Niu&Zheng, 2009 |
| Dericorythidae | Conophymacris yunnanensis Willemse Cheng, 1977 | |
| Dericorythidae | Conophymacris jiulongensis Zheng&Shi, 2009 | |
| Dericorythidae | Conophymacris cangshanensis Zheng&Mao, 1996 | |
| Dericorythidae | Conophymacris nigrofemora Liang, 1993 | |
| Dericorythidae | Sinopodisma | Sinopodisma zheng Liang&Lin, 1995 |
| Acrididae | Curvipennis | Curvipennis wixiensis (Yin,1980) |
| Acrididae | Pedopodisma | Pedopodisma emeiensis (Yin, 1980) |
| Acrididae | Pedopodisma abaensis Yin, Zheng&Ye, 2012 | |
| Acrididae | Pseudozubovskia | Pseudozubovskia xizangensis Zheng, Lin, Zhang&Zeng, 2014 |
| Acrididae | Xiangelilacris | Xiangelilacris zhongdianensis Zheng, Huang&Zhou, 2008 |
| Acrididae | Rectimargipodisma | Rectimargipodisma medogensis Zheng, Li&Wang, 2004 |
| Acrididae | Indopodisma | Indopodisma kingdoni (Uvarov, 1927) |
| Acrididae | Peseudoptygonotus | Peseudoptygonotus aipinus |
| Acrididae | Fruhstorferiola | Fruhstorferiola omei (Rehn, 1939) |
| Acrididae | Tylotropidius | Tylotropidius yunnanensis Zheng er Liang, 1990 |
| Acrididae | Rhinopodisma | Rhinopodisma eminifrontus (Huang, 1981) |
| Acrididae | Eokingdonella | Eokingdonella changtunica Yin, 1984 |
| Acrididae | Eokingdonella cyanecuda Zheng&Shi, 2009 | |
| Acrididae | Eokingdonella angqianensis Chen&Zheng, 2009 | |
| Acrididae | Eokingdonella gongbugyanda Zheng, Lin, Zhang&Zeng, 2014 | |
| Acrididae | Eokingdonella gentiana (Uvarvo,1939) | |
| Acrididae | Eokingdonella tibetana (Mistshenko, 1952) | |
| Acrididae | Eokingdonella burrusitibia Zheng, Dong&Xu, 2013 | |
| Acrididae | Eokingdonella kualbacki (Uvarvo,1939) | |
| Acrididae | Eokingdonella bayanharaensis (Huo, 1995) | |
| Acrididae | Eokingdonella luhuoensis Zheng&Shi, 2007 | |
| Acrididae | Kingdonella | Kingdonella qinghaiensis Zheng, 1990 |
| Acrididae | Kingdonella nigrotibia Zheng, 1990 | |
| Acrididae | Kingdonella bicollina Yin, 1984 | |
| Acrididae | Kingdonella magna Yin, 1984 | |
| Acrididae | Kingdonella pienbaensis Zheng, 1980 | |
| Acrididae | Kingdonella nigrofemora Yin, 1984 | |
| Acrididae | Kingdonella hanburyi Uvarov, 1939 | |
| Acrididae | Kingdonella parvula Yin, 1984 | |
| Acrididae | Kingdonella longiconica Yin, 1984 | |
| Acrididae | Kingdonella pictipes Uvarov, 1935 | |
| Acrididae | Kingdonella kozlovi Mistshenko, 1952 | |
| Acrididae | Kingdonella conica Yin, 1984 | |
| Acrididae | Kingdonella afurcula Yin, 1984 | |
| Acrididae | Kingdonella sxicola Uvarvo, 1939 | |
| Acrididae | Kingdonellarivuna Huang, 1981 | |
| Acrididae | Kingdonella wardi Uvarvo, 1939 | |
| Acrididae | Kingdonella conica Yin, 1984 | |
| Acrididae | Anepipodisma | Anepipodisma punctata Huang, 1984 |
| Dericorythidae | Conophymopsis | Conophymopsis linguspinus Huang, 1983 |
| Dericorythidae | Conophymopsis labrispinus Huang, 1983 | |
| Dericorythidae | Diricorys | Diricorys annulata roseipennis (Redtenbacher, 1889) |
| Acrididae | Leptopternis | Leptopternis gracilis (Eversmann, 1848) |
| Acrididae | Heteropternis | Heteropternis respondens (Walker, 1859) |
| Acrididae | Heteropternis robusta Bey-Bienko, 1951 | |
| Acrididae | Heteropternis motuoensis Yin, 1984 | |
| Acrididae | Heteropternis qinghaiensis Wang&Zheng, 1992 | |
| Acrididae | Heteropternis micronus Huang, 1981 | |
| Acrididae | Longipternis | Longipternis chayuensis (Yin, 1984) |
| Acrididae | Mecostethus | Mecostethus grossus (Linnaeus, 1758) |
| Acrididae | Ochyracris | Ochyracris rufotibialis Zheng, 1991 |
| Acrididae | Epacromius | Epacromius coerulipes (Ivanov, 1888) |
| Acrididae | Epacromius tergestinus tergestinus (Megerle von Mühlfeld, 1825) | |
| Acrididae | Epacromius tergestinus extimus Bey-Bienko, 1951 | |
| Acrididae | Aiolopus | Aiolopus thalassinus (Fabricius, 1798) |
| Acrididae | Aiolopus tamulus (Fabricius, 1798) | |
| Acrididae | Aiolopus markamensis Yin, 1984 | |
| Acrididae | Oedaleus | Oedaleus infernalis Saussure, 1884 |
| Acrididae | Oedaleus asiaaicus Bey-Bienko, 1941 | |
| Acrididae | Oedaleus manjius Chang, 1939 | |
| Acrididae | Acrotylus | Acrotylus longipes subfasciatus Bey-Bienko, 1984 |
| Acrididae | Aurilobulus | Aurilobulus splendens Yin, 1979 |
| Acrididae | Cyanicaudata | Cyanicaudata annulicornea Yin, 1979 |
| Acrididae | Celes | Celes skalozubovi Adelung, 1906 |
| Acrididae | Celes akitanus (Shiraki, 1910) | |
| Acrididae | Sphingonotu | Sphingonotus kueideensis Yin, 1984 |
| Acrididae | Sphingonotus qinghaiensis Yin, 1984 | |
| Acrididae | Sphingonotus yunnaneus Uvarov,1925 | |
| Acrididae | Sphingonotus (Sphingonotus) tipicus Zheng&Hang, 1974 | |
| Acrididae | Sphingonotus halophilus Bey-Bienko, 1929 | |
| Acrididae | Sphingonotus tzaidamicus Mistshenko,1937 | |
| Acrididae | Sphingonotus caerulistriatus Zheng&Ren,2007 | |
| Acrididae | Sphingonotus zadaensis Huang, 1981 | |
| Acrididae | Sphingonotus longipennis Saussure, 1884 | |
| Acrididae | Sphingonotus turcmenus Bey-Bienko, 1951 | |
| Acrididae | Tibetacris | Tibetacris changtunensis Chen, 1964 |
| Acrididae | Trilophidia annulata (Thunberg, 1815) | |
| Acrididae | Parahilethera | Parahilethera xizangensis Zheng&Ren, 2007 |
| Acrididae | Gastrimargus | Gastrimargus africanus africanus (Saussure, 1888) |
| Acrididae | Gastrimargus nubilis Uvarov, 1925 | |
| Acrididae | Gastrimargus marmoratus (Thunberg, 1815) | |
| Acrididae | Gastrimargus ommatidius Huang, 1981 | |
| Acrididae | Gastrimargus africanus parvulus Sjöstedt, 1928 | |
| Acrididae | Locusta | Locusta migratoria tibetensis Chen, 1963 |
| Acrididae | Locusta migratoria manilensis (Meyen, 1835) | |
| Acrididae | Locusta migratoria migratoria (Linnaeus, 1758) | |
| Acrididae | Pternoscirta | Pternoscirta saueri (Karny, 1915) |
| Acrididae | Pternoscirta pulchripes Uvarov,1925 | |
| Acrididae | Pternoscirta longipennis Xia, 1981 | |
| Acrididae | Bryodemella | Bryodemella xizangensis Yin, 1984 |
| Acrididae | Bryodemella tuberculatum dilutum (Stoll,1813) | |
| Acrididae | Bryodemella diamesum (Bey-Bienko, 1930) | |
| Acrididae | Bryodemella (Bryodemella) rufifemura Zhang, Zhi&Zhang, 2018 | |
| Acrididae | Bryodemella (Bryodemella) nigrifemura Yin&Wang, 2005 | |
| Acrididae | Bryodemella holdereri holdereri (Krauss, 1901) | |
| Acrididae | Bryodema luctuosum luctuosum (Stoll, 1813) | |
| Acrididae | Bryodema miramae miramae Bey-Bienko, 1930 | |
| Acrididae | Bryodema uvarovi (Bey-Bienko, 1930) | |
| Acrididae | Bryodema miramae elegantulum Bey-Bienko, 1930 | |
| Acrididae | Bryodema luctuosum inda Saussure, 1884 | |
| Acrididae | Bryodema hyalinala Zheng&Zhang, 1981 | |
| Acrididae | Bryodema pseudohyalinala Zheng, Lin&Zhang, 2012 | |
| Acrididae | Bryodema kangmarensis Zheng, Lin&Zhang, 2012 | |
| Acrididae | Bryodema nigristria Zheng&Chen, 2001 | |
| Acrididae | Bryodema gebleri gebleri (Fischer von Waldheim, 1836) | |
| Acrididae | Bryodema gansuensis Zheng, 1985 | |
| Acrididae | Bryodema qilianshanicum Lian&Zheng,1984 | |
| Acrididae | Angaracris | Angaracris barabensis (Pallas, 1773) |
| Acrididae | Angaracris rhodopa (Fischer von Waldheim, 1836) | |
| Acrididae | Angaracris morulimarginis Huang, 1981 | |
| Acrididae | Angaracris acrohylinna Bi, 1986 | |
| Acrididae | Angaracris nigripennis Lian&Zheng, 1984 | |
| Acrididae | Uvaroviola | Uvaroviola multispinosa Bey-Bienko, 1930 |
| Acrididae | Ceracris | Ceracris nigricornis nigricornis Walker, 1870 |
| Acrididae | Ceracris xizangensis xizangensis Liu, 1981 | |
| Acrididae | Ceracris versicolor (Brunner von Wattenwyl, 1893) | |
| Acrididae | Ceracris fasciata (Brunner-Wattenwyl, 1893) | |
| Acrididae | Ceracrisoides | Ceracrisoides shannanensis Bi&Xia, 1987 |
| Acrididae | Anaptygus | Anaptygus qinghaiensis Yin, 1984 |
| Acrididae | Anaptygus yueliangwan Mao, Ren&Ou, 2011 | |
| Acrididae | Anaptygus yulongensis Wang, Zheng&Lian, 2005 | |
| Acrididae | Anaptygus furculus Huo&Zheng | |
| Acrididae | Anaptygus longipennis Wang, Zheng&Lian, 2004 | |
| Acrididae | Anaptygus uvarovi (Chang, 1937) | |
| Acrididae | Arcyptera | Arcyptera coreana Shiraki, 1930 |
| Acrididae | Leuconemacris | Leuconemacris litangensis (Yin, 1983) |
| Acrididae | Leuconemacris xizangensis (Yin, 1984) | |
| Acrididae | Leuconemacris daochengensis Zheng, 1988 | |
| Acrididae | Leuconemacris longipennis Zheng, 1988 | |
| Acrididae | Leuconemacris breviptera (Yin, 1983) | |
| Acrididae | Leuconemacris xiangchengensis Zheng, 1988 | |
| Acrididae | Leuconemacris shalulishanensis Zheng, 1988 | |
| Acrididae | Leuconemacris hengduanshanensis Zheng, 1988 | |
| Acrididae | Leuconemacris xinlongensis Mao&Huang, 2023 | |
| Acrididae | Leuconemacris acuminata Mao&Huang, 2023 | |
| Acrididae | Leuconemacris muliensis Mao&Huang 2023 | |
| Acrididae | Pararcyptera | Pararcyptera microptera Ikonnikov, 1911 |
| Acrididae | Dnopherula | Dnopherula brevicornis (Bi&Xia, 1987) |
| Acrididae | Dnopherula taeniatus (Bolívar, 1902) | |
| Acrididae | Dnopherula xizangensis (Zheng&Liping, 1986) | |
| Acrididae | Dnopherula yuanmowensis (Zheng, 1977) | |
| Acrididae | Dnopherula neovittatus Otte, 1997 | |
| Acrididae | Dnopherula sinensis (Uvarov, 1925) | |
| Acrididae | Eremippus | Eremippus persicus Uvarov, 19291 |
| Acrididae | Eremippus simplex (Eversmann, 1859) | |
| Acrididae | Eremippus heimahoensis Zheng&Hang, 1974 | |
| Acrididae | Podismomorpha | Podismomorpha gibba Lian&Zheng, 1984 |
| Acrididae | Squamopenna | Squamopenna gansuensis Lian&Zheng, 1984 |
| Acrididae | Euchorthippus | Euchorthippus unicolor (Ikonnikov, 1913) |
| Acrididae | Euchorthippus weichowensis Chang, 1937 | |
| Acrididae | Euchorthippus flexucarinatus Bi e Xia, 1987 | |
| Acrididae | Euchorthippus cheui Hsia, 1965 | |
| Acrididae | Euchorthippus yungningensis Cheng&Chiu, 1965 | |
| Acrididae | Eclipophleps | Eclipophleps xizangensis Liu, 1980 |
| Acrididae | Kangacris | Kangacris rufipes Yin, 1983 |
| Acrididae | Macrokangacris | Macrokangacris luteoarmilla Yin, 1983 |
| Acrididae | Kangacrisoides | Kangacrisoides jiangdaensis Zheng, Dong, Bai&Xu, 2013 |
| Acrididae | Chorthippus | Chorthippus (Altichorthippus) intermedius (Bey-Bienko, 1926) |
| Acrididae | Chorthippus qingzangensis Yin, 1984 | |
| Acrididae | Chorthippus (Altichorthippus) latisulcus (Zheng&He, 1995) | |
| Acrididae | Chorthippus (Chorthippus) dabanshanensis Chen, Zeng&Bao, 2011 | |
| Acrididae | Chorthippus (Chorthippus) qilianshanensis Zheng&Xie, 2000 | |
| Acrididae | Chorthippus(Altichorthippus) qinghaiensis Wang&Zheng, 1992 | |
| Acrididae | Chorthippus (Altichorthippus) jishishanensis Zheng&Xie, 2000 | |
| Acrididae | Chorthippus guideensis Zheng&al, 2009 | |
| Acrididae | Chorthippus (Altichorthippus) changtunensis Yin, 1984 | |
| Acrididae | Chorthippus flexivenus (Liu, 1981) | |
| Acrididae | Chorthippus (Altichorthippus) chayuensis Yin, 1984 | |
| Acrididae | Chorthippus albonemus Zheng&Tu, 1964 | |
| Acrididae | Chorthippus intermedius (Bey-Bienko, 1926) | |
| Acrididae | Chorthippus chinensis (Tarbinsky, 1927) | |
| Acrididae | Chorthippus hsiai Cheng&Tu, 1964 | |
| Acrididae | Chorthippus longisonus (Li&Yin, 1987) | |
| Acrididae | Chorthippus (Altichorthippus) markamensis (Yin, 1984) | |
| Acrididae | Chorthippus biguttulus biguttulus (Linnaeus, 1758) | |
| Acrididae | Chorthippus chapini Chang, 1939 | |
| Acrididae | Chorthippus grahami Chang, 1937 | |
| Acrididae | Chorthippus brevipterus Yin, 1984 | |
| Acrididae | Chorthippus fallax (Zubovsky), 1899 | |
| Acrididae | Chorthippus brunneus huabeiensis Xia&Jin, 1982 | |
| Acrididae | Chorthippus maerkangensis (Zheng, 1980) | |
| Acrididae | Chorthippus xiangchengensis (Liu, 1985) | |
| Acrididae | Chorthippus louguanensis Zheng&Tu, 1964 | |
| Acrididae | Chorthippus occidentalis Xia&Jin, 1982 | |
| Acrididae | Chorthippus squamopennis Zheng, 1980 | |
| Acrididae | Chorthippus dubius (Zubovski, 1898) | |
| Acrididae | Chorthippus xiningensis Zheng&Chen, 2001 | |
| Acrididae | Chorthippus haibeiensis Zheng&Chen, 2001 | |
| Acrididae | Chorthippus helanshanensis Zheng, 1999 | |
| Acrididae | Chorthippus rufitibis (Zheng, 1989) | |
| Acrididae | Chorthippus brunneus (Thunbeng, 1815) | |
| Acrididae | Chorthippus xunhuaensis Zheng&Xie, 2000 | |
| Acrididae | Chorthippus leduensis Zheng&Xin, 1999 | |
| Acrididae | Chorthippus gongbuensis Liang&Zheng, 1991 | |
| Acrididae | Chorthippus mollis (Charpentieri, 1825) | |
| Acrididae | Chorthippus amplintersitus Liu, 1981 | |
| Acrididae | Chorthippus rubensabdomenis Liu, 1981 | |
| Acrididae | Chorthippus hemipeterus Uvarov, 1926 | |
| Acrididae | Chorthippus flavabdomenis Liu, 1981 | |
| Acrididae | Chorthippus tibetanus Uvarov, 1935 | |
| Acrididae | Chorthippus separatanus Liu, 1981 | |
| Acrididae | Chorthippus aethalinus (Zubovski, 1899) | |
| Acrididae | Chorthippus laifoveatus Xia&Jin, 1982 | |
| Acrididae | Chorthippus dichrous (Eversmann, 1859) | |
| Acrididae | Chorthippus huchengensis Xia&Jin, 1982 | |
| Acrididae | Chorthippus daitongensis Huo, 1994 | |
| Acrididae | Chorthippus fusigeniculatus Zheng&Men, 2008 | |
| Acrididae | Chorthippus xueshanensis Zheng&Mao, 1997 | |
| Acrididae | Chorthippus deqinensis Liu, 1984 | |
| Acrididae | Chorthippus nemus Liu, 1984 | |
| Acrididae | Chorthippus yajiangensis Zheng&Shi, 2007 | |
| Acrididae | Chorthippus kangdingensis Zheng&Shi, 2007 | |
| Acrididae | Chorthippus neipopennis Xia&Jin, 1982 | |
| Acrididae | Chorthippus anomopterus Liu, 1984 | |
| Acrididae | Omocestus | Omocestus cuonaensis Yin, 1984 |
| Acrididae | Omocestus gonggarensis Zheng&Chen, 1995 | |
| Acrididae | Omocestus haemorrhoidalis (Charpentier, 1825) | |
| Acrididae | Omocestus avellaeusitibia Zheng, Dong&Xu, 2013 | |
| Acrididae | Omocestus petraeus (Brisout de Barneville, 1856) | |
| Acrididae | Omocestus nyalamus Xia, 1981 | |
| Acrididae | Omocestus megaoculus Yin, 1984 | |
| Acrididae | Omocestus motuoensis Yin, 1984 | |
| Acrididae | Omocestus pinanensis Zheng&Xie, 2001 | |
| Acrididae | Omocestus qinghaihuensis Zheng&Xie, 2001 | |
| Acrididae | Omocestus hingstoni Uvarov, 1925 | |
| Acrididae | Omocestus enitor Uvarov, 1924 | |
| Acrididae | Omocestus laojunshanensis Mao&Xu, 2004 | |
| Acrididae | Nivisacris | Nivisacris zhongdianensis Liu, 1984 |
| Acrididae | Saxetophilus | Saxetophilus qinghaiensis Zheng, Chen&Li, 2012 |
| Acrididae | Saxetophilus petulans Umnov, 1930 | |
| Acrididae | Saxetophilus gansuensis Wang, Zheng&Lian, 2006 | |
| Acrididae | Oreoptygonotus | Oreoptygonotus chinghaiensis (Zheng&Hang, 1974) |
| Acrididae | Oreoptygonotus tibetanus Tarbinsky, 1927 | |
| Acrididae | Oreoptygonotus brachypterus Yin, 1984 | |
| Acrididae | Oreoptygonotus robustus Yin, 1984 | |
| Acrididae | Oreoptygonotus nigrofasciatus Yin, 1979 | |
| Acrididae | Oreoptygonotus belonocercus Liu, 1981 | |
| Acrididae | Ruganotus | Ruganotus rufipes Yin, 1979 |
| Acrididae | Ptygonotus | Ptygonotus sichuanensis Zheng, 1983 |
| Acrididae | Ptygonotus semenovi semenovi Tarbinsky, 1927 | |
| Acrididae | Ptygonotus tarbinskyi Uvarov, 1930 | |
| Acrididae | Ptygonotus gurneyi Chang, 1937 | |
| Acrididae | Ptygonotus chinghaiensis Ying, 1974 | |
| Acrididae | Ptygonotus hocashanensis Cheng&Hang, 1974 | |
| Acrididae | Ptygonotus brachypterus Yin, 1974 | |
| Acrididae | Ptygonotus gansuensis Zheng&Chang, 1994 | |
| Acrididae | Ptygonotus semenovi antennatus Mistshenko, 1951 | |
| Acrididae | Ptygonotus rangkouensis Zheng&Xu, 2009 | |
| Acrididae | Ptygonotus anurous Zheng&Xu, 2009 | |
| Acrididae | Asulconotoides | Asulconotoides sichuanensis Liu, 1984 |
| Acrididae | Asulconotoides asulcata (Zheng, 1988) | |
| Acrididae | Asulconotoides microptera (Zheng, 1988) | |
| Acrididae | Asulconotus chinghaiensis Ying, 1974 | |
| Acrididae | Asulconotus jiuzhiensis Yin, Zheng&Yin, 2012 | |
| Acrididae | Asulconotus kozlovi Mistshenko, 1981 | |
| Acrididae | Asonus | Asonus brachypterus (Ying, 1974) |
| Acrididae | Asonus amplifurculus Chen, Zheng&Zeng, 2010 |
Appendix B. Checklist of Endemic Species in the Five Distribution Areas Identified by UPGMA: Himalayas (F8, F9), Bayan Har Mountains (D12, D13), Jinsha River Basin (E14, F14), Western Sichuan (E15, E16), and Hengduan Mountains (G14, G15)
| Endemic Distribution Area | Species Name |
| Himalayas | Bryodemella xizangensis Yin, 1984 |
| Bryodema pseudohyalinala Zheng, Lin&Zhang, 2012 | |
| Bryodema kangmarensis Zheng, Lin&Zhang, 2012 | |
| Eclipophleps xizangensis Liu, 1980 | |
| Omocestus nyalamus Xia, 1981 | |
| Omocestus megaoculus Yin, 1984 | |
| Omocestus hingstoni Uvarov, 1925 | |
| Hypernephia everesti Uvarov, 1922 | |
| Dysanema irvinei Uvarov, 1925 | |
| Dysanema magna Zheng, Lin, Zhang&Zeng, 2014 | |
| Orinhippus tibetanus Uvarov, 1921 | |
| Orinhippus trisulcus Yin, 1984 | |
| Bayan Har Mountains | Eokingdonella angqianensis Chen&Zheng, 2009 |
| Eokingdonella bayanharaensis Huo, 1995 | |
| Kingdonella bicollina Yin, 1984 | |
| Kingdonella qinghaiensis Zheng, 1990 | |
| Kingdonella pienbaensis Zheng, 1980 | |
| Kingdonella pictipes Uvarov, 1935 | |
| Kingdonella kozlovi Mistshenko, 1952 | |
| Bryodemella diamesum (Bey-Bienko, 1930) | |
| Bryodema miramae elegantulum Bey-Bienko, 1930 | |
| Bryodema nigristria Zheng&Chen, 2001 | |
| Ptygonotus sichuanensis Zheng, 1983 | |
| Ptygonotus brachypterus Yin, 1974 | |
| Asulconotus chinghaiensis Ying, 1974 | |
| Asonus brachypterus (Ying, 1974) | |
| Asonus qinghaiensis Liu, 1986 | |
| Pseudoasonus yushuensis Yin, 1982 | |
| Pseudoasonus orthomarginis Zheng, Chen&Lin, 2012 | |
| Aeropedelloides altissimus Liu, 1981 | |
| Aeropedellus zadoensis Yin, 1984 | |
| Aeropedellus prominemarginis Zheng, 1981 | |
| Jinsha River Basin | Pseudoasonus baiyuensis Zheng, 1990 |
| Gonista chayuensis Yin, 1984 | |
| Tibetacris changtunensis Chen, 1964 | |
| Chorthippus (Altichorthippus) changtunensis Yin, 1984 | |
| Omocestus megaoculus Yin, 1984 | |
| Sphingonotus caerulistriatus Zheng&Ren, 2007 | |
| Chorthippus deqinensis Liu, 1984 | |
| Anepipodisma punctata Huang, 1984 | |
| Kingdonella bicollina Yin, 1984 | |
| Mekongiana gregoryi (Uvarov, 1925) | |
| Chorthippus grahami Chang, 1937 | |
| Kingdonella hanburyi Uvarov, 1939 | |
| Omocestus avellaeusitibia Zheng, Dong&Xu, 2013 | |
| Kingdonella nigrofemora Yin, 1984 | |
| Kingdonella nigrotibia Zheng, 1990 | |
| Leuconemacris hengduanshanensis Zheng, 1988 | |
| Bryodemella diamesum (Bey-Bienko, 1930) | |
| Eokingdonella burrusitibia Zheng, Dong&Xu, 2013 | |
| Kangacris rufipes Yin, 1983 | |
| Macrokangacris luteoarmilla Yin, 1983 | |
| Kangacrisoides jiangdaensis Zheng, Dong, Bai&Xu, 2013 | |
| Chorthippus chapini Chang, 1939 | |
| Chorthippus amplintersitus Liu, 1981 | |
| Chorthippus nemus Liu, 1984 | |
| Eokingdonella luhuoensis Zheng&Shi, 2007 | |
| Chorthippus (Altichorthippus) markamensis (Yin, 1984) | |
| Aiolopus markamensis Yin, 1984 | |
| Omocestus motuoensis Yin, 1984 | |
| Bryodema miramae elegantulum Bey-Bienko, 1930 | |
| Chorthippus(Altichorthippus) qinghaiensis Wang&Zheng, 1992 | |
| Asulconotoides asulcata (Zheng, 1988) | |
| Leuconemacris xizangensis (Yin, 1984) | |
| Aeropus tibetanus Uvarov, 1935 | |
| Parahilethera xizangensis Zheng&Ren, 2007 | |
| Leuconemacris xiangchengensis Zheng, 1988 | |
| Chorthippus xiangchengensis (Liu, 1985) | |
| Conophymacris xianggelilaensis Niu&Zheng, 2009 | |
| Mekongiana xiangchengensis Zheng, Huang&Zhou, 2008 | |
| Asulconotoides microptera (Zheng, 1988) | |
| Leuconemacris xinlongensis Mao&Huang, 2023 | |
| Chorthippus anomopterus Liu, 1984 | |
| Anaptygus yueliangwan Mao, Ren&Ou, 2011 | |
| Leuconemacris longipennis Zheng, 1988 | |
| Chorthippus longisonus (Li&Yin, 1987) | |
| Oreoptygonotus robustus Yin, 1984 | |
| Western Sichuan | Chorthippus (Altichorthippus) changtunensis Yin, 1984 |
| Leuconemacris breviptera (Yin, 1983) | |
| Ptygonotus gurneyi Chang, 1937 | |
| Chorthippus grahami Chang, 1937 | |
| Omocestus avellaeusitibia Zheng, Dong&Xu, 2013 | |
| Conophymacris nigrofemora Liang, 1993 | |
| Kingdonella nigrotibia Zheng, 1990 | |
| Kangacris rufipes Yin, 1983 | |
| Chorthippus occidentalis Xia&Jin, 1982 | |
| Chorthippus chapini Chang, 1939 | |
| Chorthippus kangdingensis Zheng&Shi, 2007 | |
| Pseudoasonus kangdingensis Yin, 1983 | |
| Eokingdonella cyanecuda Zheng&Shi, 2009 | |
| Leuconemacris litangensis (Yin, 1983) | |
| Chorthippus squamopennis Zheng, 1980 | |
| Chorthippus maerkangensis (Zheng, 1980) | |
| Eokingdonella luhuoensis Zheng&Shi, 2007 | |
| Ptygonotus sichuanensis Zheng, 1983 | |
| Ptygonotus anurous Zheng&Xu, 2009 | |
| Leuconemacris xinlongensis Mao&Huang, 2023 | |
| Transtympanacris xueshanensis Lian&Zheng, 1985 | |
| Chorthippus yajiangensis Zheng&Shi, 2007 | |
| Chorthippus neipopennis Xia&Jin, 1982 | |
| Habrocnemis sinensis Uvarov, 1930 | |
| Hengduan Mountains | Yunnanites albomargina Mao&Zheng, 1999 |
| Pseudoasonus baiyuensis Zheng, 1990 | |
| Caryanda bidentata Zheng&Liang, 1985 | |
| Conophymacris nigrofemora Liang, 1993 | |
| Omocestus laojunshanensis Mao&Xu, 2004 | |
| Chorthippus nemus Liu, 1984 | |
| Asulconotoides sichuanensis Liu, 1984 | |
| Curvipennis wixiensis Huang, 1984 | |
| Parapodisma astris Huang, 2006 | |
| Leuconemacris xiangchengensis Zheng, 1988 | |
| Conophymacris xianggelilaensis Niu&Zheng, 2009 | |
| Asonus parvoculus Xie & Liu, 1998 | |
| Chorthippus xueshanensis Zheng&Mao, 1997 | |
| Altichorthippus yanyuanensis Jin&Lin,1982 | |
| Anaptygus yulongensis Wang, Zheng&Lian, 2005 | |
| Paramekongiella zhongdianensis Huang, 1990 | |
| Anaptygus longipennis Wang, Zheng&Lian, 2004 | |
| Yunnanites zhengi Mao&Yang, 2003 | |
| Xiangelilacris zhongdianensis Zheng, Huang&Zhou, 2008 | |
| Nivisacris zhongdianensis Liu, 1984 | |
| Dianacris choui Yin, 1983 |
Appendix C. Checklist of Endemic Species in Three Endemic Distribution Areas Identified by the Parsimony Analysis of Endemism (PAE) Method
Jinsha River Basin (E14, F14), Hengduan Mountains (G14, G15), and Western Sichuan Hengduan Mountains Region (F15, E15, E16). The Jinsha River Basin (E14, F14) and Hengduan Mountains (G14, G15) were also identified by the Unweighted Pair Group Method with Arithmetic Mean (UPGMA) and are not repeated here.
| Endemic Distribution Area | Species Name |
| Western Sichuan Hengduan Mountains Region | Chorthippus (Altichorthippus) chayuensis Yin, 1984 |
| Chorthippus (Altichorthippus) changtunensis Yin, 1984 | |
| Leuconemacris daochengensis Zheng, 1988 | |
| Leuconemacris breviptera (Yin, 1983) | |
| Ptygonotus gurneyi Chang, 1937 | |
| Chorthippus grahami Chang, 1937 | |
| Omocestus avellaeusitibia Zheng, Dong&Xu, 2013 | |
| Conophymacris nigrofemora Liang, 1993 | |
| Kingdonella nigrotibia Zheng, 1990 | |
| Chorthippus rubensabdomenis Liu, 1981 | |
| Ochyracris rufotibialis Zheng, 1991 | |
| Kangacris rufipes Yin, 1983 | |
| Chorthippus occidentalis Xia&Jin, 1982 | |
| Leuconemacris acuminata Mao&Huang, 2023 | |
| Chorthippus chapini Chang, 1939 | |
| Conophymacris jiulongensis Zheng&Shi, 2009 | |
| Chorthippus kangdingensis Zheng&Shi, 2007 | |
| Pseudoasonus kangdingensis Yin, 1983 | |
| Eokingdonella cyanecuda Zheng&Shi, 2009 | |
| Leuconemacris litangensis (Yin, 1983) | |
| Chorthippus squamopennis Zheng, 1980 | |
| Eokingdonella luhuoensis Zheng&Shi, 2007 | |
| Chorthippus maerkangensis (Zheng, 1980) | |
| Aiolopus markamensis Yin, 1984 | |
| Leuconemacris muliensis Mao&Huang 2023 | |
| Leuconemacris shalulishanensis Zheng, 1988 | |
| Ptygonotus sichuanensis Zheng, 1983 | |
| Asulconotoides sichuanensis Liu, 1984 | |
| Ptygonotus anurous Zheng&Xu, 2009 | |
| Leuconemacris xizangensis (Yin, 1984) | |
| Leuconemacris xiangchengensis Zheng, 1988 | |
| Chorthippus xiangchengensis (Liu, 1985) | |
| Asonus parvoculus Xie & Liu, 1998 | |
| Leuconemacris xinlongensis Mao&Huang, 2023 | |
| Chorthippus xueshanensis Zheng&Mao, 1997 | |
| Chorthippus yajiangensis Zheng&Shi, 2007 | |
| Chorthippus neipopennis Xia&Jin, 1982 | |
| Transtympanacris yajiangensis Zheng, 1995 | |
| Chorthippus longisonus (Li&Yin, 1987) | |
| Habrocnemis sinensis Uvarov, 1930 |
Appendix D. Selection Results of Environmental Factors for Four Genera Within the Acridoidea
| Genus | Environmental Factors |
| Locusta | Mean Diurnal Temperature Range (MDTR) |
| Isotherm | |
| Seasonal Temperature | |
| Annual Temperature Range | |
| Annual Precipitation | |
| Precipitation in the Driest Month | |
| Seasonal Precipitation | |
| Precipitation in the Hottest Season | |
| Precipitation in the Coldest Season | |
| Elevation | |
| Wind Speed | |
| Solar Radiation | |
| Actual Evapotranspiration | |
| Potential Evapotranspiration | |
| Normalized Difference Vegetation Index (NDVI) | |
| Leuconemacris | Isothermal Temperature |
| Seasonal Temperature | |
| Annual Precipitation | |
| Precipitation in the Driest Month | |
| Precipitation in the Coldest Season | |
| Elevation | |
| Actual Evapotranspiration | |
| Solar Radiation | |
| Normalized Difference Vegetation Index (NDVI) | |
| Kingdonella | Mean Diurnal Temperature Range (MDTR) |
| Isotherm | |
| Precipitation in the Hottest Season | |
| Elevation | |
| Wind Speed | |
| Normalized Difference Vegetation Index (NDVI) | |
| Actual Evapotranspiration | |
| Chorthippus | Mean Diurnal Temperature Range (MDTR) |
| Isotherm | |
| Precipitation in the Hottest Season | |
| Elevation | |
| Wind Speed | |
| Normalized Difference Vegetation Index (NDVI) | |
| Actual Evapotranspiration |
Appendix E
Table A1.
The suitable habitat area of Locusta in current, 2050s, and 2070s based on the prediction of the MaxEnt model.
Table A1.
The suitable habitat area of Locusta in current, 2050s, and 2070s based on the prediction of the MaxEnt model.
| Non-Suitable Habitat (×104 km2) | Suitable Habitat (×104 km2) | Marginally Suitable Habitat (×104 km2) | Moderately Suitable Habitat (×104 km2) | Highly Suitable Habitat (×104 km2) | |
|---|---|---|---|---|---|
| Current | 233.27 | 17.45 | 10.51 | 3.86 | 3.08 |
| Year 2050 | 227.49 | 23.23 | 14.56 | 5.37 | 3.3 |
| Year 2070 | 227.63 | 23.1 | 14.89 | 5.31 | 2.9 |
Table A2.
Projected Changes in Suitable Habitat Area for Locusta Species by 2050 and 2070 Relative to the Current.
Table A2.
Projected Changes in Suitable Habitat Area for Locusta Species by 2050 and 2070 Relative to the Current.
| Suitable Habitat (%) | Marginally Suitable Habitat (%) | Moderately Suitable Habitat (%) | Highly Suitable Habitat (%) | |
|---|---|---|---|---|
| Year 2050 | 33.12 | 38.53 | 39.12 | 7.14 |
| Year 2070 | 32.28 | 41.67 | 37.56 | −5.84 |
Table A3.
The suitable habitat area of Leuconemacris in current, 2050s, and 2070s based on the prediction of the MaxEnt model.
Table A3.
The suitable habitat area of Leuconemacris in current, 2050s, and 2070s based on the prediction of the MaxEnt model.
| Non-Suitable Habitat (×104 km2) | Suitable Habitat (×104 km2) | Marginally Suitable Habitat (×104 km2) | Moderately Suitable Habitat (×104 km2) | Highly Suitable Habitat (×104 km2) | |
|---|---|---|---|---|---|
| Current | 239.28 | 11.48 | 5.65 | 3.15 | 2.68 |
| Year 2050 | 239.44 | 11.29 | 5.51 | 3.99 | 1.79 |
| Year 2070 | 239.88 | 10.86 | 5.43 | 3.67 | 1.76 |
Table A4.
Projected Changes in Suitable Habitat Area for Leuconemacris Species by 2050 and 2070 Relative to the Current.
Table A4.
Projected Changes in Suitable Habitat Area for Leuconemacris Species by 2050 and 2070 Relative to the Current.
| Suitable Habitat (%) | Marginally Suitable Habitat (%) | Moderately Suitable Habitat (%) | Highly Suitable Habitat (%) | |
|---|---|---|---|---|
| Year 2050 | −1.66 | −2.48 | 26.67 | −33.21 |
| Year 2070 | −5.40 | −3.89 | 16.51 | −34.33 |
Table A5.
The suitable habitat area of Kingdonella in current, 2050s, and 2070s based on the prediction of the MaxEnt model.
Table A5.
The suitable habitat area of Kingdonella in current, 2050s, and 2070s based on the prediction of the MaxEnt model.
| Non-Suitable Habitat (×104 km2) | Suitable Habitat (×104 km2) | Marginally Suitable Habitat (×104 km2) | Moderately Suitable Habitat (×104 km2) | Highly Suitable Habitat (×104 km2) | |
|---|---|---|---|---|---|
| Current | 229.4 | 21.34 | 12.84 | 5.29 | 3.21 |
| Year 2050 | 224.8 | 25.92 | 16.37 | 6.41 | 3.14 |
| Year 2070 | 225.71 | 25.03 | 15.65 | 5.95 | 3.43 |
Table A6.
Projected Changes in Suitable Habitat Area for Kingdonella Species by 2050 and 2070 Relative to the Current.
Table A6.
Projected Changes in Suitable Habitat Area for Kingdonella Species by 2050 and 2070 Relative to the Current.
| Suitable Habitat (%) | Marginally Suitable Habitat (%) | Moderately Suitable Habitat (%) | Highly Suitable Habitat (%) | |
| Year 2050 | 21.46 | 27.49 | 21.17 | −2.18 |
| Year 2070 | 17.29 | 21.88 | 12.48 | 6.85 |
Table A7.
The suitable habitat area of Chorthippus in current, 2050s, and 2070s based on the prediction of the MaxEnt model.
Table A7.
The suitable habitat area of Chorthippus in current, 2050s, and 2070s based on the prediction of the MaxEnt model.
| Non-Suitable Habitat (×104 km2) | Suitable Habitat (×104 km2) | Marginally Suitable Habitat (×104 km2) | Moderately Suitable Habitat (×104 km2) | Highly Suitable Habitat (×104 km2) | |
|---|---|---|---|---|---|
| Current | 216.31 | 34.41 | 20.21 | 8.91 | 5.29 |
| Year 2050 | 211.86 | 38.86 | 22.1 | 10.86 | 5.9 |
| Year 2070 | 212.13 | 38.6 | 21.24 | 11.26 | 6.1 |
Table A8.
Projected Changes in Suitable Habitat Area for Chorthippus Species by 2050 and 2070 Relative to the Current.
Table A8.
Projected Changes in Suitable Habitat Area for Chorthippus Species by 2050 and 2070 Relative to the Current.
| Suitable Habitat (%) | Marginally Suitable Habitat (%) | Moderately Suitable Habitat (%) | Highly Suitable Habitat (%) | |
|---|---|---|---|---|
| Year 2050 | 12.93 | 9.35 | 21.89 | 11.53 |
| Year 2070 | 12.18 | 5.10 | 26.37 | 15.31 |
Appendix F. Species List of Acridoidea in the Qinghai–Tibet Plateau
| Species Name | Collection Site | Longitude | Latitude |
| Filchnerella sunanensis Liu, 1982 | Sunan Soap Alum Gully, Gansu | 99.12 | 39.08 |
| Baidaban Grassland, Gansu | 99.30 | 38.81 | |
| Filchnerella zhengi Huo, 1994 | Ledu, Qinghai | 102.40 | 36.48 |
| Filchnerella rufitibia Yin, 1984 | Huangliangchuan, Minhe County, Huangniangchuan, Qinghai | 102.76 | 36.35 |
| Filchnerella kukunoris Bei-Bienko, 1948 | East Shore of Qinghai Lake, Gonghe County, Qinghai | 100.44 | 37.09 |
| Filchnerella beicki Ramme, 1931 | East Shore of Qinghai Lake, Gonghe County, Qinghai | 101.18 | 37.96 |
| Eoeotmethis longipennis Zheng, 1985 | Hongliu Gully, Sunan County, Gansu | 99.24 | 39.10 |
| Beybienkia amicus (Bey-Bienko, 1959) | Qilian Mountains, Gansu | 101.18 | 37.96 |
| Aularches miliaris miliaris (Linnaeus, 1758) | Mengda, Qinghai | 102.69 | 35.82 |
| Mekongiella kingdoni (Uvarov, 1937) | Linzhi Town, Linzhi City, Tibet | 94.48 | 29.57 |
| Nanyi Gully, Milin County, Tibet | 94.21 | 28.98 | |
| Mekongiella pleurodilata Yin, 1984 | Cona, Tibet | 91.96 | 28.00 |
| Lhünzhub, Tibet | 91.61 | 31.44 | |
| Mekongiella rufitibia Yin, 1984 | Cona, Tibet | 91.96 | 28.00 |
| Mekongiella xizangensis Yin, 1984 | Cona, Tibet | 91.96 | 28.00 |
| Nyingchi County, Tibet | 93.24 | 29.89 | |
| Mekongiella wardi (Uvarov, 1937) | Quese Village, Nangqên County, Tibet | 90.47 | 29.14 |
| Paramekongiella zhongdianensis Huang, 1990 | Hutiao Gorge, Shangri-La, Yunnan | 100.11 | 27.21 |
| Mekongiana gregoryi (Uvarov, 1925) | Xiangcheng County, Sichuan | 99.80 | 28.93 |
| Deqin County, Yunnan | 98.92 | 28.47 | |
| Mekongiana xiangchengensis Zheng, Huang&Zhou, 2008 | Xiangcheng County, Sichuan | 99.80 | 28.93 |
| Yunnanites coriacea Uvarov, 1925 | Xiangcheng County, Sichuan | 99.80 | 28.93 |
| Lanping County, Yunnan | 99.42 | 26.46 | |
| Yunnanites zhengi Mao&Yang, 2003 | Shangri-La, Yunnan | 99.74 | 27.84 |
| Ninglang County, Yunnan | 100.85 | 27.29 | |
| Yunnanites albomargina Mao&Zheng, 1999 | Lanping County, Yunnan | 99.42 | 26.46 |
| Tagasta indica Bolívar, 1905 | Motuo, Tibet | 95.33 | 29.33 |
| Atractomorpha melanostriga Bi, 1981 | Motuo, Tibet | 95.33 | 29.33 |
| Cona, Tibet | 91.96 | 28.00 | |
| Xiachaiyu, Tibet | 97.02 | 28.50 | |
| Atractomorpha himalayica Bolívar, 1905 | Motuo, Tibet | 95.33 | 29.33 |
| Xiachaiyu, Tibet | 97.02 | 28.50 | |
| Atractomorpha sinensis sinensis Bolívar, 1905 | Lanping County, Yunnan | 99.42 | 26.46 |
| Gongshan County, Yunnan | 98.67 | 27.74 | |
| Fugong County, Yunnan | 98.87 | 26.90 | |
| Deqin County, Yunnan | 98.92 | 28.47 | |
| Shangri-La, Yunnan | 99.74 | 27.84 | |
| Weixi County, Yunnan | 99.29 | 27.18 | |
| Zhouqu, Gansu | 104.37 | 33.79 | |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Spathosternum prasiniferum xizangensis Yin, 1982 | Motuo, Tibet | 95.33 | 29.33 |
| Beibeng, Motuo, Tibet | 95.17 | 28.91 | |
| Peripolus nepalensis Uvarov, 1942 | Zhangmu, Tibet | 85.98 | 27.99 |
| Calliptamus barbarus barbarus (Costa, 1836) | Guide County, Qinghai | 101.43 | 36.04 |
| Jianzha, Qinghai | 102.03 | 35.94 | |
| Gandu Town, Hualong County, Qinghai | 102.33 | 35.89 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Calliptamus abbreviatus Ikonnikov, 1913 | Xiahe, Gansu | 102.52 | 35.20 |
| Zhuoni County, Gansu | 103.51 | 34.59 | |
| Qilian Mountain Bai Da Ban Grassland, Gansu | 99.30 | 38.81 | |
| Xining, Qinghai | 101.78 | 36.62 | |
| Gonghe County, Qinghai | 100.62 | 36.28 | |
| Guinan Mugetan, Qinghai | 100.83 | 35.64 | |
| Tongde County Kerbu Beach, Qinghai | 100.48 | 35.34 | |
| Guide County, Qinghai | 101.43 | 36.04 | |
| Jianzha, Qinghai | 102.03 | 35.94 | |
| Tongren County, Qinghai | 102.02 | 35.52 | |
| Tongren County, Qinghai | 103.59 | 31.48 | |
| Calliptamus italicus (Linnaeus, 1758) | Guide County, Qinghai | 101.43 | 36.04 |
| Xining, Qinghai | 101.78 | 36.62 | |
| Gangca County, Qinghai | 100.14 | 37.32 | |
| Zhiduo, Qinghai | 95.62 | 33.84 | |
| Gonghe County, Qinghai | 100.62 | 36.28 | |
| Xenocatantops brachycerus (Willemse, 1932) | Chawalong Township, Tibet | 98.46 | 28.48 |
| Mendu, Chayu County, Linzhi City, Tibet | 97.03 | 28.75 | |
| Chayu, Tibet | 97.47 | 28.66 | |
| Xiachayu, Chayu County, Tibet | 96.99 | 28.50 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Zhouqu, Gansu | 104.37 | 33.79 | |
| Maqu County, Gansu | 102.07 | 34.00 | |
| Jinchuan County, Sichuan | 102.06 | 31.48 | |
| Tongren County, Qinghai | 103.59 | 31.48 | |
| Muli County, Sichuan | 100.86 | 28.42 | |
| Liziping Township, Muli County, Sichuan | 100.29 | 27.97 | |
| Shangri-La, Yunnan | 99.74 | 27.84 | |
| Weixi County, Yunnan | 99.29 | 27.18 | |
| Ninglang County, Yunnan | 100.85 | 27.29 | |
| Xenocatantops acanthraus Zheng, Li&Wang, 2004 | Motuo, Tibet | 95.33 | 29.33 |
| Xenocatantops sauteri (Ramme, 1941) | Motuo, Tibet | 95.33 | 29.33 |
| Stenocatantops splendens (Thunberg, 1815) | Xiachayu, Chayu County, Tibet | 97.02 | 28.50 |
| Chayu, Tibet | 97.40 | 28.83 | |
| Stenocatantops mistshenkoi Willemse, 1968 | Jinchuan County, Sichuan | 102.06 | 31.48 |
| Tongren County, Qinghai | 103.59 | 31.48 | |
| Catantops simlae Dirsh, 1956 | Jilong, Tibet | 85.30 | 28.86 |
| Motuo, Tibet | 95.33 | 29.33 | |
| Catantops splendena (Thunb.) | Motuo, Tibet | 95.33 | 29.33 |
| Catantops pinguis pinguis (Stål, 1860) | Shangri-La, Yunnan | 99.74 | 27.84 |
| Muli County, Sichuan | 100.86 | 28.42 | |
| Xiachayu, Chayu County, Tibet | 97.02 | 28.50 | |
| Markam, Tibet | 98.59 | 29.69 | |
| Schistocerca gregaria (Forskal, 1775) | Jilong, Tibet | 85.30 | 28.86 |
| Dingri County, Tibet | 87.12 | 28.66 | |
| Dingjie County, Tibet | 87.77 | 28.37 | |
| Zhongba County, Tibet | 84.03 | 29.77 | |
| Nyalam, Tibet | 85.98 | 28.16 | |
| Patanga japonica (Bolívar, 1898) | Chayu, Tibet | 97.47 | 28.66 |
| Chawalong Township, Tibet | 98.46 | 28.47 | |
| Bomi, Tibet | 95.77 | 29.86 | |
| Motuo, Tibet | 95.33 | 29.33 | |
| Markam, Tibet | 98.59 | 29.69 | |
| Zhouqu, Gansu | 104.37 | 33.79 | |
| Jinchuan County, Sichuan | 102.06 | 31.48 | |
| Danba County, Sichuan | 101.89 | 30.88 | |
| Tongren County, Qinghai | 103.59 | 31.48 | |
| Kangding, Sichuan | 101.95 | 30.16 | |
| Xiangcheng County, Sichuan | 99.80 | 28.93 | |
| Ninglang County, Yunnan | 100.85 | 27.29 | |
| Deqin County, Yunnan | 98.92 | 28.47 | |
| Patanga humilis Bi, 1986 | Xiachayu, Chayu County, Tibet | 97.02 | 28.50 |
| Motuo, Tibet | 95.33 | 29.33 | |
| Chondracris rosea brunneri Uvarov, 1924 | Motuo, Tibet108KM | 95.27 | 29.30 |
| Motuo, Tibet | 95.33 | 29.33 | |
| Choroedocus robusia (Servile, 1839) | Motuo, Tibet | 95.33 | 29.33 |
| Shirakiacris yunkweiensis (Chang, 1937) | Xiachayu, Chayu County, Tibet | 97.02 | 28.50 |
| Zhouqu, Gansu | 104.37 | 33.79 | |
| Jinchuan County, Sichuan | 102.06 | 31.48 | |
| Tongren County, Qinghai | 103.59 | 31.48 | |
| Shirakiacris brachyptera Zheng, 1983 | Weixi County, Yunnan | 99.29 | 27.18 |
| Eucoptacral moiuoensis Yin, 1984 | Motuo, Tibet | 95.33 | 29.33 |
| Apalacris varicornis Walker, 1870 | Motuo, Tibet | 95.33 | 29.33 |
| Apalacris xizangensis Bi, 1984 | Motuo, Tibet | 95.33 | 29.33 |
| Beibeng, Motuo, Tibet | 95.17 | 28.91 | |
| Maniyung Village, Tibet | 94.89 | 29.31 | |
| Ecphanthacris mirabilis Tinkham, 1940 | Motuo, Tibet | 95.33 | 29.33 |
| Assamacris longicercus (Huang, 1981) | Motuo, Tibet108-80KM | 95.27 | 29.30 |
| Motuo, Tibet | 95.33 | 28.83 | |
| Assamacris curticerca (Huang, 1981) | Motuo, Tibet | 95.33 | 28.83 |
| Hanmi Ani Bridge, Motuo, Tibet | 95.29 | 29.33 | |
| Promesosternus himalayicus Yin, 1984 | Motuo, Tibet | 95.33 | 29.33 |
| Promesosternus vittatus Yin, 1984 | Motuo, Tibet | 95.33 | 29.33 |
| Habrocnemis sinensis Uvarov, 1930 | Kangding, Sichuan | 101.95 | 30.16 |
| Oxya agavisa agavisa Tsai, 1931 | Jiuzhaigou Valley, Sichuan | 104.24 | 33.26 |
| Tongren County, Qinghai | 103.59 | 31.48 | |
| Oxya chinensis (Thunberg, 1815) | Motuo, Tibet | 95.33 | 29.33 |
| Oxya intricata (Stål, 1861) | Motuo, Tibet | 95.33 | 29.33 |
| Xiachayu, Chayu County, Tibet | 97.02 | 28.50 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Luqu County, Gansu | 102.49 | 34.39 | |
| Oxya adentata Willemse, 1925 | Motuo, Tibet | 95.33 | 29.33 |
| Chayu, Tibet | 97.47 | 28.66 | |
| Ledu District, Qinghai | 102.40 | 36.48 | |
| Oxya brachyptera Zheng&Huo, 1992 | Xining, Qinghai | 101.78 | 36.62 |
| Oxya velox (Fabricius, 1787) | Motuo, Tibet | 95.33 | 29.33 |
| Oxya japonica (Thunberg, 1824) | Motuo, Tibet | 95.33 | 29.33 |
| Weixi County, Yunnan | 99.29 | 27.18 | |
| Caryanda gyirongensis Huang, 1981 | Xiaojilong, Tibet | 85.33 | 28.40 |
| Caryanda bidentata Zheng&Liang, 1985 | Langfang Village, Lushui, Yunnan | 98.60 | 26.11 |
| Parapodisma astris Huang, 2006 | Zhongdian Town, Shangri-La, Yunnan | 99.79 | 27.58 |
| Baishuitai, Shangri-La, Yunnan | 100.02 | 27.60 | |
| Conophymacris chinensis Willemse, 1933 | Jade Dragon Snow Mountain, Lijiang, Yunnan | 100.17 | 27.10 |
| Xiao Zhongdian, Yunnan | 99.79 | 27.58 | |
| Deqin County, Yunnan | 98.92 | 28.47 | |
| Kangding, Sichuan | 102.33 | 30.25 | |
| Conophymacris xianggelilaensis Niu&Zheng, 2009 | Xiao Zhongdian, Yunnan | 99.79 | 27.58 |
| Zhongdian Town, Shangri-La, Yunnan | 99.79 | 27.58 | |
| Pushang, Shangri-La, Yunnan | 99.74 | 28.15 | |
| Jade Dragon Snow Mountain, Lijiang, Yunnan | 100.17 | 27.09 | |
| Conophymacris yunnanensis Cheng, 1977 | Wenhai Reservoir, Lijiang, Yunnan | 100.16 | 26.98 |
| Conophymacris jiulongensis Zheng&Shi, 2009 | Wuxuhai Village, Jiulong County, Sichuan | 101.41 | 29.15 |
| Hongba, Jiulong County, Sichuan | 102.04 | 29.23 | |
| Conophymacris cangshanensis Zheng&Mao, 1996 | Shangri-La, Yunnan | 99.74 | 27.84 |
| Yunshanping, Lijiang, Yunnan | 100.22 | 27.11 | |
| Jade Dragon Snow Mountain, Lijiang, Yunnan | 100.18 | 27.10 | |
| Laojun Mountain, Jianchuan, Yunnan | 99.59 | 26.56 | |
| Conophymacris nigrofemora Liang, 1993 | Kangding, Sichuan | 102.33 | 30.25 |
| Muli County, Sichuan | 100.86 | 28.42 | |
| Yunshanping, Lijiang, Yunnan | 100.22 | 27.11 | |
| Curvipennis wixiensis Huang, 1984 | Weixi County, Yunnan | 99.29 | 27.18 |
| Lanping County, Yunnan | 99.42 | 26.46 | |
| Laojun Mountain, Jianchuan, Yunnan | 99.59 | 26.56 | |
| Pedopodisma emeiensis (Yin, 1980) | Tongren County, Qinghai | 103.59 | 31.48 |
| Pedopodisma abaensis Yin, Zheng&Ye, 2012 | Aba County, Sichuan | 101.70 | 32.90 |
| Pseudozubovskia xizangensis Zheng, Lin, Zhang&Zeng, 2014 | Bomi, Tibet | 95.77 | 29.86 |
| Sinopodisma zheng Liang&Lin, 1995 | Ninglang County, Yunnan | 100.85 | 27.29 |
| Xiangelilacris zhongdianensis Zheng, Huang&Zhou, 2008 | Shangri-La, Yunnan | 99.74 | 27.84 |
| Rectimargipodisma medogensis Zheng, Li&Wang, 2004 | Motuo, Tibet | 95.33 | 29.33 |
| Indopodisma kingdoni (Uvarov, 1927) | Chayu, Tibet | 97.47 | 28.66 |
| Peseudoptygonotus alpinus | Jinding Town, Lanping, Yunnan | 99.41 | 26.43 |
| Fruhstorferiola omei (Rehn&Rehn, 1939) | Tongren County, Qinghai | 103.59 | 31.48 |
| Tylotropidius yunnanensis Zheng&Liang, 1990 | Shangri-La, Yunnan | 99.74 | 27.84 |
| Rhinopodisma eminifrontus (Huang, 1981) | Yigong, Bomi, Tibet | 94.82 | 30.27 |
| Pailong, Nyingchi, Tibet | 95.01 | 30.04 | |
| Eokingdonella changtunica Yin, 1984 | Chamdo, Tibet | 97.17 | 31.15 |
| Eokingdonella cyanecuda Zheng&Shi, 2009 | Yajiang, Sichuan | 101.01 | 30.03 |
| Eokingdonella angqianensis Chen&Zheng, 2009 | Yushu, Qinghai | 97.01 | 32.99 |
| Jiuzhi, Qinghai | 101.48 | 33.43 | |
| Zhiduo, Qinghai | 95.62 | 33.84 | |
| Qumarlêb, Qinghai | 95.80 | 34.13 | |
| Nagqu, Qinghai | 96.49 | 32.21 | |
| Zadoi, Qinghai | 95.30 | 32.89 | |
| Gade, Qinghai | 99.90 | 33.97 | |
| Maqin, Qinghai | 100.24 | 34.48 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Niangra Township, Qinghai | 96.84 | 32.04 | |
| Eokingdonella gongbugyanda Zheng, Lin, Zhang&Zeng, 2014 | Gongbujiangda, Tibet | 93.24 | 29.89 |
| Eokingdonella gentiana (Uvarvo, 1939) | Basu, Tibet | 96.92 | 30.05 |
| Eokingdonella tibetana (Mistshenko, 1952) | Basu, Tibet | 96.92 | 30.05 |
| Eokingdonella burrusitibia Zheng, Dong&Xu, 2013 | Jiangda County, Tibet | 98.14 | 31.44 |
| Eokingdonella kaulbacki (Uvarvo, 1939) | Zuogong, Tibet | 97.84 | 29.67 |
| Eokingdonella bayanharaensis Huo, 1995 | Yushu, Qinghai | 97.01 | 32.99 |
| Jiuzhi, Qinghai | 101.48 | 33.43 | |
| Zhiduo, Qinghai | 95.62 | 33.84 | |
| Qumarlêb, Qinghai | 95.80 | 34.13 | |
| Nagqu, Qinghai | 96.49 | 32.21 | |
| Zadoi, Qinghai | 95.30 | 32.89 | |
| Gade, Qinghai | 99.90 | 33.97 | |
| Maqin, Qinghai | 100.24 | 34.48 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Eokingdonella luhuoensis Zheng&Shi, 2007 | Ganzi County, Sichuan | 99.99 | 31.62 |
| Luhuo, Sichuan | 100.68 | 31.39 | |
| Kingdonella bicollina Yin, 1984 | Racecourse, Yushu, Qinghai | 96.96 | 33.00 |
| Nagqu, Qinghai | 96.48 | 32.19 | |
| Baizha township, Nagqu, Qinghai | 96.52 | 32.07 | |
| Duocai, Zhiduo, Qinghai | 95.42 | 33.81 | |
| Riwoqe, Tibet | 96.60 | 31.21 | |
| Dzumbala, Markam, Tibet | 98.62 | 29.72 | |
| Xiaqu Town, Biru, Tibet | 92.83 | 31.81 | |
| Kagong Township, Jiangda County, Tibet | 98.14 | 31.44 | |
| Jiangda County, Tibet | 98.23 | 31.50 | |
| Kagong Township, Jiangda County, Tibet | 97.99 | 31.29 | |
| Ganzi County, Sichuan | 99.99 | 31.62 | |
| Kingdonella qinghaiensis Zheng, 1990 | Duocai, Zhiduo, Qinghai | 95.42 | 33.81 |
| Baizha township, Nagqu, Qinghai | 96.52 | 32.07 | |
| Racecourse, Yushu, Qinghai | 96.96 | 33.00 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Jiuzhi, Qinghai | 101.48 | 33.43 | |
| Kingdonella pienbaensis Zheng, 1980 | Xiaqu Town, Biru, Tibet | 92.83 | 31.81 |
| Kagong Township, Jiangda County, Tibet | 97.99 | 31.29 | |
| Bianba, Tibet | 94.71 | 30.93 | |
| Balila Mountain, Bianba, Tibet | 95.27 | 30.79 | |
| Demula Mountain, Chayu, Qinghai | 97.03 | 29.32 | |
| Chabatang, Chayu, Qinghai | 97.22 | 32.83 | |
| Shangbatang, Chayu, Qinghai | 97.07 | 32.85 | |
| Yushu, Qinghai | 97.01 | 32.99 | |
| Bulang, Chayu, Qinghai | 96.77 | 33.35 | |
| Longbao, Chayu, Qinghai | 96.42 | 33.26 | |
| Gelong, Chayu, Qinghai | 95.56 | 32.64 | |
| Xiao Sumang, Chayu, Qinghai | 97.25 | 32.33 | |
| Duocai, Zhiduo, Qinghai | 95.42 | 33.81 | |
| Kingdonella nigrotibia Zheng, 1990 | Maharong Gully, Baiyu, Sichuan | 98.87 | 31.17 |
| Honglong Town, Yajiang, Sichuan | 100.52 | 30.11 | |
| Kingdonella magna Yin, 1984 | Dari County, Qinghai | 99.65 | 33.75 |
| Kingdonella nigrofemora Yin, 1984 | Jiangda County, Tibet | 98.10 | 31.80 |
| Jiuzhi, Qinghai | 101.48 | 33.43 | |
| Kingdonella hanburyi Uvarov, 1939 | Zuogong, Tibet | 97.84 | 29.67 |
| Basu, Tibet | 96.92 | 30.05 | |
| Ganzi County, Sichuan | 99.99 | 31.62 | |
| Kingdonella parvula Yin, 1984 | Riwoqe, Tibet | 96.60 | 31.21 |
| Kingdonella longiconica Yin, 1984 | Mount Kaji La, Tibet | 96.40 | 31.60 |
| Kingdonella pictipes Uvarov, 1935 | Ranwu Town, Tibet | 96.75 | 29.51 |
| Riwoqe, Tibet | 96.60 | 31.21 | |
| Chabatang, Chayu, Qinghai | 97.22 | 32.83 | |
| Shangbatang, Chayu, Qinghai | 97.07 | 32.85 | |
| Yushu, Qinghai | 97.01 | 32.99 | |
| Longbao, Chayu, Qinghai | 96.42 | 33.26 | |
| Xiao Sumang, Chayu, Qinghai | 97.25 | 32.33 | |
| Bulang, Chayu, Qinghai | 96.77 | 33.35 | |
| Kingdonella kozlovi Mistshenko, 1952 | Jiuzhi, Qinghai | 101.48 | 33.43 |
| Yushu, Qinghai | 97.01 | 32.99 | |
| Chamdo, Tibet | 97.17 | 31.15 | |
| Kingdonella conica Yin, 1984 | Riwoqe, Tibet | 96.60 | 31.21 |
| Kingdonella afurcula Yin, 1984 | Riwoqe, Tibet | 96.60 | 31.21 |
| Kingdonella saxicola Uvarvo, 1939 | Basu, Tibet | 96.92 | 30.05 |
| Kingdonella rivuna Huang, 1981 | Chaya Jitang, Tibet | 97.35 | 30.74 |
| Kingdonella wardi Uvarvo, 1933 | Chayu, Tibet | 97.47 | 28.66 |
| Kingdonella conica Yin, 1984 | Mount Kaji La, Tibet | 96.40 | 31.60 |
| Anepipodisma punctata Huang, 1984 | Markam, Tibet | 98.59 | 29.69 |
| Chayu, Tibet | 97.47 | 28.66 | |
| Deqin County, Yunnan | 98.92 | 28.47 | |
| Conophymopsis linguspinus Huang, 1983 | Tashkurgan Tajik Autonomous County, Xinjiang | 75.23 | 37.77 |
| Akto County, Xinjiang | 75.37 | 38.84 | |
| Conophymopsis labrispinus Huang, 1983 | Yecheng County, Xinjiang | 76.70 | 37.22 |
| Dericorys annulata roseipennis (Redtenbacher, 1889) | Xiahe, Gansu | 102.52 | 35.20 |
| Leptopternis gracilis (Eversmann, 1848) | Qilian Mountains, Jiayuguan, Gansu | 98.29 | 39.32 |
| Heteropternis respondens (Walker, 1859) | Markam, Tibet | 98.59 | 29.69 |
| Jinchuan County, Sichuan | 102.06 | 31.48 | |
| Deqin County, Yunnan | 98.92 | 28.47 | |
| Heteropternis robusta Bey-Bienko, 1951 | Markam, Tibet | 98.59 | 29.69 |
| Xiangcheng County, Sichuan | 99.80 | 28.93 | |
| Kangding, Sichuan | 101.95 | 30.16 | |
| Luding County, Sichuan | 102.23 | 29.92 | |
| Yanyuan County, Sichuan | 101.39 | 27.69 | |
| Lanping County, Yunnan | 99.42 | 26.46 | |
| Shangri-La, Yunnan | 99.74 | 27.84 | |
| Deqin County, Yunnan | 98.92 | 28.47 | |
| Ninglang County, Yunnan | 100.85 | 27.29 | |
| Heteropternis motuoensis Yin, 1984 | Motuo, Tibet | 95.33 | 29.33 |
| Heteropternis qinghaiensis Wang&Zheng, 1992 | Qumarlêb, Qinghai | 95.80 | 34.13 |
| Heteropternis micronus Huang, 1981 | Xiachayu, Chayu County, Tibet | 97.02 | 28.50 |
| Songgula Ga, Tibet | 97.24 | 29.16 | |
| Shangri-La, Yunnan | 99.74 | 27.84 | |
| Pusana chayuensis (Yin, 1984) | Chayu, Tibet | 97.47 | 28.66 |
| Mecostethus grossus (Linnaeus, 1758) | Qilian County, Qinghai | 100.25 | 38.18 |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Kangding, Sichuan | 101.95 | 30.16 | |
| Ruoergai County, Sichuan | 102.96 | 33.58 | |
| Ochyracris rufotibialis Zheng, 1991 | Jinping Mountain, Mianyang County, Sichuan | 101.82 | 28.46 |
| Epacromius coerulipes (Ivanov, 1888) | Guinan County, Qinghai | 100.75 | 35.59 |
| Daotang River, Gonghe County, Qinghai | 100.97 | 36.40 | |
| Guide County, Qinghai | 101.43 | 36.04 | |
| Xining, Qinghai | 101.78 | 36.62 | |
| Ledu District, Qinghai | 102.40 | 36.48 | |
| Ping’an County, Qinghai | 102.11 | 36.50 | |
| Xunhua County, Qinghai | 102.49 | 35.85 | |
| Epacromius tergestinus tergestinus (Megerle von Mühlfeld, 1825) | Wutumeiren Township, Geermu, Qinghai | 93.16 | 36.91 |
| Tashkurgan Tajik Autonomous County, Xinjiang | 75.23 | 37.77 | |
| Xiachayu, Chayu County, Tibet | 97.02 | 28.50 | |
| Chayu, Tibet | 97.40 | 28.83 | |
| Epacromius tergestinus extimus Bey-Bienko, 1951 | Ledu District, Qinghai | 102.40 | 36.48 |
| Geermu, Qinghai | 94.90 | 36.40 | |
| Dachaidan, Qinghai | 95.36 | 37.85 | |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Xining, Qinghai | 101.78 | 36.62 | |
| Muli Tibetan Autonomous County, Sichuan | 100.86 | 28.42 | |
| Aiolopus thalassinus (Fabricius, 1798) | Deqin County, Yunnan | 98.92 | 28.47 |
| Aiolopus tamulus (Fabricius, 1798) | Luhuo, Sichuan | 100.68 | 31.39 |
| Chawalong Township, Tibet | 98.46 | 28.47 | |
| Chayu, Tibet | 97.47 | 28.66 | |
| Aiolopus markamensis Yin, 1984 | Markam, Tibet | 98.59 | 29.69 |
| Xiachayu, Chayu County, Tibet | 97.02 | 28.50 | |
| Muli County, Sichuan | 100.86 | 28.42 | |
| Derong County, Sichuan | 99.29 | 28.72 | |
| Oedaleus infernalis Saussure, 1884 | Xiahe, Gansu | 102.52 | 35.20 |
| Qilian Shan Grassland, Bai Da Ban Prairie, Gansu | 99.30 | 38.81 | |
| Daotang River, Gonghe County, Qinghai | 100.97 | 36.40 | |
| Jianzha, Qinghai | 102.03 | 35.94 | |
| Hualong Hui Autonomous County, Qinghai | 102.26 | 36.09 | |
| Xunhua County, Qinghai | 102.49 | 35.85 | |
| Guide County, Qinghai | 101.43 | 36.04 | |
| Tongren County, Qinghai | 102.02 | 35.52 | |
| Xining, Qinghai | 101.78 | 36.62 | |
| Datong Hui and Tu Autonomous County, Qinghai | 101.69 | 36.93 | |
| Ping’an County, Qinghai | 102.11 | 36.50 | |
| Huzhu Tuzu Zizhixian, Qinghai | 101.96 | 36.84 | |
| Menyuan Hui Autonomous County, Qinghai | 101.61 | 37.39 | |
| Oedaleus asiaaicus Bey-Bienko, 1941 | Xining, Qinghai | 101.78 | 36.62 |
| Gonghe County, Qinghai | 100.97 | 36.40 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Guide County, Qinghai | 101.43 | 36.04 | |
| Jianzha, Qinghai | 102.03 | 35.94 | |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Hualong Hui Autonomous County, Qinghai | 102.26 | 36.09 | |
| Tongren County, Qinghai | 102.02 | 35.52 | |
| Datong Hui and Tu Autonomous County, Qinghai | 101.69 | 36.93 | |
| Ping’an County, Qinghai | 102.11 | 36.50 | |
| Huzhu Tuzu Zizhixian, Qinghai | 101.96 | 36.84 | |
| Menyuan Hui Autonomous County, Qinghai | 101.61 | 37.39 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Xiahe, Gansu | 102.44 | 35.06 | |
| Jinchuan County, Sichuan | 102.06 | 31.48 | |
| Daocheng County, Sichuan | 100.29 | 28.72 | |
| Tongren County, Qinghai | 103.59 | 31.48 | |
| Maerkang County, Sichuan | 102.20 | 31.91 | |
| Oedaleus manjius Chang, 1939 | Muli County, Sichuan | 100.86 | 28.42 |
| Ganzi County, Sichuan | 99.99 | 31.62 | |
| Tongren County, Qinghai | 103.59 | 31.48 | |
| Acrotylus longipes subfasciatus Bey-Bienko, 1984 | Zhangmu, Tibet | 85.98 | 27.99 |
| Aurilobulus splendens Yin, 1979 | Zanda Xiangquan River Collection Site, Tibet | 80.30 | 31.11 |
| Cyanicaudata annulicornea Yin, 1979 | Zanda Xiangquan River Collection Site, Tibet | 80.30 | 31.11 |
| Celes skalozubovi Adelung, 1906 | Jianzha, Qinghai | 102.03 | 35.94 |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Lintan County, Gansu | 103.53 | 34.79 | |
| Celes akitanus (Shiraki, 1910) | Jianzha, Qinghai | 102.03 | 35.94 |
| Sphingonotus kueideensis Yin, 1984 | Guide County, Qinghai | 101.43 | 36.04 |
| Sphingonotus qinghaiensis Yin, 1984 | Jianzha, Qinghai | 102.03 | 35.94 |
| Sphingonotus zhengi Huo, 1994 | Ledu District, Qinghai | 102.40 | 36.48 |
| Sphingonotus yunnaneus Uvarov,1925 | Markam, Tibet | 98.59 | 29.69 |
| Xiangcheng County, Sichuan | 99.80 | 28.93 | |
| Deiqen Moon Bay, Yunnan | 99.31 | 28.20 | |
| Shangri-La, Yunnan | 99.74 | 27.84 | |
| Sphingonotus tipicus Zheng&Hang, 1974 | Gonghe Tiebujia, Qinghai | 99.58 | 37.03 |
| Sphingonotus halophilus Bey-Bienko, 1929 | Dachaidan Administrative Committee, Haixi Tibetan Autonomous Prefecture, Qinghai | 95.36 | 37.86 |
| Sphingonotus tzaidamicus Mistshenko,1937 | Delhi City, Qinghai | 97.36 | 37.37 |
| Dachaidan, Qinghai | 95.36 | 37.85 | |
| Geermu, Qinghai | 94.90 | 36.40 | |
| Dulan County, Qinghai | 98.10 | 36.30 | |
| Tanggula, Qinghai | 92.44 | 34.21 | |
| Sphingonotus caerulistriatus Zheng&Ren,2007 | Yanjing, Markam, Tibet | 98.61 | 29.61 |
| Sphingonotus zadaensis Huang, 1981 | Zanda Mayang, Tibet | 78.96 | 31.79 |
| Sphingonotus longipennis Saussure, 1884 | Chayu, Tibet | 97.47 | 28.66 |
| Xiachaiyu, Tibet | 97.02 | 28.50 | |
| Sphingonotus mengolicus Saussure, 1884 | Diebu County, Gansu | 103.22 | 34.06 |
| Sphingonotus obscuratus latissimus Uvarvo, 1925 | Tashkurgan Tajik Autonomous County, Xinjiang | 75.23 | 37.77 |
| Sphingonotus elegans Mistshenko, 1937 | Tashkurgan Tajik Autonomous County, Xinjiang | 75.23 | 37.77 |
| Sphingonotus turcmenus Bey-Bienko, 1951 | Dachaidan, Qinghai | 95.36 | 37.85 |
| Tibetacris changtunensis Chen, 1964 | Chamdo, Tibet | 97.17 | 31.15 |
| Chayagang, Tibet | 97.35 | 30.74 | |
| Yanjing, Markam, Tibet | 98.61 | 29.61 | |
| Trilophidia annulata (Thunberg, 1815) | Motuo, Tibet | 95.33 | 29.33 |
| Qilian Moutain, Qinghai | 100.34 | 38.30 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Zhouqu, Gansu | 104.37 | 33.79 | |
| Xiahe, Gansu | 102.44 | 35.06 | |
| Muli County, Sichuan | 100.86 | 28.42 | |
| Weixi County, Yunnan | 99.29 | 27.18 | |
| Lanping County, Yunnan | 99.42 | 26.46 | |
| Parahilethera xizangensis Zheng&Ren, 2007 | Chamdo, Tibet | 97.17 | 31.15 |
| Yanjing, Markam, Tibet | 98.61 | 29.61 | |
| Gastrimargus africanus africanus (Saussure, 1888) | Zhangmu, Tibet | 85.98 | 27.99 |
| Motuo, Tibet | 95.33 | 29.33 | |
| Jinchuan County, Sichuan | 102.06 | 31.48 | |
| Luding County, Sichuan | 102.23 | 29.92 | |
| Weixi County, Yunnan | 99.29 | 27.18 | |
| Laojun Mountain, Jianchuan, Yunnan | 99.59 | 26.56 | |
| Shangri-La, Yunnan | 99.74 | 27.84 | |
| Gastrimargus nubilis Uvarov, 1925 | Chamdo, Tibet | 97.17 | 31.15 |
| Markam, Tibet | 98.59 | 29.69 | |
| Maerkang County, Sichuan | 102.20 | 31.91 | |
| Jinchuan County, Sichuan | 102.06 | 31.48 | |
| Lixian County, Sichuan | 103.16 | 31.44 | |
| Batang County, Sichuan | 99.11 | 30.01 | |
| Kangding, Sichuan | 102.33 | 30.50 | |
| Daocheng County, Sichuan | 100.29 | 28.72 | |
| Ganzi County, Sichuan | 99.99 | 31.62 | |
| Shangri-La, Yunnan | 99.74 | 27.84 | |
| Deqin County, Yunnan | 98.92 | 28.47 | |
| Ninglang County, Yunnan | 100.85 | 27.29 | |
| Weixi County, Yunnan | 99.29 | 27.18 | |
| Lanping County, Yunnan | 99.42 | 26.46 | |
| Gastrimargus marmoratus (Thunberg, 1815) | Jinchuan County, Sichuan | 102.06 | 31.48 |
| Luding County, Sichuan | 102.23 | 29.92 | |
| Jinsha River, Markam, Tibet | 99.01 | 29.62 | |
| Gastrimargus ommatidius Huang, 1981 | Chayab, Tibet | 97.62 | 30.61 |
| Gastrimargus africanus parvulus Sjöstedt, 1928 | Deiqen Moon Bay, Yunnan | 99.31 | 28.20 |
| Locusta migratoria migratorioides (Reiche & Fairmaire, 1849) | Qingduo Township, Bomi, Tibet | 95.60 | 30.06 |
| Dongga Township, Shigatse, Tibet | 88.83 | 29.36 | |
| Qubuxiong, Shigatse, Tibet | 88.87 | 29.18 | |
| Tongmai, Bomi, Tibet | 95.08 | 30.10 | |
| Guxiang, Bomi, Tibet | 95.50 | 29.91 | |
| Dongjiu, Nyingchi, Tibet | 94.80 | 29.94 | |
| Bayi Town North, Nyingchi, Tibet | 94.36 | 29.65 | |
| Dingri, Shigatse, Tibet | 88.85 | 29.00 | |
| Gongbujiangda, Tibet | 93.24 | 29.89 | |
| Linzhi Town, Linzhi City, Tibet | 94.48 | 29.57 | |
| Baiba, Nyingchi, Tibet | 94.30 | 29.72 | |
| Yigong, Bomi, Tibet | 95.03 | 30.15 | |
| Milin County, Tibet | 94.22 | 29.21 | |
| Pulan County, Tibet | 81.18 | 30.29 | |
| Sangri County, Tibet | 92.02 | 29.26 | |
| Zhanang, Tibet | 91.34 | 29.24 | |
| Zada Naga, Tibet | 79.74 | 31.54 | |
| Shigatse, Tibet | 88.88 | 29.27 | |
| Zhada Xiangzi, Tibet | 79.63 | 31.83 | |
| Rutog County, Tibet | 79.73 | 33.38 | |
| Gar County, Tibet | 80.09 | 32.49 | |
| Namling County, Tibet | 89.10 | 29.69 | |
| Gar County, Tibet | 89.73 | 28.86 | |
| Sakya County, Tibet | 88.02 | 28.90 | |
| Tongmen Township, Tibet | 88.20 | 29.42 | |
| Geji County, Tibet | 81.14 | 32.40 | |
| Gongjue, Tibet | 98.27 | 30.86 | |
| Zuogong, Tibet | 97.84 | 29.67 | |
| Yanjing, Tibet | 98.61 | 29.61 | |
| Jilong, Tibet | 85.30 | 28.86 | |
| Motuo, Tibet | 95.33 | 29.33 | |
| Linzho County, Tibet | 91.26 | 29.90 | |
| Lhasa, Tibet | 91.17 | 29.66 | |
| Chayu, Tibet | 97.47 | 28.66 | |
| Chayu, Tibet Shangcaoyu | 96.78 | 28.72 | |
| Nedong County, Zetang, Tibet | 91.76 | 29.23 | |
| Lazi County, Tibet | 87.64 | 29.08 | |
| Bai Zha Township, Nagqu, Qinghai | 96.52 | 32.07 | |
| Nagqu, Qinghai | 96.48 | 32.19 | |
| Yushu, Qinghai | 97.01 | 32.99 | |
| Luhuo, Sichuan | 100.68 | 31.39 | |
| Yimu Township, Luhuo, Sichuan | 100.77 | 31.29 | |
| Luoshu, Shiqiu County, Sichuan | 98.00 | 32.47 | |
| Gengqing Town, Dege County, Sichuan | 98.58 | 31.82 | |
| Ganzi County, Sichuan | 99.99 | 31.62 | |
| Zhengke Township, Shiqiu County, Sichuan | 97.83 | 32.52 | |
| Litang County, Ganzi, Sichuan | 100.27 | 30.00 | |
| Zangba Township, Litang County, Sichuan | 100.44 | 29.65 | |
| Xiangcheng County, Sichuan | 99.80 | 28.93 | |
| Bamei, Daofu, Sichuan | 101.48 | 30.48 | |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Lintan County, Gansu | 103.53 | 34.79 | |
| Jinchuan County, Sichuan | 102.06 | 31.48 | |
| Ganzi County, Sichuan | 99.99 | 31.62 | |
| Muli County, Sichuan | 100.86 | 28.42 | |
| Shangri-La, Yunnan | 99.74 | 27.84 | |
| Deqin County, Yunnan | 98.92 | 28.47 | |
| Weixi County, Yunnan | 99.29 | 27.18 | |
| Ninglang County, Yunnan | 100.85 | 27.29 | |
| Locusta migratoria migratoria (Linnaeus, 1758) | Xining, Qinghai | 101.78 | 36.62 |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Pternoscirta saueri (Karny, 1915) | Motuo, Tibet | 95.33 | 29.33 |
| Pternoscirta pulchripes Uvarov, 1925 | Weixi County, Yunnan | 99.29 | 27.18 |
| Lanping County, Yunnan | 99.30 | 26.59 | |
| Pternoscirta longipennis Xia, 1981 | Xiaojilong, Jilong, Tibet | 85.33 | 28.40 |
| Bryodemella xizangensis Yin, 1984 | Shigatse, Tibet | 88.88 | 29.27 |
| Chamdo, Tibet | 96.35 | 30.51 | |
| Dingri County, Tibet | 86.98 | 28.47 | |
| Saga, Tibet | 85.27 | 29.38 | |
| Sakya, Tibet | 88.02 | 28.90 | |
| Bryodemella tuberculatum dilutum (Stoll, 1813) | Xiahe, Gansu | 102.52 | 35.20 |
| Luqu County, Gansu | 102.49 | 34.39 | |
| Heiyatou, Qilian Mountains, Sunan, Gansu | 100.00 | 38.50 | |
| Dongliugou, Sunan, Gansu | 99.64 | 38.77 | |
| Zaofangou, Sunan, Gansu | 99.12 | 39.08 | |
| Nedong County, Zetang, Tibet | 91.76 | 29.23 | |
| Namtso, Damxung, Tibet | 90.64 | 30.66 | |
| Quse, Nangqên County, Tibet | 90.47 | 29.14 | |
| Yamdrok Lake, Nangqên County, Tibet | 90.62 | 29.19 | |
| Bairang Monastery, Tibet | 89.23 | 29.14 | |
| Lumbu Monastery, Jiangzi County, Tibet | 89.73 | 28.86 | |
| Yamdrok Lake, Tibet | 90.71 | 28.92 | |
| Jiangtang Village, Sangri County, Tibet | 91.98 | 29.25 | |
| Rongbu Town, Sog County, Tibet | 94.65 | 31.55 | |
| Duina Township, Yatung County, Tibet | 89.23 | 27.98 | |
| Heika Mountain, Xinghai County, Qinghai | 100.05 | 35.78 | |
| Daotang River, Gonghe County, Qinghai | 100.97 | 36.40 | |
| Qilian County, Qinghai | 100.25 | 38.18 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Jianzha, Qinghai | 101.88 | 35.93 | |
| Ruoergai County, Sichuan Erlang Mountain | 102.79 | 34.03 | |
| Ruoergai County, Sichuan | 102.96 | 33.58 | |
| Maerkang County, Sichuan | 102.20 | 31.91 | |
| Hongyuan County, Sichuan | 102.54 | 32.79 | |
| Jinchuan County, Sichuan | 102.06 | 31.48 | |
| Ganzi County, Sichuan | 99.99 | 31.62 | |
| Bryodemella diamesum (Bey-Bienko, 1930) | Batang, Yushu, Qinghai | 97.15 | 32.83 |
| Racecourse, Yushu, Qinghai | 96.96 | 33.00 | |
| Zadoi, Qinghai | 95.30 | 32.89 | |
| Jiangda County, Tibet | 98.22 | 31.50 | |
| Riwoqe, Tibet | 96.60 | 31.21 | |
| Chamdo, Tibet | 97.17 | 31.15 | |
| Bryodemella (Bryodemella) rufifemura Zhang, Zhi&Zhang, 2018 | Qushe Village, Lhazê County, Tibet | 90.47 | 29.14 |
| Bryodemella (Bryodemella) nigrifemura Yin&Wang, 2005 | Nedong County, Zetang, Tibet | 91.76 | 29.23 |
| Lhasa, Tibet | 91.13 | 29.66 | |
| Lhazê County, Shannan, Tibet | 90.40 | 28.97 | |
| Yamdrok Lake, Tibet | 90.71 | 28.92 | |
| Bryodemella holdereri holdereri (Krauss, 1901) | Xiahe, Gansu | 102.52 | 35.20 |
| Dongliugou, Gansu | 99.64 | 38.77 | |
| Xining, Qinghai | 101.78 | 36.62 | |
| Daotang River, Gonghe County, Qinghai | 100.97 | 36.40 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Jianzha, Qinghai | 102.03 | 35.94 | |
| Xinghai, Qinghai | 99.99 | 35.59 | |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Guide County, Qinghai | 101.43 | 36.04 | |
| Tongren County, Qinghai | 102.02 | 35.52 | |
| Datong Hui and Tu Autonomous County, Qinghai | 101.69 | 36.93 | |
| Ping’an County, Qinghai | 102.11 | 36.50 | |
| Huzhu Tuzu Zizhixian, Qinghai | 101.96 | 36.84 | |
| Menyuan Hui Autonomous County, Qinghai | 101.61 | 37.39 | |
| Bryodema luctuosum luctuosum (Stoll, 1813) | Rutog County, Tibet | 79.73 | 33.38 |
| Amdo, Tibet | 91.33 | 32.33 | |
| Gar County, Tibet | 80.09 | 32.49 | |
| Pulan County, Tibet | 81.18 | 30.29 | |
| Damxung County, Tibet | 91.10 | 30.48 | |
| Yangbajing, Tibet | 90.54 | 30.10 | |
| Namtso, Damxung, Tibet | 90.64 | 30.66 | |
| Nagqu City, Tibet | 92.05 | 31.48 | |
| Nima County, Heping Township, Tibet | 88.63 | 32.13 | |
| Shenzha County, Tibet | 88.71 | 30.93 | |
| Biru County, Datang Township, Tibet | 93.11 | 31.53 | |
| Bange County, Beila Town, Tibet | 90.83 | 31.40 | |
| Bange County, Mendang Township, Tibet | 89.75 | 31.42 | |
| Gegye County, Wenbu Dang Sang Township, Tibet | 82.90 | 32.39 | |
| Geji County, Salt Lake Township, Tibet | 82.46 | 32.53 | |
| Amdo, Tibet | 91.68 | 32.26 | |
| Nagqu Bange County, Tibet | 90.01 | 31.39 | |
| Duina Township, Yatung County, Tibet | 89.23 | 27.98 | |
| Gaize County, Dongcuo Township, Tibet | 84.88 | 32.10 | |
| Lhasa Linzhou, Tibet | 91.26 | 29.90 | |
| Lhasa, Tibet | 91.17 | 29.66 | |
| Pulan County, Mapam Yumco, Tibet | 81.47 | 30.66 | |
| Pulan County, Huo’er Township, Tibet | 81.61 | 30.77 | |
| Gaize County, Tibet | 84.05 | 32.30 | |
| Zanda County, Xiangzi Township, Tibet | 79.63 | 31.83 | |
| Cuoqin County, Tibet | 85.15 | 31.02 | |
| Cuoqin County, Xiabula Mountain, Tibet | 85.04 | 31.16 | |
| Lhazê County, Tibet | 90.40 | 28.97 | |
| Maqin, Qinghai | 100.24 | 34.48 | |
| Guinan County, Qinghai | 100.87 | 35.73 | |
| Racecourse, Yushu, Qinghai | 96.96 | 33.00 | |
| Zhiduo, Qinghai | 95.61 | 33.87 | |
| Duocai, Zhiduo, Qinghai | 95.42 | 33.81 | |
| Batang, Yushu, Qinghai | 97.15 | 32.83 | |
| Madoi County, Golog Tibetan Autonomous Prefecture, Qinghai | 98.21 | 34.92 | |
| Nian Di Township, Gonghe County, Qinghai | 100.39 | 36.38 | |
| Gonghe Tiebujia, Qinghai | 99.58 | 37.03 | |
| Hei Ma He Township, Gonghe County, Qinghai | 99.78 | 36.73 | |
| Ziketan, Xinghai County, Qinghai | 99.98 | 35.59 | |
| Batan, Qinghai | 101.31 | 35.32 | |
| Xining, Qinghai | 101.78 | 36.62 | |
| Jianzha, Qinghai | 102.03 | 35.94 | |
| Qilian County, Qinghai | 100.25 | 38.18 | |
| Tongren County, Qinghai | 102.02 | 35.52 | |
| Tianjun County, Qinghai | 99.02 | 37.30 | |
| Madoi County, Qinghai | 98.21 | 34.91 | |
| Qinghai Lake | 100.35 | 36.75 | |
| Altun Mountain Range, Xinjiang | 90.16 | 38.39 | |
| Shaziquan, Xinjiang | 88.48 | 38.12 | |
| Xiahe, Gansu | 102.44 | 35.06 | |
| Bryodema miramae miramae Bey-Bienko, 1930 | Dongliugou, Sunan, Gansu | 99.64 | 38.77 |
| Heiyatou, Qilian Mountains, Sunan, Gansu | 100.00 | 38.50 | |
| Gonghe County, Qinghai | 100.62 | 36.28 | |
| Qilian County, Qinghai | 100.25 | 38.18 | |
| Xining, Qinghai | 101.78 | 36.62 | |
| Datong Hui and Tu Autonomous County, Qinghai | 101.69 | 36.93 | |
| Ping’an County, Qinghai | 102.11 | 36.50 | |
| Huzhu Tuzu Zizhixian, Qinghai | 101.96 | 36.84 | |
| Menyuan Hui Autonomous County, Qinghai | 101.61 | 37.39 | |
| Bryodema uvarovi (Bey-Bienko, 1930) | Dongliugou, Sunan, Gansu | 99.64 | 38.77 |
| Jianzha, Qinghai | 102.03 | 35.94 | |
| Tongren County, Qinghai | 102.02 | 35.52 | |
| Guide County, Qinghai | 101.43 | 36.04 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Bryodema miramae elegantulum Bey-Bienko, 1930 | Yushu, Qinghai | 97.01 | 32.99 |
| Qilian County, Qinghai | 100.25 | 38.18 | |
| Gongjue, Tibet | 98.27 | 30.86 | |
| Chamdo, Tibet | 97.17 | 31.15 | |
| Bryodema luctuosum inda Saussure, 1884 | Ziketan, Xinghai County, Qinghai | 99.98 | 35.59 |
| Taga, Tibet | 91.79 | 28.38 | |
| Shannan City, Tibet | 91.89 | 28.77 | |
| Linzhi, Tibet | 95.46 | 29.39 | |
| Cona, Tibet | 92.17 | 28.09 | |
| Motuo, Tibet | 95.00 | 29.33 | |
| Chayu, Tibet | 97.40 | 28.83 | |
| Lhünzhub County, Tibet | 92.79 | 28.45 | |
| Himalayas, Tibet | 85.17 | 28.66 | |
| Bryodema hyalinala Zheng&Zhang, 1981 | Guanjiao Beach, Tianjun County, Qinghai | 98.91 | 37.22 |
| Bryodema pseudohyalinala Zheng, Lin&Zhang, 2012 | Langma, Gongba County, Tibet | 88.51 | 28.27 |
| Bryodema kangmarensis Zheng, Lin&Zhang, 2012 | Kangma Gala, Tibet | 89.68 | 28.56 |
| Bryodema nigristria Zheng&Chen, 2001 | Yushu, Qinghai | 97.01 | 32.99 |
| Bryodema gebleri gebleri (Fischer von Waldheim, 1836) | Zhimen Da, Yushu, Qinghai | 97.26 | 33.01 |
| Yushu, Qinghai | 97.01 | 32.99 | |
| Bryodema gansuensis Zheng, 1985 | Diebu County, Gansu | 103.22 | 34.06 |
| Bryodema ochropenna Zheng&Xi, 1985 | Dachagai Ranch, Subei Mongol Autonomous County, Gansu | 99.51 | 38.66 |
| Bryodema qilianshanicum Lian&Zheng, 1984 | Xining, Qinghai | 101.78 | 36.62 |
| Datong Hui and Tu Autonomous County, Qinghai | 101.69 | 36.93 | |
| Ping’an County, Qinghai | 102.11 | 36.50 | |
| Huzhu Tuzu Zizhixian, Qinghai | 101.96 | 36.84 | |
| Menyuan Hui Autonomous County, Qinghai | 101.61 | 37.39 | |
| Angaracris barabensis (Pallas, 1773) | Tianjun County, Qinghai | 99.02 | 37.30 |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Gonghe County, Qinghai | 100.62 | 36.28 | |
| Qilian County, Qinghai | 100.25 | 38.18 | |
| Baiyang Gully, Qilian County, Qinghai | 100.28 | 38.24 | |
| Gangcha County, Qinghai | 100.14 | 37.32 | |
| Xinghai, Qinghai | 99.99 | 35.59 | |
| Xunhua County, Qinghai | 102.45 | 35.68 | |
| Baoan, Tongren County, Qinghai | 102.07 | 35.62 | |
| Mai Xiu Forest Farm, Zeku County, Qinghai | 101.93 | 35.26 | |
| Luqu County, Gansu | 102.49 | 34.39 | |
| Maqu County, Gansu | 102.07 | 34.00 | |
| Lintan, Gansu | 103.53 | 34.79 | |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Angaracris rhodopa (Fischer von Waldheim, 1836) | Xining, Qinghai | 101.78 | 36.62 |
| Daotang River, Gonghe County, Qinghai | 100.97 | 36.40 | |
| Gonghe Tiebujia, Qinghai | 99.58 | 37.03 | |
| Nian Di Township, Gonghe County, Qinghai | 100.39 | 36.38 | |
| Santa La, Gonghe County, Qinghai | 100.27 | 36.06 | |
| Ziketan, Xinghai County, Qinghai | 99.98 | 35.59 | |
| Dahe Dam, Xinghai, Qinghai | 100.01 | 35.52 | |
| Wenquan Township, Xinghai, Qinghai | 99.67 | 35.22 | |
| Heika Mountain, Xinghai County, Qinghai | 100.05 | 35.78 | |
| Batan, Qinghai | 101.31 | 35.32 | |
| Guinan County, Qinghai | 100.74 | 35.57 | |
| Guinan Mugetan, Qinghai | 100.48 | 35.34 | |
| Duogun Township, Tongde, Qinghai | 100.37 | 34.94 | |
| Qingmei Township, Tongde, Qinghai | 101.21 | 34.82 | |
| Guide County, Qinghai | 101.43 | 36.04 | |
| Xunhua County, Qinghai | 102.49 | 35.85 | |
| Tianjun County, Qinghai | 99.02 | 37.30 | |
| Huzhu Tuzu Zizhixian, Qinghai | 101.96 | 36.84 | |
| Datong Hui and Tu Autonomous County, Qinghai | 101.69 | 36.93 | |
| Ping’an County, Qinghai | 102.11 | 36.50 | |
| Menyuan Hui Autonomous County, Qinghai | 101.61 | 37.39 | |
| Lintan, Gansu | 103.53 | 34.79 | |
| Bai Da Ban Prairie, Qilian Mountains, Gansu | 99.30 | 38.81 | |
| Maqu County, Gansu | 102.07 | 34.00 | |
| Luqu County, Gansu | 102.49 | 34.59 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Zhuoni County, Gansu | 103.51 | 34.59 | |
| Zhouqu, Gansu | 104.37 | 33.79 | |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Angaracris morulimarginis Huang, 1981 | Lhasa, Tibet | 91.17 | 29.66 |
| Gangcha County, Qinghai | 100.14 | 37.32 | |
| Angaracris acrohylinna Bi, 1986 | Lhasa, Tibet | 91.17 | 29.66 |
| Angaracris nigripennis Lian&Zheng, 1984 | Xiahe, Gansu | 102.52 | 35.20 |
| Uvaroviola multispinosa Bey-Bienko, 1930 | Xiahe, Gansu | 102.52 | 35.20 |
| Gonghe County, Qinghai | 100.22 | 36.13 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Maqin, Qinghai | 100.24 | 34.48 | |
| Tongren County, Qinghai | 102.02 | 35.52 | |
| Ceracris nigricornis nigricornis Walker, 1870 | Motuo, Tibet | 95.33 | 29.33 |
| Zhouqu, Gansu | 104.37 | 33.79 | |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Ceracris xizangensis xizangensis Liu, 1981 | Tongmai, Bomi, Tibet | 95.08 | 30.10 |
| Yumei, Nyingchi, Tibet | 94.95 | 29.59 | |
| Zayü, Nyingchi, Tibet | 95.14 | 29.90 | |
| Ceracris versicolor (Brunner von Wattenwyl, 1893) | Motuo, Tibet | 95.33 | 29.33 |
| Ceracrisoides shannanensis Bi&Xia, 1987 | Shannan, Tibet | 91.77 | 29.24 |
| Anaptygus qinghaiensis Yin, 1984 | Zeku, Qinghai | 101.46 | 35.04 |
| Xunhua County, Qinghai | 102.49 | 35.85 | |
| Anaptygus yueliangwan Mao, Ren&Ou, 2011 | Deiqen Moon Bay, Yunnan | 99.31 | 28.20 |
| Anaptygus yulongensis Wang, Zheng&Lian, 2005 | Jade Dragon Snow Mountain, Lijiang, Yunnan | 100.17 | 27.10 |
| Anaptygus furculus Huo&Zheng | Xunhua County, Qinghai | 102.49 | 35.85 |
| Anaptygus longipennis Wang, Zheng&Lian, 2004 | Laojun Mountain, Jianchuan, Yunnan | 99.59 | 26.56 |
| Anaptygus uvarovi (Chang, 1937) | Zhongzha Town, Batang County, Sichuan | 99.17 | 29.50 |
| Ganzi County, Sichuan | 100.86 | 28.42 | |
| Luding County, Sichuan | 102.23 | 29.92 | |
| Kangding, Sichuan | 101.95 | 29.98 | |
| Arcyptera coreana Shiraki, 1930 | Gonghe, Qinghai | 100.62 | 36.28 |
| Maerkang County, Sichuan | 102.20 | 31.91 | |
| Leuconemacris litangensis (Yin, 1983) | Litang County, Sichuan | 100.27 | 30.00 |
| Leuconemacris xizangensis (Yin, 1984) | Markam, Tibet | 98.59 | 29.69 |
| Xiangcheng County, Sichuan | 100.05 | 29.12 | |
| Xiangcheng County, Sichuan | 99.75 | 29.12 | |
| Batang County, Sichuan | 99.11 | 30.01 | |
| Leuconemacris daochengensis Zheng, 1988 | Daocheng County, Sichuan | 100.30 | 29.04 |
| Daocheng County, Sichuan | 100.18 | 28.55 | |
| Leuconemacris longipennis Zheng, 1988 | Batang County, Sichuan | 99.11 | 30.01 |
| Leuconemacris breviptera (Yin, 1983) | Litang County, Sichuan | 100.27 | 30.00 |
| Yajiang, Sichuan | 101.01 | 30.03 | |
| Dechai Township, Yajiang, Sichuan | 100.76 | 29.67 | |
| Kangding, Sichuan | 101.95 | 30.16 | |
| Leuconemacris xiangchengensis Zheng, 1988 | Xiangcheng County, Sichuan | 99.80 | 28.93 |
| Liziping Township, Muli County, Sichuan | 100.29 | 27.97 | |
| Muli County, Sichuan | 100.86 | 28.42 | |
| Leuconemacris shalulishanensis Zheng, 1988 | Daocheng County, Sichuan | 100.30 | 29.04 |
| Leuconemacris hengduanshanensis Zheng, 1988 | Xiangcheng County, Sichuan | 99.80 | 28.93 |
| Leuconemacris xinlongensis Mao&Huang, 2023 | Tongxiao Township, Xinlong, Sichuan | 100.08 | 30.89 |
| Xinlong County, Sichuan | 99.90 | 30.85 | |
| Leuconemacris acuminata Mao&Huang, 2023 | Bowo Township, Muli County, Sichuan | 101.12 | 28.78 |
| Leuconemacris muliensis Mao&Huang 2023 | Bowo Township, Muli County, Sichuan | 101.12 | 28.78 |
| Muli County, Sichuan | 101.10 | 29.00 | |
| Pararcyptera microptera Ikonnikov, 1911 | Guide County, Qinghai | 101.43 | 36.04 |
| Bai Da Ban Prairie, Qilian Mountains, Gansu | 99.30 | 38.81 | |
| Dnopherula brevicornis (Bi&Xia, 1987) | Jilong, Tibet | 85.30 | 28.86 |
| Dnopherula taeniatus (Bolívar, 1902) | Chayu, Tibet | 97.47 | 28.66 |
| Dnopherula xizangensis (Zheng&Liping, 1986) | Chayu, Tibet | 97.47 | 28.66 |
| Dnopherula yuanmowensis (Zheng, 1977) | Aba County, Sichuan | 101.70 | 32.90 |
| Dnopherula neovittatus Otte, 1997 | Chayu, Tibet | 97.47 | 28.66 |
| Dnopherula sinensis (Uvarov, 1925) | Deqin County, Yunnan | 98.92 | 28.47 |
| Lanping County, Yunnan | 99.42 | 26.46 | |
| Ninglang County, Yunnan | 100.85 | 27.29 | |
| Eremippus persicus Uvarov, 19291 | Zanda, Tibet | 79.80 | 31.48 |
| Eremippus simplex (Eversmann, 1859) | Xining, Qinghai | 101.78 | 36.62 |
| Eremippus heimahoensis Zheng&Hang, 1974 | Hei Ma He Township, Gonghe County, Qinghai | 99.78 | 36.73 |
| Podismomorpha gibba Lian&Zheng, 1984 | Ma’ai, Luqu, Gansu | 102.52 | 35.20 |
| Squamopenna gansuensis Lian&Zheng, 1984 | Ganjia Town, Xiahe County, Gansu | 102.52 | 35.40 |
| Euchorthippus unicolor (Ikonnikov, 1913) | Jianzha, Qinghai | 102.03 | 35.94 |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Xining, Qinghai | 101.78 | 36.62 | |
| Euchorthippus weichowensis Chang, 1937 | Tongren County, Qinghai | 103.59 | 31.48 |
| Euchorthippus flexucarinatus Bi&Xia, 1987 | Qusongbu, Tibet | 80.73 | 31.13 |
| Lhünzhub County, Tibet | 92.46 | 28.41 | |
| Milin, Tibet | 94.21 | 29.22 | |
| Qusang Nama, Tibet | 82.24 | 32.06 | |
| Euchorthippus cheui Hsia, 1965 | Diebu County, Gansu | 103.22 | 34.06 |
| Luqu, Gansu | 102.49 | 34.59 | |
| Zhuoni, Gansu | 103.51 | 34.59 | |
| Lintan, Gansu | 103.35 | 34.69 | |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Euchorthippus yungningensis Cheng&Chiu, 1965 | Zhuoni, Gansu | 103.51 | 34.59 |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Eclipophleps xizangensis Liu, 1980 | Ngamring County, Tibet | 88.57 | 29.17 |
| Kangacris rufipes Yin, 1983 | Litang County, Sichuan | 100.27 | 30.00 |
| Kuiyong Gully, Danba County, Sichuan | 101.77 | 30.56 | |
| Garzê Town, Sichuan | 99.99 | 31.63 | |
| Kangding, Sichuan | 101.95 | 30.16 | |
| Pengta Township, Kangding, Sichuan | 102.29 | 30.43 | |
| Dechai, Yajiang, Sichuan | 101.35 | 29.95 | |
| Macrokangacris luteoarmilla Yin, 1983 | Batang County, Sichuan | 99.11 | 30.01 |
| Kangacrisoides jiangdaensis Zheng, Dong, Bai&Xu, 2013 | Jiangda County, Tibet | 98.22 | 31.50 |
| Chorthippus intermedius (Bey-Bienko, 1926) | Kagong Township, Jiangda County, Tibet | 98.14 | 31.44 |
| Guxiang, Bomi, Tibet | 95.50 | 29.91 | |
| Chayu, Tibet | 97.47 | 28.66 | |
| Basu, Tibet | 96.92 | 30.05 | |
| Gujing, Tibet | 97.22 | 29.15 | |
| Bomi, Tibet | 95.77 | 29.86 | |
| Riwoqe, Tibet | 96.60 | 31.21 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Zeku, Qinghai | 101.46 | 35.04 | |
| Tongren County, Qinghai | 102.02 | 35.52 | |
| Qilian County, Qinghai | 100.25 | 38.18 | |
| Menyuan Hui Autonomous County, Qinghai | 101.61 | 37.39 | |
| Datong Hui and Tu Autonomous County, Qinghai | 101.69 | 36.93 | |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Xunhua County, Qinghai | 102.49 | 35.85 | |
| Huzhu Tuzu Zizhixian, Qinghai | 101.96 | 36.84 | |
| Nagqu, Qinghai | 96.48 | 32.19 | |
| Maqu County, Gansu | 102.07 | 34.00 | |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Jiuzhaigou Valley, Sichuan | 104.24 | 33.26 | |
| Luhuo, Sichuan | 100.64 | 31.43 | |
| Kangding, Sichuan | 101.95 | 30.16 | |
| Batang County, Sichuan | 99.11 | 30.01 | |
| Chorthippus qingzangensis Yin, 1984 | Cháiwéi Township, Chamdo, Tibet Cháiwéi Township, Chamdo, TibeSangduo Town, Riwoqe, Tibet: Sangduo Town, Riwoqe, Tibet | 97.14 | 31.57 |
| Kagong Township, Jiangda County, Tibet | 98.14 | 31.44 | |
| Cháiwéi Township, Chamdo, TibeSangduo Town, Riwoqe, Tibet | 96.60 | 31.21 | |
| Ranwu Town, Basu, Tibet | 96.75 | 29.50 | |
| Riwoqe, Tibet | 96.60 | 31.21 | |
| Chamdo, Tibet | 97.17 | 31.15 | |
| Basu, Tibet | 96.92 | 30.05 | |
| Qilian County, Qinghai | 100.25 | 38.18 | |
| Ping’an County, Qinghai | 102.11 | 36.50 | |
| Yushu, Qinghai | 97.01 | 32.99 | |
| Xining, Qinghai | 101.78 | 36.62 | |
| Datong Hui and Tu Autonomous County, Qinghai | 101.69 | 36.93 | |
| Ganzi County, Sichuan | 99.99 | 31.62 | |
| Jiuzhaigou Valley, Sichuan | 104.24 | 33.26 | |
| Chorthippus latisulcus (Zheng&He, 1995) | Datong Hui and Tu Autonomous County, Qinghai | 101.69 | 36.93 |
| Haiyan, Qinghai | 101.00 | 36.90 | |
| Chorthippus dabanshanensis Chen, Zeng&Bao, 2011 | Tianjun County, Qinghai | 99.02 | 37.30 |
| Gonghe, Qinghai | 100.62 | 36.28 | |
| Menyuan Hui Autonomous County, Qinghai | 101.61 | 37.39 | |
| Huzhu Tuzu Zizhixian, Qinghai | 101.96 | 36.84 | |
| Daban Mountain, Qinghai | 102.09 | 37.06 | |
| Xunhua County, Qinghai | 102.49 | 35.85 | |
| Qilian County, Qinghai | 100.25 | 38.18 | |
| Chorthippus qilianshanensis Zheng&Xie, 2000 | Menyuan Hui Autonomous County, Qinghai | 101.61 | 37.39 |
| Tianjun County, Qinghai | 99.02 | 37.30 | |
| Gonghe, Qinghai | 100.62 | 36.28 | |
| Qilian County, Qinghai | 100.25 | 38.18 | |
| Huzhu Tuzu Zizhixian, Qinghai | 101.96 | 36.84 | |
| Xunhua County, Qinghai | 102.49 | 35.85 | |
| Chorthippus qinghaiensis Wang&Zheng, 1992 | Qumarlêb, Qinghai | 95.80 | 34.13 |
| Mǎnigān Gēng, Dégé, Sichuan | 99.20 | 31.92 | |
| Chorthippus jishishanensis Zheng&Xie, 2000 | Xunhua County, Qinghai | 102.49 | 35.85 |
| Chorthippus guideensis Zheng&al, 2009 | Gārdé Qiānhù Village, Qinghai | 101.55 | 36.29 |
| Chorthippus changtunensis Yin, 1984 | Beibeng, Motuo, Tibet | 95.17 | 28.91 |
| Chamdo, Tibet | 97.17 | 31.15 | |
| Jiāngdá, Chamdo, Tibet | 98.22 | 31.50 | |
| Ganzi County, Sichuan | 99.99 | 31.62 | |
| Batang County, Sichuan | 99.11 | 30.01 | |
| Chorthippus flexivenus (Liu, 1981) | Ranwu Lake, Basu, Tibet | 96.79 | 29.45 |
| Jilong County, Shigatse, Tibet | 85.30 | 28.86 | |
| Chorthippus chayuensis Yin, 1984 | Linzhi Town, Tibet | 94.48 | 29.57 |
| Chayu, Tibet | 97.47 | 28.66 | |
| Linzhi, Tibet | 94.36 | 29.65 | |
| Muli County, Sichuan | 100.86 | 28.42 | |
| Chorthippus albonemus Zheng&Tu, 1964 | Nagqu, Qinghai | 96.48 | 32.19 |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Guide County, Qinghai | 101.43 | 36.04 | |
| Xining, Qinghai | 101.78 | 36.62 | |
| Jianzha, Qinghai | 102.03 | 35.94 | |
| Xunhua County, Qinghai | 102.49 | 35.85 | |
| Huzhu Tuzu Zizhixian, Qinghai | 101.96 | 36.84 | |
| Ledu District, Qinghai | 102.40 | 36.48 | |
| Datong Hui and Tu Autonomous County, Qinghai | 101.69 | 36.93 | |
| Tongren County, Qinghai | 102.02 | 35.52 | |
| Ping’an County, Qinghai | 102.11 | 36.50 | |
| Ganzi County, Sichuan | 99.99 | 31.62 | |
| Muli County, Sichuan | 100.86 | 28.42 | |
| Bai Da Ban Prairie, Qilian Mountains, Gansu | 99.30 | 38.81 | |
| Maqu County, Gansu | 102.07 | 34.00 | |
| Luqu County, Gansu | 102.49 | 34.59 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Lintan County, Gansu | 103.35 | 34.69 | |
| Zhuoni County, Gansu | 103.51 | 34.59 | |
| Zhouqu, Gansu | 104.37 | 33.79 | |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Chorthippus chinensis (Tarbinsky, 1927) | Xiahe, Gansu | 102.52 | 35.20 |
| Ganzi County, Sichuan | 99.99 | 31.62 | |
| Muli County, Sichuan | 100.86 | 28.42 | |
| Jiuzhaigou Valley, Sichuan | 103.95 | 33.30 | |
| Chorthippus hsiai Cheng&Tu, 1964 | Litang County, Sichuan | 100.27 | 30.00 |
| Xining, Qinghai | 101.78 | 36.62 | |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Gonghe County, Qinghai | 100.62 | 36.28 | |
| Xinghai, Qinghai | 99.99 | 35.59 | |
| Jianzha, Qinghai | 102.03 | 35.94 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Guide County, Qinghai | 101.43 | 36.04 | |
| Tongren County, Qinghai | 102.02 | 35.52 | |
| Maqu County, Gansu | 102.07 | 34.00 | |
| Luqu County, Gansu | 102.49 | 34.59 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Lintan County, Gansu | 103.35 | 34.69 | |
| Zhuoni County, Gansu | 103.51 | 34.59 | |
| Zhouqu, Gansu | 104.37 | 33.79 | |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Chorthippus longisonus (Li&Yin, 1987) | Xunhua County, Qinghai | 102.49 | 35.85 |
| Litang, Sichuan | 100.12 | 29.83 | |
| Chorthippus markamensis (Yin, 1984) | Markam, Tibet | 98.59 | 29.69 |
| Jiangda County, Tibet | 98.22 | 31.50 | |
| Chorthippus biguttulus biguttulus (Linnaeus, 1758) | Chamdo, Tibet | 97.17 | 31.15 |
| Riwoqe, Tibet | 96.60 | 31.21 | |
| Chayab, Tibet | 97.57 | 30.66 | |
| Kagong Township, Jiangda County, Tibet | 98.14 | 31.44 | |
| Zhiduo, Qinghai | 95.62 | 33.84 | |
| Yushu, Qinghai | 97.01 | 32.99 | |
| Gonghe County, Qinghai | 100.62 | 36.28 | |
| Xinghai, Qinghai | 99.99 | 35.59 | |
| Tianjun County, Qinghai | 99.02 | 37.30 | |
| Luqu, Gansu | 102.49 | 34.39 | |
| Zhuoni, Gansu | 103.51 | 34.59 | |
| Chorthippus chapini Chang, 1939 | Kangding, Sichuan | 101.95 | 30.16 |
| Luding County, Sichuan | 102.23 | 29.92 | |
| Garzê Town, Sichuan | 99.99 | 31.63 | |
| Chorthippus grahami Chang, 1937 | Maerkang County, Sichuan | 102.20 | 31.91 |
| Xiaojin, Sichuan | 102.36 | 31.00 | |
| Jinchuan County, Sichuan | 102.06 | 31.48 | |
| Batang County, Sichuan | 99.11 | 30.01 | |
| Chorthippus brevipterus Yin, 1984 | Aba County, Sichuan | 101.70 | 32.90 |
| Mai Xiu Forest Farm, Zeku County, Qinghai | 101.93 | 35.26 | |
| Jiuzhi, Qinghai | 101.48 | 33.43 | |
| Chorthippus fallax (Zubovsky), 1899 | Xiahe, Gansu | 102.52 | 35.20 |
| Maqu County, Gansu | 102.07 | 34.00 | |
| Luqu County, Gansu | 102.49 | 34.59 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Lintan County, Gansu | 103.35 | 34.69 | |
| Zhuoni County, Gansu | 103.51 | 34.59 | |
| Zhouqu, Gansu | 104.37 | 33.79 | |
| Bai Da Ban Prairie, Qilian Mountains, Gansu | 99.30 | 38.81 | |
| Ganzi County, Sichuan | 99.99 | 31.62 | |
| Muli County, Sichuan | 100.86 | 28.42 | |
| Daotang River, Gonghe County, Qinghai | 100.97 | 36.40 | |
| Hei Ma He Township, Gonghe County, Qinghai | 99.78 | 36.73 | |
| Gonghe Tiebujia, Qinghai | 99.58 | 37.03 | |
| Nian Di Township, Gonghe County, Qinghai | 100.39 | 36.38 | |
| Santa La, Gonghe County, Qinghai | 100.27 | 36.06 | |
| Heika Mountain, Xinghai County, Qinghai | 100.05 | 35.78 | |
| Guinan County, Qinghai | 100.74 | 35.57 | |
| Mùgé Tān, Guinan County, Qinghai | 100.83 | 35.64 | |
| Guinan Mugetan, Qinghai | 100.48 | 35.34 | |
| Mai Xiu Forest Farm, Zeku County, Qinghai | 101.93 | 35.26 | |
| Xining, Qinghai | 101.78 | 36.62 | |
| Huangyuan County, Qinghai | 101.26 | 36.68 | |
| Jianzha, Qinghai | 102.03 | 35.94 | |
| Ping’an County, Qinghai | 102.11 | 36.50 | |
| Ledu District, Qinghai | 102.40 | 36.48 | |
| Guide County, Qinghai | 101.43 | 36.04 | |
| Maqin, Qinghai | 100.24 | 34.48 | |
| Menyuan Hui Autonomous County, Qinghai | 101.61 | 37.39 | |
| Xunhua County, Qinghai | 102.49 | 35.85 | |
| Datong Hui and Tu Autonomous County, Qinghai | 101.69 | 36.93 | |
| Tongren County, Qinghai | 102.02 | 35.52 | |
| Huzhu Tuzu Zizhixian, Qinghai | 101.96 | 36.84 | |
| Chorthippus brunneus huabeiensis Xia&Jin, 1982 | Ledu District, Qinghai | 102.40 | 36.48 |
| Jianzha, Qinghai | 102.03 | 35.94 | |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Bai Da Ban Prairie, Qilian Mountains, Gansu | 99.30 | 38.81 | |
| Maqu County, Gansu | 102.07 | 34.00 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Lintan County, Gansu | 103.35 | 34.69 | |
| Zhuoni County, Gansu | 103.51 | 34.59 | |
| Zhouqu, Gansu | 104.37 | 33.79 | |
| Luqu, Gansu | 102.49 | 34.59 | |
| Kangding, Sichuan | 101.95 | 30.16 | |
| Maerkang County, Sichuan | 102.20 | 31.91 | |
| Chorthippus maerkangensis (Zheng, 1980) | Maerkang County, Sichuan | 102.20 | 31.91 |
| Jinchuan County, Sichuan | 102.06 | 31.48 | |
| Chorthippus xiangchengensis (Liu, 1985) | Xiangcheng County, Sichuan | 99.80 | 28.93 |
| Chorthippus louguanensis Zheng&Tu, 1964 | Jinchuan County, Sichuan | 102.06 | 31.48 |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Luqu County, Gansu | 102.49 | 34.59 | |
| Zhuoni County, Gansu | 103.51 | 34.59 | |
| Maqu County, Gansu | 102.48 | 34.59 | |
| Lintan County, Gansu | 103.35 | 34.69 | |
| Chorthippus occidentalis Xia&Jin, 1982 | Maerkang County, Sichuan | 102.20 | 31.91 |
| Xiaojin, Sichuan | 102.36 | 31.00 | |
| Kangding, Sichuan | 101.95 | 30.16 | |
| Jinchuan County, Sichuan | 102.06 | 31.48 | |
| Chorthippus squamopennis Zheng, 1980 | Jinchuan County, Sichuan | 102.06 | 31.48 |
| Chorthippus dubius (Zubovski, 1898) | Daotang River, Gonghe County, Qinghai | 100.97 | 36.40 |
| Hei Ma He Township, Gonghe County, Qinghai | 99.78 | 36.73 | |
| Ziketan, Xinghai County, Qinghai | 99.98 | 35.59 | |
| Batan, Qinghai | 101.31 | 35.32 | |
| Guinan Mugetan, Qinghai | 100.48 | 35.34 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Tianjun County, Qinghai | 99.02 | 37.30 | |
| Guide County, Qinghai | 101.43 | 36.04 | |
| Bai Da Ban Prairie, Qilian Mountains, Gansu | 99.30 | 38.81 | |
| Maqu County, Gansu | 102.07 | 34.00 | |
| Luqu County, Gansu | 102.49 | 34.59 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Lintan County, Gansu | 103.35 | 34.69 | |
| Zhuoni County, Gansu | 103.51 | 34.59 | |
| Zhouqu, Gansu | 104.37 | 33.79 | |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Chorthippus xiningensis Zheng&Chen, 2001 | Xishan, Xining, Qinghai | 101.57 | 36.49 |
| Chorthippus haibeiensis Zheng&Chen, 2001 | Dauyinshan, Qilian County, Qinghai | 100.21 | 38.15 |
| Chorthippus helanshanensis Zheng, 1999 | Nagqu, Qinghai | 96.49 | 32.21 |
| Chorthippus rufitibis (Zheng, 1989) | Ping’an County, Qinghai | 102.11 | 36.50 |
| Huzhu Tuzu Zizhixian, Qinghai | 101.96 | 36.84 | |
| Xunhua County, Qinghai | 102.49 | 35.85 | |
| Chorthippus brunneus brunneus (Thunbeng, 1815) | Jianzha, Qinghai | 102.03 | 35.94 |
| Jiangda County, Tibet | 98.22 | 31.50 | |
| Riwoqe, Tibet | 96.60 | 31.21 | |
| Chamdo, Tibet | 97.17 | 31.15 | |
| Bomi, Tibet | 95.77 | 29.86 | |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Chorthippus xunhuaensis Zheng&Xie, 2000 | Xunhua County, Qinghai | 102.49 | 35.85 |
| Chorthippus leduensis Zheng&Xin, 1999 | Ledu District, Qinghai | 102.40 | 36.48 |
| Chorthippus gongbuensis Liang&Zheng, 1991 | Gongbujiangda, Tibet | 93.24 | 29.89 |
| Chorthippus mollis (Charpentieri, 1825) | Jiangda County, Tibet | 98.22 | 31.50 |
| Ganzi County, Sichuan | 99.99 | 31.62 | |
| Chorthippus amplintersitus Liu, 1981 | Riwoqe, Tibet | 96.60 | 31.21 |
| Xiangcheng County, Sichuan | 99.72 | 29.11 | |
| Chorthippus rubensabdomenis Liu, 1981 | Jilong, Tibet | 85.30 | 28.86 |
| Mai Xiu Forest Farm, Zeku County, Qinghai | 101.93 | 35.26 | |
| Daocheng County, Sichuan | 100.29 | 28.72 | |
| Chorthippus hemipeterus Uvarov, 1926 | Qusong, Zanda, Tibet | 79.18 | 32.20 |
| Chorthippus flavabdomenis Liu, 1981 | Jitang, Chayab, Tibet | 97.35 | 30.74 |
| Chorthippus tibetanus Uvarov, 1935 | Shannan City, Tibet | 91.77 | 29.24 |
| Linzhi, Tibet | 94.36 | 29.65 | |
| Chamdo, Tibet | 97.17 | 31.15 | |
| Bomi, Tibet | 95.77 | 29.86 | |
| Chorthippus separatanus Liu, 1981 | Jigong, Chayu, Tibet | 97.07 | 28.84 |
| Chorthippus aethalinus (Zubovski, 1899) | Xiahe, Gansu | 102.52 | 35.20 |
| Chorthippus laifoveatus Xia&Jin, 1982 | Motuo, Tibet | 95.33 | 29.33 |
| Chorthippus dichrous (Eversmann, 1859) | Huzhu Tuzu Zizhixian, Qinghai | 101.96 | 36.84 |
| Chorthippus huchengensis Xia&Jin, 1982 | Datong, Xining, Qinghai | 101.69 | 36.93 |
| Qilian County, Qinghai | 100.25 | 38.18 | |
| Geermu, Qinghai | 94.90 | 36.40 | |
| Guide County, Qinghai | 101.43 | 36.04 | |
| Jianzha, Qinghai | 102.03 | 35.94 | |
| Tongren County, Qinghai | 102.02 | 35.52 | |
| Lazikou, Diebu County, Gansu | 103.90 | 34.06 | |
| Chorthippus daitongensis Huo, 1994 | Haiyan, Qinghai | 101.00 | 36.90 |
| Chorthippus fusigeniculatus Zheng&Men, 2008 | Chamdo, Tibet | 97.17 | 31.15 |
| Chorthippus xueshanensis Zheng&Mao, 1997 | Lanping County, Yunnan | 99.42 | 26.46 |
| Shangri-La, Yunnan | 99.74 | 27.84 | |
| Laojun Mountain, Jianchuan, Yunnan | 99.59 | 26.56 | |
| Ninglang County, Yunnan | 100.85 | 27.29 | |
| Hengduan Mountains, Deqin County, Yunnan | 98.99 | 27.58 | |
| Yanyuan County, Sichuan | 101.51 | 27.43 | |
| Chorthippus deqinensis Liu, 1984 | Deqin County, Yunnan | 98.92 | 28.47 |
| Chorthippus nemus Liu, 1984 | Shangri-La, Yunnan | 99.74 | 27.84 |
| Chongjiang River, Shangri-La, Yunnan | 100.01 | 27.27 | |
| Deqin County, Yunnan | 98.92 | 28.47 | |
| Chorthippus yajiangensis Zheng&Shi, 2007 | Dechai, Yajiang, Sichuan | 101.35 | 29.95 |
| Chorthippus kangdingensis Zheng&Shi, 2007 | Pengta, Kangding, Sichuan | 102.29 | 30.43 |
| Chorthippus neipopennis Xia&Jin, 1982 | Xi’e Luo, Sichuan | 100.72 | 30.02 |
| Chorthippus anomopterus Liu, 1984 | Deqin County, Yunnan | 98.92 | 28.47 |
| Chorthippus yanyuanensis Jin&Lin, 1982–1983 | Yanyuan County, Sichuan | 101.39 | 27.69 |
| Hongxing Township, Ruoergai County, Sichuan | 102.74 | 34.10 | |
| Omocestus cuonaensis Yin, 1984 | Qingduo Township, Bomi, Tibet | 95.60 | 30.06 |
| Guxiang, Bomi, Tibet | 95.50 | 29.91 | |
| Ranwu Town, Basu, Tibet | 96.75 | 29.50 | |
| Milin, Tibet | 94.21 | 29.22 | |
| Nyingchi, Tibet | 94.36 | 29.65 | |
| Lang County, Tibet | 93.07 | 29.05 | |
| Cona, Tibet | 91.96 | 28.00 | |
| Omocestus gonggarensis Zheng&Chen, 1995 | Gonggar County, Tibet | 90.98 | 29.29 |
| Omocestus haemorrhoidalis (Charpentier, 1825) | Kagong Township, Jiangda County, Tibet | 98.14 | 31.44 |
| Chamdo, Tibet | 97.17 | 31.15 | |
| Riwoqe, Tibet | 96.60 | 31.21 | |
| Datong County, Xining, Qinghai | 101.69 | 36.93 | |
| Yushu, Qinghai | 97.01 | 32.99 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Tongren County, Qinghai | 102.02 | 35.52 | |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Zeku, Qinghai | 101.46 | 35.04 | |
| Qilian County, Qinghai | 100.25 | 38.18 | |
| Litang County, Sichuan | 100.27 | 30.00 | |
| Ri’erlang Mountain, Ruoergai County, Sichuan | 102.79 | 34.03 | |
| Mǎnigān Gēng, Dégé, Sichuan | 99.20 | 31.92 | |
| Ganzi County, Sichuan | 99.99 | 31.62 | |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Lintan, Gansu | 103.53 | 34.79 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Zhuoni County, Gansu | 103.51 | 34.59 | |
| Maqu County, Gansu | 102.07 | 34.00 | |
| Luqu, Gansu | 102.49 | 34.59 | |
| Omocestus avellaeusitibia Zheng, Dong&Xu, 2013 | Ganzi County, Sichuan | 99.99 | 31.62 |
| Omocestus petraeus (Brisout de Barneville, 1856) | Dechai, Yajiang, Sichuan | 101.35 | 29.95 |
| Menyuan Hui Autonomous County, Qinghai | 101.61 | 37.39 | |
| Menyuan Hui Autonomous County, Qinghai | 101.62 | 37.37 | |
| Omocestus nyalamus Xia, 1981 | Damxung County, Tibet | 91.10 | 30.49 |
| Nyalam, Tibet | 85.98 | 28.16 | |
| Dingri County, Tibet | 87.12 | 28.66 | |
| Lazi County, Tibet | 87.64 | 29.08 | |
| Omocestus megaoculus Yin, 1984 | Nyalam, Tibet | 86.04 | 28.52 |
| Markam, Tibet | 98.59 | 29.69 | |
| Omocestus motuoensis Yin, 1984 | Motuo, Tibet | 95.33 | 29.33 |
| Milin, Tibet | 94.21 | 29.22 | |
| Bomi, Tibet | 95.77 | 29.86 | |
| Delomagnigou, Sichuan | 99.20 | 31.92 | |
| Omocestus pinanensis Zheng&Xie, 2001 | Ping’an County, Qinghai | 102.11 | 36.50 |
| Omocestus qinghaihuensis Zheng&Xie, 2001 | South Shore of Qinghai Lake, Qinghai | 100.54 | 36.56 |
| Tianjun County, Qinghai | 99.02 | 37.30 | |
| Gonghe, Qinghai | 100.62 | 36.28 | |
| Guide County, Qinghai | 101.43 | 36.04 | |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Huzhu Tuzu Zizhixian, Qinghai | 101.96 | 36.84 | |
| Omocestus hingstoni Uvarov, 1925 | Dongjiu, Nyingchi, Tibet | 94.80 | 29.94 |
| Mount Qomolangma, Tibet | 86.86 | 28.13 | |
| Omocestus enitor Uvarov, 1924 | Lhünzhub County, Tibet | 92.46 | 28.41 |
| Omocestus laojunshanensis Mao&Xu, 2004 | Laojun Mountain, Jianchuan, Yunnan | 99.59 | 26.56 |
| Jade Dragon Snow Mountain, Lijiang, Yunnan | 100.18 | 27.10 | |
| Nivisacris zhongdianensis Liu, 1984 | Shangri-La, Yunnan | 99.74 | 27.84 |
| Saxetophilus qinghaiensis Zheng, Chen&Li, 2012 | Kunlun Mountains, Xinjiang | 81.67 | 35.47 |
| Qumarlêb, Qinghai | 95.80 | 34.13 | |
| Hei Ma He Township, Gonghe County, Qinghai | 99.78 | 36.73 | |
| Nian Di Township, Gonghe County, Qinghai | 100.39 | 36.38 | |
| Wenquan Township, Xinghai, Qinghai | 99.67 | 35.22 | |
| Heika Mountain, Xinghai County, Qinghai | 100.05 | 35.78 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Maqin, Qinghai | 100.24 | 34.48 | |
| Qilian County, Qinghai | 100.25 | 38.18 | |
| Xining, Qinghai | 101.78 | 36.62 | |
| Jianzha, Qinghai | 102.03 | 35.94 | |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Guide County, Qinghai | 101.43 | 36.04 | |
| Saxetophilus petulans Umnov, 1930 | Luqu, Gansu | 102.49 | 34.59 |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Saxetophilus gansuensis Wang, Zheng&Lian, 2006 | Luqu, Gansu | 102.49 | 34.59 |
| Oreoptygonotus chinghaiensis (Zheng&Hang, 1974) | Madoi County, Golog Tibetan Autonomous Prefecture, Qinghai | 98.21 | 34.92 |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Hei Ma He Township, Gonghe County, Qinghai | 99.78 | 36.73 | |
| Heka Mountain, Xinghai, Qinghai | 100.05 | 35.78 | |
| Tongren County, Qinghai | 102.02 | 35.52 | |
| Zeku, Qinghai | 101.46 | 35.04 | |
| Henan, Qinghai | 101.62 | 34.73 | |
| Maqin, Qinghai | 100.24 | 34.48 | |
| Tianjun County, Qinghai | 99.02 | 37.30 | |
| Oreoptygonotus tibetanus Tarbinsky, 1927 | Guinan County, Qinghai | 100.75 | 35.59 |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Eling Lake, Qinghai | 97.74 | 34.92 | |
| Zaling Lake, Qinghai | 97.91 | 35.08 | |
| Madoi County, Qinghai | 98.21 | 34.91 | |
| Qumarlêb, Qinghai | 95.80 | 34.13 | |
| Oreoptygonotus brachypterus Yin, 1984 | Heka Mountain, Xinghai, Qinghai | 100.05 | 35.78 |
| Oreoptygonotus robustus Yin, 1984 | Zeku, Qinghai | 101.46 | 35.04 |
| Henan, Qinghai | 101.62 | 34.73 | |
| Jiangda County, Tibet | 98.10 | 31.80 | |
| Oreoptygonotus nigrofasciatus Yin, 1979 | Gaize County, Tibet | 84.68 | 32.24 |
| Oreoptygonotus belonocercus Liu, 1981 | Cuomei County, Tibet | 91.43 | 28.42 |
| Ruganotus rufipes Yin, 1979 | Pulan County, Tibet | 81.18 | 30.29 |
| Ptygonotus sichuanensis Zheng, 1983 | Jiuzhaigou Valley, Sichuan | 104.24 | 33.26 |
| Lixian County, Sichuan | 103.16 | 31.44 | |
| Qingzhenzhan Mountain, Gade County, Qinghai | 100.12 | 34.17 | |
| Batang, Yushu, Qinghai | 97.15 | 32.83 | |
| Ptygonotus semenovi semenovi Tarbinsky, 1927 | Songpan County, Sichuan | 103.60 | 32.64 |
| Ptygonotus tarbinskyi Uvarov, 1930 | Songpan County, Sichuan | 103.60 | 32.64 |
| Ma’ai, Luqu, Gansu | 102.52 | 35.20 | |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Diebu County, Gansu | 103.21 | 34.05 | |
| Ptygonotus gurneyi Chang, 1937 | Kangding, Sichuan | 101.95 | 30.16 |
| Dechai, Yajiang, Sichuan | 101.35 | 29.95 | |
| Ptygonotus chinghaiensis Ying, 1974 | Tongde, Qinghai | 100.58 | 35.25 |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Ptygonotus hocashanensis Cheng&Hang, 1974 | Heika Mountain, Xinghai County, Qinghai | 100.05 | 35.78 |
| Luqu, Gansu | 102.49 | 34.59 | |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Maqu County, Gansu | 102.07 | 34.00 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Ptygonotus brachypterus Yin, 1974 | Yushu, Qinghai | 97.01 | 32.99 |
| Zhiduo, Qinghai | 95.62 | 33.84 | |
| Amdo, Tibet | 91.68 | 32.26 | |
| Ptygonotus gansuensis Zheng&Chang, 1994 | Diebu County, Gansu | 103.22 | 34.06 |
| Ptygonotus semenovi antennatus Mistshenko, 1951 | Luqu, Gansu | 102.49 | 34.59 |
| Ptygonotus rangkouensis Zheng&Xu, 2009 | Raokou Township, Sichuan | 102.50 | 32.21 |
| Ptygonotus anurous Zheng&Xu, 2009 | Heishui County, Sichuan | 102.99 | 32.06 |
| Asulconotoides sichuanensis Liu, 1984 | Daocheng County, Sichuan | 100.10 | 29.00 |
| Haizi Mountain, Litang, Sichuan | 100.10 | 29.38 | |
| Hengduan Mountains, Yunnan | 99.00 | 27.50 | |
| Asulconotoides asulcata (Zheng, 1988) | Batang County, Sichuan | 99.11 | 30.01 |
| Asulconotoides microptera (Zheng, 1988) | Batang County, Sichuan | 99.11 | 30.01 |
| Asulconotus chinghaiensis Ying, 1974 | Duocai, Zhiduo, Qinghai | 95.42 | 33.81 |
| Gande County, Guoluo, Qinghai | 99.90 | 33.97 | |
| Yushu, Qinghai | 97.01 | 32.99 | |
| Jiuzhi, Qinghai | 101.48 | 33.43 | |
| Qumarlêb, Qinghai | 95.80 | 34.13 | |
| Maqin, Qinghai | 100.24 | 34.48 | |
| Zadoi, Qinghai | 95.30 | 32.89 | |
| Aba County, Sichuan | 101.70 | 32.90 | |
| Asulconotus jiuzhiensis Yin, Zheng&Yin, 2012 | Jiuzhi, Qinghai | 101.48 | 33.43 |
| Asulconotus kozlovi Mistshenko, 1981 | Yequ River, Yushu, Qinghai | 95.57 | 34.22 |
| Madoi County, Qinghai | 98.21 | 34.91 | |
| Qumarlêb, Qinghai | 95.80 | 34.13 | |
| Asonus brachypterus (Ying, 1974) | Zhiduo, Qinghai | 95.61 | 33.87 |
| Racecourse, Yushu, Qinghai | 96.96 | 33.00 | |
| Duocai, Zhiduo, Qinghai | 95.42 | 33.81 | |
| Zhiduo, Qinghai | 95.62 | 33.84 | |
| Yushu, Qinghai | 97.01 | 32.99 | |
| Amdo, Tibet | 91.68 | 32.26 | |
| Nagqu City, Tibet | 92.05 | 31.48 | |
| Riwoqe, Tibet | 96.60 | 31.21 | |
| Asonus amplifurculus Chen, Zheng&Zeng, 2010 | Gārdé Qiānhù Village, Qinghai | 101.55 | 36.29 |
| Asonus microfurculus Yin, 1984 | Kagila Mountain, Chamdo, Tibet | 96.40 | 31.60 |
| Asonus parvoculus Xie & Liu, 1998 | Muli County, Sichuan | 101.28 | 27.93 |
| Asonus qinghaiensis Liu, 1986 | Yushu, Qinghai | 97.01 | 32.99 |
| Asonus longisulcus Yin, 1984 | Riwoqe, Tibet | 96.60 | 31.21 |
| Pseudoasonus yushuensis Yin, 1982 | Qumarlêb, Qinghai | 95.80 | 34.13 |
| Zadoi, Qinghai | 95.30 | 32.89 | |
| Nagqu, Qinghai | 96.49 | 32.21 | |
| Gade, Qinghai | 99.90 | 33.97 | |
| Maqin, Qinghai | 100.24 | 34.48 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Jiuzhi, Qinghai | 101.48 | 33.43 | |
| Yushu, Qinghai | 97.01 | 32.99 | |
| Duocai, Zhiduo, Qinghai | 95.42 | 33.81 | |
| Pseudoasonus kangdingensis Yin, 1983 | Kangding, Sichuan | 101.95 | 30.16 |
| Xinduqiao, Sichuan | 101.49 | 30.05 | |
| Pseudoasonus baiyuensis Zheng, 1990 | Baiyu County, Sichuan | 98.82 | 31.21 |
| Hengduan Mountains, Yunnan | 99.00 | 27.50 | |
| Pseudoasonus orthomarginis Zheng, Chen&Lin, 2012 | Mozhugongka County, Tibet | 95.42 | 33.81 |
| Transtympanacris xueshanensis Lian&Zheng, 1985 | Kangding, Sichuan | 101.95 | 30.16 |
| Transtympanacris yajiangensis Zheng, 1995 | Yajiang, Sichuan | 101.01 | 30.03 |
| Hypernephia everesti Uvarov, 1922 | Mount Qomolangma, Tibet | 86.86 | 28.13 |
| Hypernephia xizangensis Zheng, 1980 | Gar County, Tibet | 80.09 | 32.49 |
| Atympanum nigrofasciatum Yin, 1984 | Yangbajing, Tibet | 90.54 | 30.10 |
| Amdo, Tibet | 91.68 | 32.26 | |
| Gaize County, Tibet | 84.05 | 32.30 | |
| Damxung County, Tibet | 91.09 | 30.47 | |
| Atympanum gonggarensis Zheng&Chen, 1995 | Gonggar County, Tibet | 90.98 | 29.29 |
| Atympanum belonocercum (Liu, 1981) | Karila Mountain Pass, Cuomei, Tibet | 91.48 | 28.51 |
| Yangbajing, Tibet | 90.54 | 30.10 | |
| Atympanum antennatum Yin, 1984 | Nedong County, Tibet | 91.76 | 29.22 |
| Atympanum carinotus (Yin, 1979) | Gaize County, Tibet | 84.05 | 32.30 |
| Shenzha County, Tibet | 88.71 | 30.93 | |
| Atympanum comainensis (Liu), 1981 | Karila Mountain Pass, Cuomei, Tibet | 91.48 | 28.51 |
| Oknosacris gyirogensis Liu, 1981 | Cuoluo Long, Jilong, Tibet | 85.42 | 29.13 |
| Hebetacris amplinota Liu, 1981 | Cuoluo Long, Jilong, Tibet | 85.42 | 29.13 |
| Dysanema brevitibia Niu, Zheng&Lin, 2015 | Cuoqin County, Tibet | 85.15 | 31.02 |
| Dysanema irvinei Uvarov, 1925 | Mount Qomolangma, Tibet | 86.86 | 28.13 |
| Dingjie County, Tibet | 87.76 | 28.37 | |
| Gamba County, Tibet | 88.51 | 28.27 | |
| Lhünzhub County, Lin Karzong, Tibet | 90.27 | 28.82 | |
| Lhünzhub County, Lin Karzong, Tibet | 89.15 | 27.72 | |
| Dysanema malloryi Uvarov, 1925 | Lhünzhub County, Lin Karzong, Tibet | 89.15 | 27.72 |
| Dysanema magna Zheng, Lin, Zhang&Zeng, 2014 | Dingri County, Tibet | 87.12 | 28.66 |
| Stristernum ratogensis Liu, 1981 | Rutog County, Dogmarsum, Tibet | 80.38 | 33.71 |
| Aeropus licenti Chang, 1939 | Heika Mountain, Xinghai County, Qinghai | 100.05 | 35.78 |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Qilian County, Qinghai | 100.25 | 38.18 | |
| Dongliugou, Sunan, Gansu | 99.64 | 38.77 | |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Ranwu Town, Basu, Tibet | 96.75 | 29.50 | |
| Zongbala Mountain, Markam, Tibet | 98.62 | 29.72 | |
| Cháiwéi Township, Chamdo, TibeSangduo Town, Riwoqe, Tibet | 96.60 | 31.21 | |
| Jiangtang Village, Sangri County, Tibet | 91.98 | 29.25 | |
| Gartok Town, Markam, Tibet | 98.59 | 29.68 | |
| Ganzi County, Sichuan | 99.99 | 31.62 | |
| Aeropus tibetanus Uvarov, 1935 | Ranwu Town, Tibet | 96.75 | 29.51 |
| Jida Township, Basu, Tibet | 96.68 | 29.91 | |
| Kagong Township, Jiangda County, Tibet | 98.14 | 31.44 | |
| Gartok Town, Markam, Tibet | 98.59 | 29.68 | |
| Zuogong, Tibet | 97.84 | 29.67 | |
| Markam, Tibet | 98.59 | 29.69 | |
| Jiangda County, Tibet | 98.22 | 31.50 | |
| Riwoqe, Tibet | 96.60 | 31.21 | |
| Basu, Tibet | 96.92 | 30.05 | |
| Xinghai, Qinghai | 99.99 | 35.59 | |
| Gonghe, Qinghai | 100.62 | 36.28 | |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Gomphoceroides flavutibia Zheng&Chen, 2001 | Menyuan Hui Autonomous County, Qinghai | 101.59 | 37.32 |
| Dasyhippus barbipes (Fischer von Waldheim, 1846) | Daotang River, Gonghe County, Qinghai | 100.97 | 36.40 |
| Guinan County, Qinghai | 100.75 | 35.59 | |
| Xinghai, Qinghai | 99.99 | 35.59 | |
| Myrmeleotettix palpalis Zubovski, 1900 | Quanji, Gangcha County, Qinghai | 99.88 | 37.27 |
| Heika Mountain, Xinghai County, Qinghai | 100.05 | 35.78 | |
| Dahe Dam, Xinghai, Qinghai | 100.01 | 35.52 | |
| Hei Ma He Township, Gonghe County, Qinghai | 99.78 | 36.73 | |
| Gonghe Tiebujia, Qinghai | 99.58 | 37.03 | |
| Santa La, Gonghe County, Qinghai | 100.27 | 36.06 | |
| Nian Di Township, Gonghe County, Qinghai | 100.39 | 36.38 | |
| Batan, Qinghai | 101.31 | 35.32 | |
| Mùgé Tān, Guinan County, Qinghai | 100.83 | 35.64 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Zhiduo, Qinghai | 95.61 | 33.87 | |
| Tongren County, Qinghai | 102.02 | 35.52 | |
| Qilian County, Qinghai | 100.25 | 38.18 | |
| Yushu, Qinghai | 97.01 | 32.99 | |
| Menyuan Hui Autonomous County, Qinghai | 101.61 | 37.39 | |
| Haiyan, Qinghai | 101.00 | 36.90 | |
| Geermu, Qinghai | 94.90 | 36.40 | |
| Zhiduo, Qinghai | 95.62 | 33.84 | |
| Tianjun County, Qinghai | 99.02 | 37.30 | |
| Xining, Qinghai | 101.78 | 36.62 | |
| Quanji, Gangca County, Qinghai | 99.88 | 37.27 | |
| Bai Da Ban Prairie, Qilian Mountains, Gansu | 99.30 | 38.81 | |
| Diebu County, Gansu | 103.22 | 34.06 | |
| Maqu County, Gansu | 102.07 | 34.00 | |
| Zhuoni County, Gansu | 103.51 | 34.59 | |
| Xiahe, Gansu | 102.52 | 35.20 | |
| Myrmeleotettix kunlunensis Huang, 1987 | Dongkunlun Karl Cave, Ruoqiang County, Xinjiang | 89.03 | 38.20 |
| Aeropedelloides tibetanus Uvarov, 1939 | Zuogong, Tibet | 97.84 | 29.67 |
| Bianba, Tibet | 94.71 | 30.93 | |
| Basu, Tibet | 96.92 | 30.05 | |
| Aeropedelloides altissimus Liu, 1981 | Yushu, Qinghai | 97.01 | 32.99 |
| Riwoqe, Tibet | 96.60 | 31.21 | |
| Basu, Tibet | 96.92 | 30.05 | |
| Chayab, Tibet | 97.57 | 30.66 | |
| Aeropedelloides nigrocaudus Liu, 1981 | Chayab, Tibet | 97.57 | 30.66 |
| Aeropedellus zadoensis Yin, 1984 | Zadoi, Qinghai | 97.01 | 32.99 |
| Yushu, Qinghai | 97.01 | 32.99 | |
| Qumarlêb, Qinghai | 95.80 | 34.13 | |
| Nagqu, Qinghai | 96.49 | 32.21 | |
| Gade, Qinghai | 99.90 | 33.97 | |
| Maqin, Qinghai | 100.24 | 34.48 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Jiuzhi, Qinghai | 101.48 | 33.43 | |
| Aeropedellus changtunensis (Yin, 1984) | Chamdo, Tibet | 97.17 | 31.15 |
| Aeropedellus prominemarginis Zheng, 1981 | Yushu, Qinghai | 97.01 | 32.99 |
| Zhiduo, Qinghai | 95.62 | 33.84 | |
| Qumarlêb, Qinghai | 95.80 | 34.13 | |
| Zadoi, Qinghai | 95.30 | 32.89 | |
| Nagqu, Qinghai | 96.49 | 32.21 | |
| Gade, Qinghai | 99.90 | 33.97 | |
| Maqin, Qinghai | 100.24 | 34.48 | |
| Tongde, Qinghai | 100.58 | 35.25 | |
| Jiuzhi, Qinghai | 101.48 | 33.43 | |
| Orinhippus tibetanus Uvarov, 1921 | Bailang County, Tibet | 89.26 | 29.11 |
| Lumbu Monastery, Jiangzi County, Tibet | 89.73 | 28.86 | |
| Dingri County, Tibet | 87.12 | 28.66 | |
| Sar Township, Shigatse, Tibet | 87.74 | 28.19 | |
| Dingjie County, Shigatse, Tibet | 87.76 | 28.37 | |
| Dingri, Shigatse, Tibet | 88.85 | 29.00 | |
| Lazi County, Tibet | 87.64 | 29.08 | |
| Mount Qomolangma, Tibet | 86.86 | 28.13 | |
| Orinhippus trisulcus Yin, 1984 | Lazi County, Tibet | 87.64 | 29.08 |
| Yamdrok Lake, Tibet | 90.71 | 28.92 | |
| Dongga Township, Shigatse, Tibet | 88.83 | 29.36 | |
| Pacris xizangensis Zhang, Zhang&Yin, 2012 | Damxung County, Tibet | 91.09 | 30.47 |
| Foveolatacris qinghaiensis (Yin, 1984) | Tongde, Qinghai | 100.58 | 35.25 |
| Mongolotettix japonicus (Bolívar, 1898) | Qingzhenzhan Mountain, Gade County, Qinghai | 100.12 | 34.17 |
| Huzhu Tuzu Zizhixian, Qinghai | 101.96 | 36.84 | |
| Mongolotettix anomopterus (Caudell, 1921) | Diebu County, Gansu | 103.22 | 34.06 |
| Zhouqu, Gansu | 104.37 | 33.79 | |
| Dianacris choui Yin, 1983 | Jade Dragon Snow Mountain, Lijiang, Yunnan | 100.17 | 27.10 |
| Pseudoeoscyllina golmudensis Zheng&Chen, 2012 | Geermu, Qinghai | 94.90 | 36.40 |
| Dulan County, Qinghai | 98.10 | 36.30 | |
| Carinulaenotus motuoensis Yin, 1982 | Motuo, Tibet | 95.33 | 29.33 |
| Phlaeoba sikkimensis Ramme, 1941 | Motuo, Tibet | 95.33 | 29.33 |
| Xiachayu, Chayu County, Tibet | 97.02 | 28.50 | |
| Chayu, Tibet | 97.47 | 28.66 | |
| Phlaeoba medogensis Liu, 1981 | Motuo, Tibet | 95.33 | 29.33 |
| Phlaeoba tenebrosa (Walker, 1871) | Motuo, Tibet | 95.33 | 29.33 |
| Chayu, Tibet | 97.40 | 28.83 | |
| Phlaeoba albonema Zheng, 1981 | Motuo, Tibet | 95.33 | 29.33 |
| Phlaeoba infumata Brunner von Wattenwyl, 1893 | Weixi County, Yunnan | 99.29 | 27.18 |
| Chlorophlaeoba nigripennis Zheng, Lin, Xu&Ma, 2013 | Motuo, Tibet | 95.33 | 29.33 |
| Sikkimiana darjeelingensis (Bolívar, 1914) | Xiachaiyu, Tibet | 97.02 | 28.50 |
| Motuo, Tibet | 95.33 | 29.33 | |
| Jilong, Tibet | 85.30 | 28.86 | |
| Acrida kozlovi Mishchenko, 1951 | Jianzha, Qinghai | 102.03 | 35.94 |
| Xunhua County, Qinghai | 102.49 | 35.85 | |
| Guide County, Qinghai | 101.43 | 36.04 | |
| Hualong Hui Autonomous County, Qinghai | 102.26 | 36.09 | |
| Acrida curticnema Huang, 1981 | Motuo, Tibet | 95.33 | 29.33 |
| Xiachayu, Chayu County, Tibet | 97.02 | 28.50 | |
| Chiba Gully, Chayu, Tibet | 97.07 | 28.58 | |
| Acrida cinerea (Thunberg, 1815) | Zhouqu, Gansu | 104.37 | 33.79 |
| Xiachayu, Chayu County, Tibet | 97.02 | 28.50 | |
| Acrida oxycephala (Pallas, 1771) | Ledu District, Qinghai | 102.40 | 36.48 |
| Acrida exaltata (Walker, 1859) | Chayu, Tibet | 97.47 | 28.66 |
| Acrida willemsei Dirsh, 1954 | Lhasa, Tibet | 91.12 | 29.65 |
| Gonista chayuensis Yin, 1984 | Chayu, Tibet | 97.47 | 28.66 |
| Xiachayu, Chayu County, Tibet | 97.02 | 28.50 | |
| Gonista bicolor (De Hann, 1842) | Motuo, Tibet | 95.33 | 29.33 |
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Figure 1.
The Qinghai–Tibet Plateau and its external boundaries. This figure illustrates the geographic extent of the Qinghai–Tibet Plateau, marked in red, showcasing its position within the regional topography. The delineated area represents the plateau, and the boundaries distinguish it from adjacent landscapes. The map provides a visual overview of the plateau’s location within its broader geographical context, highlighting its significance as a distinct geographical feature.
Figure 1.
The Qinghai–Tibet Plateau and its external boundaries. This figure illustrates the geographic extent of the Qinghai–Tibet Plateau, marked in red, showcasing its position within the regional topography. The delineated area represents the plateau, and the boundaries distinguish it from adjacent landscapes. The map provides a visual overview of the plateau’s location within its broader geographical context, highlighting its significance as a distinct geographical feature.
Figure 2.
Sparse curves of Acridoidea in different habitats of the Qinghai–Tibet Plateau. This figure presents the species diversity indices of Acridoidea individuals randomly sampled from various habitats within the plateau using different algorithms in R studio. The y-axis represents the number of species, while the x-axis denotes the number of specimens. The point at which the dilution curves cease to show a significant upward trend indicates adequate sampling. Except for the evergreen broad-leaved forest in the warm temperate zone, which shows insufficient sampling, all other habitats demonstrate adequate sampling. Legend: SEBF—Subtropical Evergreen Broad-leaved Forest; TG—Temperate Grassland; TD—Temperate Desert; ACV—Alpine Cold Vegetation of Qinghai–Tibet; WTEBF—Warm Temperate Evergreen Broad-leaved Forest; TMR—Temperate Monsoon Rainforest.
Figure 2.
Sparse curves of Acridoidea in different habitats of the Qinghai–Tibet Plateau. This figure presents the species diversity indices of Acridoidea individuals randomly sampled from various habitats within the plateau using different algorithms in R studio. The y-axis represents the number of species, while the x-axis denotes the number of specimens. The point at which the dilution curves cease to show a significant upward trend indicates adequate sampling. Except for the evergreen broad-leaved forest in the warm temperate zone, which shows insufficient sampling, all other habitats demonstrate adequate sampling. Legend: SEBF—Subtropical Evergreen Broad-leaved Forest; TG—Temperate Grassland; TD—Temperate Desert; ACV—Alpine Cold Vegetation of Qinghai–Tibet; WTEBF—Warm Temperate Evergreen Broad-leaved Forest; TMR—Temperate Monsoon Rainforest.
Figure 3.
Alpha diversity index of Acridoidea in Qinghai–Tibet Plateau under different habitat types. Different habitat types; (a) number of species; (b) number of individuals; (c) Margalef Index (R); (d) Simpson Index (D); (e) Shannon–Wiener Index (H); (f) Pielou Index (J).
Figure 3.
Alpha diversity index of Acridoidea in Qinghai–Tibet Plateau under different habitat types. Different habitat types; (a) number of species; (b) number of individuals; (c) Margalef Index (R); (d) Simpson Index (D); (e) Shannon–Wiener Index (H); (f) Pielou Index (J).
Figure 4.
Species richness pattern of Acridoidea in the Qinghai–Tibet Plateau. This figure illustrates the distribution of species richness for Acridoidea across the Qinghai–Tibet Plateau, analyzed using ArcMap 10.8 software at a 0.5° grid scale. The species richness is categorized into four classes using the natural breaks (Jenks) classification method, with darker shades indicating a higher number of species within each grid. The data show a concentration of Acridoidea species in the eastern and southern regions of the plateau (between 94–104° E longitude and 26–34° N latitude), suggesting a pattern of higher species richness in these areas.
Figure 4.
Species richness pattern of Acridoidea in the Qinghai–Tibet Plateau. This figure illustrates the distribution of species richness for Acridoidea across the Qinghai–Tibet Plateau, analyzed using ArcMap 10.8 software at a 0.5° grid scale. The species richness is categorized into four classes using the natural breaks (Jenks) classification method, with darker shades indicating a higher number of species within each grid. The data show a concentration of Acridoidea species in the eastern and southern regions of the plateau (between 94–104° E longitude and 26–34° N latitude), suggesting a pattern of higher species richness in these areas.
Figure 5.
Species richness pattern of endemic species in the Qinghai–Tibet Plateau. This figure illustrates the distribution of endemic species richness across the Qinghai–Tibet Plateau, utilizing the same analytical methods as Figure 4.
Figure 5.
Species richness pattern of endemic species in the Qinghai–Tibet Plateau. This figure illustrates the distribution of endemic species richness across the Qinghai–Tibet Plateau, utilizing the same analytical methods as Figure 4.
Figure 6.
Specific distribution areas identified by two methods on a 1° grid scale. The non-weighted average group method reveals five endemic areas: Himalayas (F8, F9), Bayan Har Mountains (D12, D13), Jinsha River (E14, F14), Western Sichuan (E15, E16), and Hengduan Mountains (G14, G15). The endemicity parsimony analysis (PAE) identifies three areas: Jinsha River (E14, F14), Hengduan Mountains (G14, G15), and Western Sichuan Hengduan region (G14, G15). Both methods highlight the Jinsha River and Hengduan Mountains as key endemic regions, with the non-weighted method also noting the Himalayas and Bayan Har, and PAE marking Western Sichuan.
Figure 6.
Specific distribution areas identified by two methods on a 1° grid scale. The non-weighted average group method reveals five endemic areas: Himalayas (F8, F9), Bayan Har Mountains (D12, D13), Jinsha River (E14, F14), Western Sichuan (E15, E16), and Hengduan Mountains (G14, G15). The endemicity parsimony analysis (PAE) identifies three areas: Jinsha River (E14, F14), Hengduan Mountains (G14, G15), and Western Sichuan Hengduan region (G14, G15). Both methods highlight the Jinsha River and Hengduan Mountains as key endemic regions, with the non-weighted method also noting the Himalayas and Bayan Har, and PAE marking Western Sichuan.
Figure 7.
MaxEnt model predictions of suitable habitat areas for four genera of Acridoidea on the Qinghai–Tibet Plateau under current conditions (a), by 2050 (b), and by 2070 (c). The genera assessed are (A) Locusta, (B) Leuconemacris, (C) Kingdonella, and (D) Chorthippus. The model projections indicate shifts and changes in the distribution of suitable habitats for these genera over time, reflecting potential impacts of climate change on their biogeography.
Figure 7.
MaxEnt model predictions of suitable habitat areas for four genera of Acridoidea on the Qinghai–Tibet Plateau under current conditions (a), by 2050 (b), and by 2070 (c). The genera assessed are (A) Locusta, (B) Leuconemacris, (C) Kingdonella, and (D) Chorthippus. The model projections indicate shifts and changes in the distribution of suitable habitats for these genera over time, reflecting potential impacts of climate change on their biogeography.
Table 1.
Distribution data of Acridoidea species in the Qinghai–Tibetan Plateau, China.
| Family | Number of Distribution Records | Percentage of Total Family Distribution Records |
|---|---|---|
| Pamphagidae | 7 | 0.45% |
| Pyrgomorphidae | 32 | 2.08% |
| Dericorythidae | 25 | 1.62% |
| Acrididae | 1476 | 95.84% |
| Total | 1540 | 100% |
Table 2.
Species composition of Acridoidea in the Qinghai–Tibetan Plateau.
| Family | Number of Species | Percentage of Total Family | Number of Endemic Species |
|---|---|---|---|
| Pamphagidae | 7 | 1.79% | 1 |
| Pyrgomorphidae | 16 | 4.10% | 10 |
| Dericorythidae | 10 | 2.56% | 5 |
| Acrididae | 357 | 91.54% | 205 |
| Total | 390 | 100% | 227 |
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MDPI and ACS Style
Bao, B.; Wang, X.; Peng, Z.; Zhu, Q.; Li, X.; Zhang, D. Pattern of Diversity and Prediction of Suitable Areas of Grasshoppers from the Qinghai–Tibet Plateau in China (Orthoptera: Acridoidea). Insects 2025, 16, 191. https://doi.org/10.3390/insects16020191
AMA Style
Bao B, Wang X, Peng Z, Zhu Q, Li X, Zhang D. Pattern of Diversity and Prediction of Suitable Areas of Grasshoppers from the Qinghai–Tibet Plateau in China (Orthoptera: Acridoidea). Insects. 2025; 16(2):191. https://doi.org/10.3390/insects16020191
Chicago/Turabian StyleBao, Bowen, Xicheng Wang, Zhenrui Peng, Qingyao Zhu, Xinjiang Li, and Daochuan Zhang. 2025. "Pattern of Diversity and Prediction of Suitable Areas of Grasshoppers from the Qinghai–Tibet Plateau in China (Orthoptera: Acridoidea)" Insects 16, no. 2: 191. https://doi.org/10.3390/insects16020191
APA StyleBao, B., Wang, X., Peng, Z., Zhu, Q., Li, X., & Zhang, D. (2025). Pattern of Diversity and Prediction of Suitable Areas of Grasshoppers from the Qinghai–Tibet Plateau in China (Orthoptera: Acridoidea). Insects, 16(2), 191. https://doi.org/10.3390/insects16020191
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