**Author Contributions**

Conceived and designed the experiments: HDW, DWK, YSH, BMC, JHP, JWK, JHY, HWC, JHL, MJK, YMK, JHS, JK. Contributed to the acquisition of data: BMC, JHP, JWK, JHY, HWC, JHL, MJK, YMK, JHS. Analyzed the data: HDW, DWK, JK. Wrote the paper: HDW, JK.
