Landscape Approach towards Integrated Conservation and Use of Primeval Forests: The Transboundary Kovda River Catchment in Russia and Finland

Regional clear-felling of naturally dynamic boreal forests has left remote forest landscapes in northern Europe with challenges regarding rural development based on wood mining. However, biodiversity conservation with higher levels of ambition than what is possible in regions with a long forest history, and cultural heritage, offer opportunities for developing new value chains that support rural development. We explored the opportunities for pro-active integrated spatial planning based on: (i) landscapes’ natural and cultural heritage values in the transboundary Kovda River catchment in Russia and Finland; (ii) forest canopy loss as a threat; and (iii) private, public and civil sector stakeholders’ views on the use and non-use values at local to international levels. After a 50-year history of wood mining in Russia, the remaining primeval forest and cultural heritage remnants are located along the pre-1940 Finnish-Russian border. Forest canopy loss was higher in Finland (0.42%/year) than in Russia (0.09%/year), and decreased from the south to the north in both countries. The spatial scales of stakeholders’ use of forest landscapes ranged from stand-scale to the entire catchment of Kovda River in Russia and Finland (~2,600,000 ha). We stress the need to develop an integrated landscape approach that includes: (i) forest landscape goods; (ii) other ecosystem services and values found in intact forest landscapes; and (iii) adaptive local and regional forest landscape governance. Transboundary collaboration offers opportunities for effective knowledge production and learning.


Introduction
The circumboreal forest biome provides natural resources in terms of wood, minerals, oil, gas and hydroelectric energy that support human welfare and quality of life. Northern natural boreal forest ecosystems dominated by primeval and old forests, bogs and fens form an enormous biomass and carbon sink [1,2], which is sensitive to climate change. Parts of these regions in Canada and Russia

Study Area
In the European boreal forest biome, the Finnish-Russian border is a particularly interesting region to explore the opportunities for conservation of primeval forest, i.e., ecological sustainability with a high level of ambition, and to satisfy the economic, social and cultural dimensions of sustainable landscape management and governance. Berglund [51] and Lehtinen [52,53] have shown how the Finnish-Russian border area changed from a political periphery into a focus for international ecopolitics. The reason is that landscapes on either side have been treated under different geopolitically informed regimes of government. On the Finnish side, landscapes have been transformed into forests industrially managed for maximum sustained yield forestry [54], whereas in Russia, the border zone forests have been harvested and then left largely unmanaged, or remained more or less intact, due to their remoteness [53,55]. Historical social links across the border were re-activated after the end of the Cold War, and young Finnish and Russian forest conservation activists in particular have created social links between East and West [16,56]. Subsequently, a hybrid set of actors including environmental nongovernmental organisations, publishing companies, certification agencies, market researchers and citizens have brought conservation forward [57]. Such activities have challenged the accepted ideas of sovereign territory [51].
As a case study, we chose the cross-border Kovda River catchment (Koutajoki in Finnish) ( Figure 1). The catchment covers 26,100 km 2 , and extends from easternmost Finland to the northern part of the Republic of Karelia and the southern part of the Murmansk region in Russia. The catchment belongs to  [50] and Hansen et al. [60] methodology) within the Kovda River catchment. Note that the protected area borders are generalised.  [50] and Hansen et al. [60] methodology) within the Kovda River catchment. Note that the protected area borders are generalised.

Overview of Landscape History
To provide an overview the landscape history of the Kovda River catchment, and how forest landscape goods, services and values have been used over time, we reviewed the English, Finnish and Russian literature. Inspired by Cioc's [61] approach to the economic history of the Rhine, we traced the life story, or biography, of the Kovda River catchment, starting from its estuary in the White Sea. We also used information related to the landscape history from the interviews taken with the stakeholders.

Forest Canopy Loss
We mapped the combined forest canopy loss within the Kovda River catchment using remote sensing data from Hansen et al. [60] as a visualisation display ( Figure 2). Mean annual forest canopy loss is defined as "a stand-replacement disturbance, or a change from a (high) forest to non-forest state" [60]. Second, we calculated the mean annual forest canopy loss from 2001-2018 [60] by dividing the total forest canopy lost for the period 2001-2018 by the total forest area and the number of years. This was made for two scales: (a) the Kovda River catchment, using a 25 km 2 grid (5 × 5 km); and (b) the regional Russian-Finnish transboundary region, using a 625 km 2 grid (25 × 25 km). The mean annual forest canopy loss was calculated using GIS (ArcMap 10.7). Assuming a harvest rotation of 100 years, average forest losses of >1% would indicate an unsustainable loss of forest canopy. Losses of forest due to fire or wind in the selected study areas were negligible during the study period [60].

Stakeholders' Use of Forest Landscape
To map stakeholders' use of forest goods, services and landscape values, we used multiple methods (see [62]). Based on snowballing to secure stakeholder representativeness, we made 36 open-ended face-to-face interviews in the study area. The interviews lasted 60-180 minutes, were recorded digitally, and transcribed. The interviews focused on the landscape use profiles from within civil, private and public sectors, forest management and governance trends, and socio-economic developments in the lower part of Kovda River catchment. The interviews were supplemented with analyses of socio-economic statistical data and archives of local and regional administrations [63].

Stakeholders' Use of Forest Landscape
To map stakeholders' use of forest goods, services and landscape values, we used multiple methods (see [62]). Based on snowballing to secure stakeholder representativeness, we made 36 open-ended face-to-face interviews in the study area. The interviews lasted 60-180 minutes, were recorded digitally, and transcribed. The interviews focused on the landscape use profiles from within civil, private and public sectors, forest management and governance trends, and socio-economic developments in the lower part of Kovda River catchment. The interviews were supplemented with analyses of socio-economic statistical data and archives of local and regional administrations [63]. The bottom or street-level in the stakeholder interview study was the lower and most densely populated part of the Kovda River catchment, which is located in Murmansk oblast in north-west Russia [55] (Figure 1). There the main management units are eight settlements, the Kovdozersky forest management unit (4006 km 2 ), and the Kovda hydropower station. In 2006, the population in this forest management unit was about 15,000, and the population density was 3.7 people per km 2 [63,64]. Since then there has been a >15% population decline. A majority of the population live in the town Zelenoborskiy, and following the period of Soviet-period wood harvesting there are many abandoned logging villages [55].
All stakeholders were divided according to three variables. First, we defined three groups according to their rights to use natural resources. These were: (a) leasers of state land (e.g., the The bottom or street-level in the stakeholder interview study was the lower and most densely populated part of the Kovda River catchment, which is located in Murmansk oblast in north-west Russia [55] (Figure 1). There the main management units are eight settlements, the Kovdozersky forest management unit (4006 km 2 ), and the Kovda hydropower station. In 2006, the population in this forest management unit was about 15,000, and the population density was 3.7 people per km 2 [63,64]. Since then there has been a >15% population decline. A majority of the population live in the town Zelenoborskiy, and following the period of Soviet-period wood harvesting there are many abandoned logging villages [55].
All stakeholders were divided according to three variables. First, we defined three groups according to their rights to use natural resources. These were: (a) leasers of state land (e.g., the Kovdozersky state forest management unit or the Kovda hydropower station); (b) forest contractors leasing forest areas from the Kovdozersky state forest management unit for a period up to 49 years; and (c) forest users who bought rights to harvest wood at auctions. Second, drawing upon methods used by Mingione [65], we defined three groups of stakeholders according to the societal sector which they represent, i.e., (i) the civil sector (i.e., a broad range of organizations outside of government, including civil associations, non-profit organizations, churches and neighbourhood clubs that contribute to the public good [66]); (ii) the private sector (e.g., businesses controlled or owned by private individuals, Land 2020, 9, 144 7 of 27 directly or through stock ownership); and (iii) the public sector (e.g., stakeholders representing public interests through governmental agencies and local government units). Third, stakeholder activities were divided into four groups according to the spatial extents of their management or business activities. These included those who conduct their activity on: (1) local (e.g., the local garden society); (2) regional (e.g., logging forest companies); (3) national (e.g., the hydropower station, the state forest management unit); and (4) international levels (tourism enterprises).
To survey the products derived from different kinds of natural resources, we divided them into use and non-use values [33]. Direct use values include consumptive (e.g., wood and non-wood goods), as well as non-consumptive direct use values (e.g., landscape quality for recreation and tourism). Indirect use values include regulating ecosystem services (e.g., watershed protection, water purification and carbon sequestration). Non-use values are closely linked to conservation interests of the landscape. Two examples are bequest values arising from placing a value on the conservation of natural or cultural elements of the landscape for future generations, and existence values derived from the knowledge of conserved ecosystems, habitats or species. To capture the diversity of landscape components in the study area, it was stratified into five parts: (a) the airmass, (b) the open land, (c) the forested land, (d) the water (rivers and lakes), (e) the ground (quarternary deposits, bedrock and groundwater).
Discarding the early colonisation of humans, which took place several thousand years ago, the Kovda River catchment began developing economically in a modern sense during the 15th century when the northern White Sea coast was colonised by Russians. Fishing, fish pickling, reindeer farming and fish barrel production were key activities [55,69]. The Russian timber merchant Rusanov built the first sawmill in 1890 at the Kovda River mouth, which made harvesting and export of high-quality lumber possible. Additional saw-mills were built a decade later by the Swede G. Oslund and the Brit K. Stuart from (see [70]). Wood harvesting increased when another sawmill was built in 1915 at the Kovda Lake. This supplied wood for building the "October" railway between Murmansk and Petrozavodsk in 1915-17 [69]. After the Russian Revolution in 1917, the new railway allowed intensive wood harvesting. In the 1920s, several new mills were built to produce sawn wood for export. The construction of the Kovdozersky hydroelectric power station in the 1950s facilitated industrial development of the area [71,72]. Subsequently, three large water reservoirs (Knyazhegubskoye, Iovskoye and Kumskoye) were built to provide electric energy to the metallurgic industry, such as the aluminium smelter in Kandalaksha. This caused a major transformation of the aquatic system, to the extent that the Kovda River catchment became one of the most regulated large river catchments in Europe. While the chemical status of the Kovda River is good, ecological problems arise from shore erosion, organic matter decay in the submersed forests and winter low-water conditions for fish that spawn under ice in the littoral zone [73]. Further planned water regulation upstream involving Kitkajärvi and Paanajärvi was stopped in 1987.
The Kovdozersky state forest management unit in the Kovda River catchment was created in 1955. Thinning, assigning felling areas, harvesting control and safeguarding regeneration were the key activities. Logging in the lower Kovda River catchment was made by the state timber industry enterprises, which were created in the 1950s [55,69]. The forest industry was the main consumer of forest Land 2020, 9, 144 8 of 27 landscape resources during the Soviet period, especially from the 1950s to the 1970s. The harvesting of the naturally dynamic boreal forest was intense, and the annual allowable cutting quotas were exceeded. This resulted from competition among the state timber enterprises fulfilling their five-year plans of the socialistic planned economy at the time.
During the end of the 1980s, a forest industry decline began, because of the lack of accessible wood resources [55,69]. The final shut-down started in the early 1990s, due to the transition from a socialistic planned toward a market economy, as well as the remote location and previous overharvesting. During the past decades, ideas about use and restoration of wood resources have been discussed.
The decline in employment once created by hydro-electric development, mining and forestry has led to depopulation [74]. For instance, the rate of population decline (−36.9%) in the Loukhsky district was the highest in the Republic of Karelia in 2008-2019 [75,76]. During times of economic transition villages have confronted privatization, economic problems and unemployment, especially in the Russian part of the Kovda River catchment [77]. Self-sufficiency based on local livelihoods has become the most important means of survival for villagers who depend on many wood and non-wood resources (e.g., firewood and wood for house-building, berries, mushrooms and wild food) [55,78]. This calls for developing new value chains. Tourism can be presented as promising for regional development, which can mitigate social challenges, and have an important role for the conservation and reproduction of the natural, historical and cultural potential of the territory [79,80].
Moving upstream, the human influence in the Paanajärvi area was studied by Bondestam [81]. Karelians were the first to use cultivated grasses and a slash-and-burn method, but abandoned the area after a cultivation period of about one hundred years. After this, the area was only occasionally cultivated for about another hundred years. Then Finns came in with permanent field cultivation. Their period lasted for about 100 years until end of the 1930s.
With increasing altitude and remoteness from the White Sea coast and the railway along it, the area of primeval old-growth forests increases [3,82]. The Kutsa-Paanajärvi wilderness along the pre-1940 Finnish-Russian border in the west-central part of the Kovda River catchment represents a shift in the view of forestry in the region [83]. During the late 1980s, Russian scientists were involved with the establishment of Paanajärvi National Park. Finnish NGOs also reported a low human impact in areas located north of the Paanajärvi National Park [16], reaching the previous Finnish, and now Russian, Kutsa nature reserve (http://www.biodiversity.ru/kola/html/kutsa/kutsa.html).
When the Paanajärvi National Park was created in 1992 a significant part of the Pyaozersky lespromkhoz's in the Republic of Karelia, a base for logging primeval forest, was excluded from harvest. Employing more than 900 people in a community with about 4000 inhabitants, the company was vividly against further forest protection. The Pyazoresky forest industry unit made attempts to carry out massive logging in the area, while NGOs insisted on conservation of the forest area as an example of unique undisturbed nature. However, after a long conflict, in late 2004, the head of the forestry unit Pyaozersky lespromkhoz, and representatives of International Socio-ecological Union, Biodiversity Conservation Center and Greenpeace-Russia signed an agreement about a 25-year long moratorium for an old-growth territory of 109,200 ha in the area leased by the Pyaozersky forest industry unit.
As a consequence of this concentration of primeval forests, many species found or declining in managed terrestrial landscapes now remain [84]. Similarly, Huusko et al. [85], who studied the structure of fish pelagic fish populations in Lake Paanajärvi, located above the large water reservoirs, found that the size distribution of this resembled un-fished natural lakes. This stresses the importance of nature conservation as a means of maintaining the attraction of large fish for sport fishing. This overview of the biography of the Kovda River catchment clearly indicates that the use of forest landscape products has thus varied considerably during the past 500 years ( Table 2).

Forest Canopy Loss and Intact Forest Landscapes
The 2000-2018 forest canopy loss analysis (25 km 2 grid cells) for the Kovda River catchment showed that the Finnish part had the highest mean annual forest canopy loss at 0.42%/year. In contrast, the Russian part of the Kovda River catchment only recorded a mean forest canopy loss of 0.09%/year. The mean annual forest canopy loss of the local Kovda River catchment ranged from 0.00 to 2.87% for the Russian part and 0.01 to 1.16% for the Finish part ( Figure 2). Expanding the analysis to the regional Russian-Finnish transboundary scale (625 km 2 grid cells) showed a two-way division ( Figure 3). First, there was a west-east border division with the mean forest canopy loss of 0.72% for Finland and 0.18% for Russia. Second, in each country there was a south-north decreasing gradient in forest canopy loss. For Finland, the mean forest canopy loss ranged from 1.60% loss/year in the south to 0.01% in the north. In comparison, the mean forest canopy loss for Russia ranged from 1.03 % loss/year south to 0.0 in the north of the Kovda River catchment.
Land 2020, 9, x FOR PEER REVIEW 9 of 28 importance of nature conservation as a means of maintaining the attraction of large fish for sport fishing. This overview of the biography of the Kovda River catchment clearly indicates that the use of forest landscape products has thus varied considerably during the past 500 years ( Table 2).

Forest Canopy Loss and Intact Forest Landscapes
The 2000-2018 forest canopy loss analysis (25 km 2 grid cells) for the Kovda River catchment showed that the Finnish part had the highest mean annual forest canopy loss at 0.42%/year. In contrast, the Russian part of the Kovda River catchment only recorded a mean forest canopy loss of 0.09%/year. The mean annual forest canopy loss of the local Kovda River catchment ranged from 0.00 to 2.87% for the Russian part and 0.01 to 1.16% for the Finish part ( Figure 2). Expanding the analysis to the regional Russian-Finnish transboundary scale (625 km 2 grid cells) showed a two-way division ( Figure 3). First, there was a west-east border division with the mean forest canopy loss of 0.72% for Finland and 0.18% for Russia. Second, in each country there was a south-north decreasing gradient in forest canopy loss. For Finland, the mean forest canopy loss ranged from 1.60% loss/year in the south to 0.01% in the north. In comparison, the mean forest canopy loss for Russia ranged from 1.03 % loss/year south to 0.0 in the north of the Kovda River catchment.  [60] using a 25 × 25 km grid, with protected areas larger than 10 km 2 (data from the website protectedplanet.net) and remaining intact forest [50]. It should be noted that for the Russian area there were only two protected areas <10 km 2 , both located North of the Kovda River catchment. Note that the protected area borders are generalised.

Landscape Stakeholders
A total of 31 stakeholders were identified in the lower part of the Kovda River catchment. These were land leasers (13), forest contractors (9), forest users (6) and potential forest contractors (3). Private sector stakeholders made up 55% (Figure 4). They represented enterprises working with small-scale logging, tourism and agriculture. The civil sector was represented only by a local gardening society. Public sector stakeholders represented managed forests (Kovdozersky state forest management unit), water (the Kovda hydropower station) and protected areas. Private sector stakeholders stressed the need to develop new business activities. These were very diverse and ranged from small-scale based on local wood resources, farming and fisheries, tourism (e.g., nature-based, sport, fishing including maintenance and restoration of fish population and hunting) and to the construction of bio-fuel stations for central heating in towns and villages.

Landscape Stakeholders
A total of 31 stakeholders were identified in the lower part of the Kovda River catchment. These were land leasers (13), forest contractors (9), forest users (6) and potential forest contractors (3). Private sector stakeholders made up 55% (Figure 4). They represented enterprises working with small-scale logging, tourism and agriculture. The civil sector was represented only by a local gardening society. Public sector stakeholders represented managed forests (Kovdozersky state forest management unit), water (the Kovda hydropower station) and protected areas. Private sector stakeholders stressed the need to develop new business activities. These were very diverse and ranged from small-scale based on local wood resources, farming and fisheries, tourism (e.g., nature-based, sport, fishing including maintenance and restoration of fish population and hunting) and to the construction of bio-fuel stations for central heating in towns and villages. The interviews also showed that the stakeholders in the area used landscape goods, services and values in different ways ( Table 3). The stakeholders with direct use of natural resources were from public and private sectors, which were concerned with the extraction of natural resources including wood, fish and game, as well as the use of kinetic energy of water to produce electricity by the establishment of several large water reservoirs and dams. These stakeholders had the most substantial effect on the physical landscape. Additionally, sport hunting for grouse, ducks and moose was the aim for several landscape stakeholders. Over-intensive hunting could affect the population size and productivity of prey species, and thus create customer bad-will. It was therefore felt necessary to assure that hunting pressure can be maintained, as well as to identify areas with suitable habitat for different species.  The interviews also showed that the stakeholders in the area used landscape goods, services and values in different ways ( Table 3). The stakeholders with direct use of natural resources were from public and private sectors, which were concerned with the extraction of natural resources including wood, fish and game, as well as the use of kinetic energy of water to produce electricity by the establishment of several large water reservoirs and dams. These stakeholders had the most substantial effect on the physical landscape. Additionally, sport hunting for grouse, ducks and moose was the aim for several landscape stakeholders. Over-intensive hunting could affect the population size and productivity of prey species, and thus create customer bad-will. It was therefore felt necessary to assure that hunting pressure can be maintained, as well as to identify areas with suitable habitat for different species.
There were many challenges for stakeholders representing the forest industry, based on local forest resources. After a 50-year history of intensive wood harvesting amounting to 700,000 m 3 /year in the 1980s, currently, only about 10,000 m 3 is harvested annually [55]. According to the current state forest management plan, about 100,000 m 3 can be harvested annually. However, the possible final felling stands are located far away from today's transport infrastructure. The reason is that logging in the area was made as a frontier into un-harvested old-growth forest from each of the different logging villages. This is an illustration of the problem of the inaccessibility of the estimated annually allowable harvest. To combine silviculture by commercial thinning to improve stands, and to produce bio-energy assortments, there were discussions about introducing central heating based on wood chips. Another group of stakeholders delivered services in terms of recreation and tourism. This natural resource use was largely non-consumptive. Indirect use values were derived from the landscape values linked to primary forest remnants. For example, wilderness tourism has been established (e.g., https://vk.com/bazaiova). Annually >5000 people visit the Paanajärvi National Park. The area used for these tours was large and included long-distance river trips, and snowmobile safaris from Finland to the White Sea. At the same time the largest group of visitors of the Loukhsky District (65% of total inbound tourism) comes from Moscow and Saint Petersburg, and organise the visit themselves [86], a group which is difficult to reach in processes of multi-stakeholder collaboration.
The non-use values of the landscape were of importance to a fewer number of stakeholders. The recognised non-use values consisted of existence values, which were closely linked to nature conservation interests, and bequest values that, in addition, encompass cultural heritage conservation interests. Protected areas aimed at biodiversity conservation and forests with protective [87] and social functions [88] form a key Green Belt infrastructure. For example, 34% of the total forest area of the Kovdozersky forest management unit was excluded from forest exploitation for ecological reasons [55]. However, interviewees expressed the opinion that biodiversity conservation was not an urgent issue in the area. Simultaneously, stakeholders reported that culture tourism was emerging. The village of Kestenga in the Loukhsky District is an example of the rune-song villages that have preserved their ancient traditions [47]. The shores of the White Sea have a very long history of human settlement. The Pomor people culture was well represented in the Kovda village, located at the mouth of the Kovda River. For those interested in the history of forest use, another interesting village was Lesozavodskiy, which was created to produce sawn wood and to provide a port for export. The Russian-Finnish border moved considerably to the west after the Finnish winter war in the 1940s. The land lost to Russia hosts several ancient Finnish settlements, which are important for upholding cultural traditions and as tourist attractions.
The final group of stakeholders was formed by control authorities. While these stakeholders did not use the natural resources actively, they had an important law enforcement function counteracting unwanted use of natural resources. The administration of the Kovdozersky forest management unit, local administration, railroad and road authorities were some examples.
All stakeholders in the Russian part of Kovda River catchment had many opportunities for collaboration. Our study of the management interactions among landscape stakeholders ( Figure 5) indicated that: (1) there was a group of "well-integrated" stakeholders like the administration of the forest management unit and the local administrations; (2) some stakeholders had "one-way" interaction, which means that they coordinated their management activity with only one stakeholder (for example, administration of the Kutsa nature reserve and tourism enterprises); (3) "outsiders"-stakeholders, who did not integrate their activities with others. For example, the administration of hydro-power stations did not interact with landscapes' stakeholders at all. The interviews with landscapes' stakeholders allowed us to make suggestions about their role in local development of different dimensions of sustainability ( Figure 6). The majority of stakeholders contributed to the economic and social development, which means that there is a need to encourage and increase the attention to ecological and cultural issues.
forest management unit and the local administrations; (2) some stakeholders had "one-way" interaction, which means that they coordinated their management activity with only one stakeholder (for example, administration of the Kutsa nature reserve and tourism enterprises); (3) "outsiders"-stakeholders, who did not integrate their activities with others. For example, the administration of hydro-power stations did not interact with landscapes' stakeholders at all. The interviews with landscapes' stakeholders allowed us to make suggestions about their role in local development of different dimensions of sustainability ( Figure 6). The majority of stakeholders contributed to the economic and social development, which means that there is a need to encourage and increase the attention to ecological and cultural issues.

Spatial Levels of Forest Landscape Use
The spatial extents of stakeholder activities ranged from local to international. However, almost half of the stakeholders focused on the Murmansk region ( Figure 7). Thus, the goods, services and values of the landscape cover multiple spatial scales (Table 4).

Spatial Levels of Forest Landscape Use
The spatial extents of stakeholder activities ranged from local to international. However, almost half of the stakeholders focused on the Murmansk region ( Figure 7). Thus, the goods, services and values of the landscape cover multiple spatial scales (Table 4).   Table 4. Spatial scales for different types of planning identified from interviews with 31 categories of users of forest landscapes' goods, services and values in the lower part of the Kovda River catchment, as determined by the stakeholders' use profiles.

Spatial Scale Type of Planning Landscape Stakeholders
Trees in stands (~1-100 ha) Operational planning (e.g., general considerations in forest management, stream and riparian management)

Forest leasers that harvest wood
Stands in management sub-unit (=landscape, leasing area) (~2,000 to 100,000 ha) Tactical planning (e.g., forest management, landscape planning for game species) Forest management unit, nature protection units Landscapes (~500,000 ha) Strategic planning (e.g., secure use of wood and non-wood goods, energy, tourism) Forest management unit, municipality, district River catchment in the boreal forest and its administrative units (~5,000,000 ha) Regional planning for sustainable development (e.g., hydro power, tourism, conservation of "green belt" forest) Hydroelectricity production, authorities working with nature conservation 0 Social Cultural Economic Figure 7. Representation of the spatial scales of stakeholders' activities in the lower part of Kovda River catchment. The main planning units in the Russian part of the Kovda River catchment were the four forest management units Kandalakshsky, Kovdozersky in Murmansk, and Pyaozersky and Chupinsky in the Republic of Karelia. An important task for them was to enhance co-existence of forest land leasers operating at different spatial scales. For example, in the Kovdozersky forest management unit, 17 stakeholders leased different parts of this area to operate businesses based on wood harvesting, hunting and recreation. To secure sustainable use of goods, services and values, their states and spatial distributions need to be mapped. This applies to wood resources for timber and biomass for bioenergy, habitat availability for game and fish, nature and culture values as bases for tourism, opportunities for hydro-electric development and for nature conservation through protected area networks. Some users expressed a need for regional trans-boundary planning covering the entire Kovda River catchment, i.e., in Murmansk oblast, the Republic of Karelia and Finland. Three examples are hydro-electric development, tourism linked to the Finnish and Russian cultural heritage in the border zone between the two countries, and the conservation of one of Fennoscandia's intact forest landscapes located entirely within the Kovda River catchment.

A Diversity of Boreal Forest Landscape Perspectives
In the light of climate change and globalisation, boreal forest conservation has become a key issue [50,[89][90][91]. To sustain primeval forests as a core value in large intact forest landscape remnants, there is a need to reconcile the needs and desires of landscape stakeholders at the local and regional level with national and international policies related to biodiversity conservation and sustainable development [37,92,93]. Given a range of drivers, like biodiversity conservation, globalisation of economies, energy supply and climate change and Russian wood export tariffs, decision-makers at multiple levels need to learn about these issues and how to handle them, and how to cope with uncertainty and risk [94,95].
From the Kovda River catchment's perspective the economic use of once primeval forests since the ceding of the Salla area to the Soviet Union in 1940 (see Figure 1) spread from the White Sea in the east and upstream. While forest logging in the catchment began in the late 19th century along the White Sea coast [70], it only became intense in the 1950s, and at present, forestry for wood production has declined dramatically [55]. As a consequence of its regional remoteness and associated values Russian-Finnish cooperation has prospered [96,97]. Nevertheless, rural settlements are in decline, and locals rely on a wide range of wood and non-wood resources [44,52,98,99]. While hydropower production continues, planning measures are discussed to increase the reliability and safety of hydraulic structures, due to the poor quality of constructions and the limited consideration of local operating conditions [100], as well as to develop compensation measures for lost fish industry opportunities [101]. At the same time, tourism based on natural and cultural values is emerging [102].
The development since the early 1950s can thus be described as a transition from an industrial society based on hydro-electric development, mining and the clear-felling of primeval forests, which indeed gave many people jobs and created resource-dependent rural communities during a relatively short period of time, to declining local societies in Russia. In contrast, the combined use of wood and non-wood goods, services and values are emerging in Finland [103][104][105].
There are also attempts to develop cross-border tourism between Finnish Lapland and southern Kola region based on natural and cultural heritage [106]. Tourism development program work is currently also going on in the Murmansk region. This strategy concentrates mostly on development of cross-border tourism in the Kovda River catchment. The main attractions for tourism are nature protected areas with unique boreal primeval forests and wild rivers.
Our study shows that the range of values people associate with forest landscapes is wide. We conclude that: (a) the values are new and evolving, and many stakeholders are not prepared for this; (b) the demands to deliver goods, services and values from forest landscapes are steadily increasing; (c) the rate of range is accelerating, and there is a need for continued assessment of ecological, economic and socio-cultural consequences; (d) there is an urgent need for collaboration among sectors within a geographical area to avoid losing desired values.
Regarding the history and culture of forest landscapes in the border zone between Finland and Russia, Lehtinen [44] stated "The violent past of the region stands as clear proof that this multitude of can only be supported and developed by prioritizing culturally and ecologically sensitive means of cooperation across the border, instead of large-scale industrial, ecological or geopolitical projections controlled from the outside".
The interviews with stakeholders in the Kovda River catchment clearly show a much wider range of spatial scale of forest landscape use than in the past. This is consistent with the pattern of globally emerging efforts towards multi-level participation of actors and stakeholders representing private businesses, public organisations and civil society. Therefore, we conclude that a sustainable forest landscape requires integrated spatial planning at multiple spatial scales, and landscape stewardship based on multi-level governance [95]. The emergence of new and the deepening of existing cross-border links between sectors (forest, mining and tourism) is pivotal for sustainable local land-use and natural resource use policies [107].

Trees in Stands
Data about species, density, size and volume of trees, as well as site conditions form the basis for operational economic forest planning aimed at wood harvest and sustained yield wood production. In addition to site conditions affecting growth and transport infrastructure, tree data forms the base for dividing a management unit into forest stands with homogenous properties [108]. To mitigate problems caused by forestry intensification for biodiversity conservation retention, forest certification has promoted set-aside of woodland key biotopes, such as paludified forest patches, buffers around water bodies, habitats of red-listed species, as well as of dispersed retention of living trees, snags and dead wood [109,110]. However, Blumröder et al. [111] stressed that the effectiveness of stand-scale conservation instruments has to be assessed, with respect to edge effects and secondary dieback of retention trees and patches. For example, while Russia has a unique history of forest zoning separating production and environmental functions, however, the Russian Forest Code of 2006 increased the focus on wood production. In turn, this increased harvest rates in stream buffer zones [87]. Moreover, even if effective, these stand-level conservation efforts do not help to solve conservation problems related to birds, mammals and fish, all of which have landscape scale requirements.

Stands in a Management Sub-Unit
To carry out logging operations, there is a need to develop a sufficiently dense network of forest roads that can be used at least seasonally for wood mining. Given that the cost for terrain transport using forwarders is higher than road-transport hauling with trucks, there is a need for a dense road network, especially if multiple forest silvicultural treatments are planned within a management unit. The cost to transport timber with an all-terrain forwarder rapidly increases for distances over 500m [108]. Therefore, focusing on intensive forest management, the Swedish road plan 1990 aimed at leaving no point in the managed forests more than 500 m away from a forest road, allowing hauling by trucks [112]. Sundberg & Silversides [108] estimated the total density of main artery roads, secondary and feeder roads required to 15-25 m per hectare. In the Republic of Karelia and the Murmansk region, the road density is about 2 m per hectare [113].
International and national biodiversity conservation policy commits forest actors to maintain viable populations of naturally occurring species. The emergence of spatial planning of forest stands with different properties in a forest management unit in Nordic forestry is linked to the maintenance of species that cannot cope with the simplification of the forest ecosystem that sustained yield forest forestry causes [114,115]. The reduction of dead wood, old deciduous trees, old forest stands and functional habitat networks are four examples. Termed ecological landscape planning or landscape ecological planning Swedish and Finnish private and state forest companies developed approaches to support the presence of different lichen, moss, fungal and insect species, by setting aside stands with natural forest structures (e.g., woodland key habitats) [116].
However, to maintain boreal biodiversity according to policy means, at least, that a minimum amount of habitat patches with sufficient quality, size and concentration is needed for each forest ecosystem type in every ecoregion. The Swedish state forest company Sveaskog is a good example. This company has used ecological knowledge to set a target of 20% of the productive (i.e., more than one m 3 /ha and year) forest landscape to be set aside for nature conservation, and to design local Land 2020, 9,144 16 of 27 and regional networks of different forest ecosystems that represent the five main different natural ecoregions in Sweden. The target of 20% in each ecoregion is reached by leaving individual trees, tree corridors, groups and patches of trees, as well entire forest stands. About half of the ecoregional targets are made up by sub-stand level retention, and half by setting aside entire stands (S. Bleckert pers. comm.). In Russia, the national FSC standard could be viewed as an example of bringing biodiversity considerations with high level of ambitions to the forest management. The indicators in the Russian FSC standard require the maintenance of communities of naturally occurring species and to some extent ecological integrity of the representative ecosystems of an eco-region [109,117]. On this scale, the planning for social-cultural issues could be done for the maintenance of non-wood forest products, which are important for local people during the transition period [88].

Landscapes in Management Unit
As is the case of early industrial use of wood resources anywhere, Soviet wood harvesting was based on what can be called wood mining. The transition from this system, combined with a planned economy, to the current broad range of forest landscape goods, services and values, puts new demands on spatial planning across several forest owners, leasers and users. This general trend is clearly corroborated by the different forest users in the Kovda River catchment.
After a 50-year history of old-growth logging, final felling stands are located far away from today's transport infrastructure. The reason is that logging was made as a wood mining frontier from each of the different logging villages, now by and large abandoned. This is an illustration of the problem of the inaccessibility of the allowable harvest, which often in reality result in severe local overharvesting. As a consequence, thousands of hectares of forests around abandoned forest villages are young to middle-aged never pre-commercially cleaned nor thinned forest stands. To combine silviculture for improved stands and to produce bioenergy, there are discussions about introducing central heating in larger settlements based on wood chips, which are produced locally from the wood after thinning to improve stands. Currently produced wood chips are mainly sold abroad. Sport hunting for grouse, ducks and moose is the aim for several landscape stakeholders. Intensive hunting does affect the population size and productivity of prey species. It is therefore necessary to assure that hunting pressure can be maintained, as well as to identify areas with a suitable habitat for different species. Another issue is the representation of different forest types for biodiversity conservation. While the Paanajärvi and Kutsa primeval forest massifs are indeed large, most of them are located at >300 m a.s.l., which is less preferred by focal species such as the flying squirrel [118]. It is thus necessary to assess the functionality of different forest habitat networks at the scale of landscapes at the regional level [22].
The mapping of high conservation value forests (e.g., intact forest landscape with primeval forests) needs to be made at multiple scales, from that of large region, local landscapes, to the scale of trees in stands at the local level [119]. An interesting approach to integrated landscape planning within ecoregions was developed by Sveaskog Co. in Sweden. Based on the insight that habitat network functionality is alleviated by concentrating set-asides of high conservation value forest, Sveaskog Co. made analyses of all their management units to map high conservation value areas in Sweden. The management units with the highest natural values in each ecoregion are termed Ekopark [120]. Here, the local proportion of set-aside areas for biodiversity conservation is 30%-50%. Indeed, this approach has led to better habitat network functionality than other forest owner categories focusing on wood production.
At this spatial scale, planning for maintenance of cultural landscapes, which have high cultural, social and ecological values, according to international and national policies [25], is an important issue. The traditional villages in the Kovda River catchment reflect the ancient cultural and land use traditions of the Pomor people. Spatial planning for protection of cultural landscapes and traditional village systems should be a milestone in a regional program designated for keeping cultural diversity and social stability of forest landscapes. The village system intactness would be a good partial indicator of success [121]. Since the 1990s, when the benefits of tourism development became apparent, the Republic of Karelia developed, and has continuously improved, a system of strategic management. The 2007 General Layout of Tourist Sights and Tourism Infrastructure in the Republic of Karelia identified twelve tourism zones, one of which being the Pyaozersky zone, located in the Loukhsky District border districts [47,80]. At the municipal level, the significance of tourism development has been emphasised in the programme for the socio-economic development of the Louksky District. In general, the implementation of the programme for tourism development will increase the contribution of the industry to the socio-economic development of the area, enhance their investment attractiveness and create a competitive tourism product [122].

Ecoregional Level
Several landscape stakeholders used goods, services and landscape values at a spatial scale encompassing the entire Kovda River catchment from Finland to the White Sea and further. Three examples are stakeholders working with the conservation of primeval forests, cross-border tourism and hydro-electric development.
The main focus of conservation is the primeval forests along the Russian-Finish border. Although very large, the boreal forest region has been transformed from its naturally dynamic state by various anthropogenic effects. Gradually expanding frontiers of clear-felling is a major impact. Being remote from centres from economic development in Russia, and not accessible from the Finnish side, a "Green Belt" of large intact forest landscapes is located immediately east of the Finnish-Russian border [44]. Focusing on the northern boreal forest ecoregion, one of these large intact forest landscapes is formed by the Paanajärvi-Kutsa wilderness in the upper part of the Kovda River catchment. Apart from the "Green Belt" forests, other main reference areas to understand the natural range of variability are the forest massifs at the border between the Republic of Karelia and the Arkhangelsk oblast, the Dvina-Pinega watershed, and the forest complex along the western slopes of the Ural Mountains, made up by Yugud-Va National Park and the Pechora-Ilych zapovednik, and its buffer zone in the Unya River catchment in the Komi Republic [12,123]. Efimov [124] presented a masterplan for a collection of primeval forest areas in Northwest Russia to provide ecosystem corridors in both east-west and south-north directions and for the Barents Region. It is essential that assessments of the functionality of protected area networks are made [125,126]. For example, a review of protected areas in Murmansk Oblast by Borovichev et al. [127] concluded that effective environmental protection is provided only by state nature reserves and national parks, (4.2% of the Murmansk Oblast), and that "it is hardly possible to guarantee the proper level of biodiversity conservation and the stability of the ecosystem of the region".
The appearing cross-border tourism development is based mainly on ecological and cultural values of forest landscapes [5,105,128]. To reconcile the interests of forestry, nature conservation and rural development spatial planning for entire the Kovda River catchment in needed. Similarly, the production of electricity by the Kovda hydro-power stations for the customers on national and international levels require spatial planning to integrate the interests of tourism development, conservation of biodiversity, local fisheries and forestry. Table 5 outlines how planning for ecological, economic and socio-cultural dimensions of sustainable use and governance of forest landscapes takes place at multiple spatial scales. Table 5. Outline of how different planning dimensions can be framed using planning horizons (short-term to long-term) across multiple spatial scales for: (a) hierarchical forest planning of wood supply-roads-logging-silviculture; (b) securing ecological dimensions through biodiversity conservation planning, including ecosystem services; and (c) to satisfy socio-cultural dimensions, based on regional comprehensive planning.

Landscape Approach to Conservation and Use of Primeval Forests
European regions holding valuable natural and cultural heritage are often located across state borders, especially if linked to topography or other features that hampered economic development [129][130][131]. In general, East European countries have more intact biodiversity compared to Western Europe, including species, habitat networks and natural processes [132] and cultural values [121,133]. However, at present, regions located along the eastern border of EU stand at a crossroad [134]. First, this is between continued decline of biological and cultural diversity, aggravated by continued wood mining [22,87] and bio-economic policy, aimed at intensified biomass production [135]. Second, it is about developing new approaches to landscape stewardship and the management of landscapes and regions that are built not only on wood products, but also on a wide range of landscape values as a base for nature-based tourism [91,136]. For example, in Poland, conflicts between existing ecological habitat networks and improved road networks [137] and intensified forestry and tourism development [24,138,139] are currently occurring along the Polish-Belarus border. This urgent conservation challenge calls for the identification of landscape stewardship platforms for co-operation and implementation of cross-border management mechanisms at local, regional, national and transnational levels. Steering towards sustainable landscape stewardship means "a place-based, landscape-scale expression of broader ecosystem stewardship" [140] that maintains natural capital. This requires knowledge and skills to navigate the complexity of interactions within landscapes' social systems, through an inclusive societal learning process [30]. These efforts must also support communication, education and public awareness about the state and trends of all dimensions of sustainability. Working towards the sustainable development of landscapes using governance and landscape stewardship requires special emphasis on finding a common platform for the inclusion of ecological, economic, socio-cultural values, as well as approaches towards a system of good governance. Adaptation and learning requires an iterated procedure that involves multi-level co-operation among disciplines, sectors and actors [141,142]. To achieve this, a transdisciplinary approach should be applied [143]. This means that both human and natural science disciplines need to be included, as well as stakeholders using natural resources, all with the aim to balance landscape values from policy to practice, and back again [144]. Transdisciplinary research has four features that separate it from disciplinary sciences [142]: (1) it develops an evolving framework to guide problem solving efforts, which focuses on achieving holistic understanding; (2) it develops its own theoretical structure, research methods and modes of practice, based on the input from different disciplines; (3) unlike the disciplinary sciences, where results are communicated through institutions, knowledge production is communicated by those (practitioners and scientists) who have participated in the work; (4) it is about problem solving on the move. Thus, communication in ever new configurations is crucial. Landscape is an important concept within humanities and social sciences, as well as natural sciences [36,145]. The landscape concept can thus be used as an interface for improved communication between human and natural sciences, as well as between policy and practice, to increase the understanding of dependencies between social and ecological systems. Such a landscape approach is characterised by a focus on five core features: (1) a geographical area, (2) collaboration among stakeholders, (3) a commitment to sustainable development, (4) knowledge production, and (5) sharing of knowledge and experiences [146]. In the downstream part of the Kovda River catchment, the Kovdozersky Model Forest was such an initiative [55,147]. Cross-border co-operation (CBC) is another new form of governance, as an essential part of the Pan-European integration process for overcoming historic divisions, eliminating stereotypes in mutual perception, strengthening good-neighbourly relations between nations, ensuring stability, peace and socio-economic development [121,148]. The need to consider political, institutional and organisational dimensions of such initiatives, their abilities to create a landscape strategy for guiding the future development, and what they achieve on the ground in actual landscapes, is a major task for policy and decision makers and for research [62,95,149]. Financing a suite of cross border landscape strategies with the purpose of gaining experiences with such collaborative processes would be valuable. Recent reviews about the landscape approach concepts model forest [147] and Long-Term Socio-Ecological Research (LTSER) platform [150] provide systematic analyses of barriers and bridges [151].

Conclusions
This case study about the eastern external border of the European Union as a transition zone between Europe's East and West is a clear example of how different political and economic histories, as well as governance systems, have created large contrasts in social-ecological systems between adjacent regions. Following the Soviet era of wood mining, the remaining primeval forest and cultural heritage remnants are located along the Russian side of Finnish-Russian border. That the recent forest canopy loss is higher in Finland (0.42%/year) than in Russia (0.09%/year) is advantageous for primeval forest conservation in Russia. We argue that the planning of a collaborative development direction involving different landscape stakeholders, regional and municipal authorities, as well as local citizens, is crucial for future sustainable multiple landscape uses in the Kovda River catchment. Wilderness based on primeval forest conservation in large intact forest landscapes and cultural landscapes are key assets, and product development built on non-wood forest goods is emerging in new kinds of value chains. To avoid conflicts between non-use and use values, there is a need for an integrated social-ecological approach. This involves: (1) assessment and communication about the states and trends of different dimensions of sustainable forest landscapes; (2) empowering stakeholders towards collaborative learning and landscape stewardship built on multi-level governance; and (3) undertaking spatial planning at multiple spatial scales to reconcile primeval forest conservation with the needs and interests of different landscape stakeholders.