Water Conserving Irrigation Practices, Plant Growth, Seasonal Crop Coe ﬃ cients, and Nutrition of Container-Grown Woody Ornamentals

: Irrigation practices for container nursery crops often result in over-application and can lead to leaching of nutrients and reduced growth. Our objectives were to: (1) compare growth and foliar nutrient content for plants under daily water use (DWU) based irrigation treatments, (2) determine DWU of 14 woody ornamental taxa, and (3) classify taxa into irrigation functional groups based on crop coe ﬃ cients (K C ). Irrigation was applied daily to 8 taxa in 2009 and 2010 using a control of 19 mm and three irrigation treatments: (1) replacing 100% plant DWU (100DWU) each day, (2) alternating 100% DWU with 75% DWU in a 2-day cycle (100-75DWU), and (3) a 3-day cycle replacing 100% DWU the ﬁrst day and 75% DWU on the second and third days (100-75-75DWU). In 2009, seasonal average DWU ranged between 8.8 and 17.3 mm depending on taxa and treatment. Most DWU-based treatments resulted in less water applied than the control, yet plant growth was not reduced, and for one taxon ( Hydrangea paniculata ‘Limelight’) the 100DWU increased plant growth index. Lower foliar P and K concentrations were found for several taxa in control versus DWU treatments. In 2010, DWU for the season ranged between 2.1- and 22.0-mm d − 1 depending on taxa and treatment. Growth was lower only for 100-75-75DWU Hydrangea paniculata ‘Limelight’ compared to other treatments and there were no di ﬀ erences in foliar nutrient content.


Introduction
Container landscape nurseries grow a large number of plants using a correspondingly large amount of inputs per hectare, especially water, to produce an attractive product for consumers. Since container substrates are designed to drain rapidly and containers provide a relatively small reservoir, irrigation is often applied once or more each day. Overhead irrigation is the most common system for container production [1]. Only 13% to 26% of the applied overhead irrigation is retained in the container, the remainder lands between containers or is leached out [2]. Additionally, irrigators tend to over-irrigate in order to avoid the negative consequences of drought stress without considering the more subtle negative consequences of over-irrigating [3,4]. In addition to increasing water withdrawals, inefficient irrigation practices also result in greater runoff and nutrient losses through leaching, which can move offsite to contaminate adjacent water resources [5][6][7]. Moreover, increasing competition between urban and agricultural water use and greater interest in environmental fate of water necessitate improved irrigation practices be developed for agricultural production [8][9][10].
Improving irrigation management in container production often reduces the amount of water applied, decreases the volume of runoff and reduces nutrient movement in runoff [6,7,[11][12][13].
Sensor-controlled irrigation systems can reduce irrigation applications between 30% to 75%, depending on species, with little to no effect on plant growth [3,6,7,13,14]. Sensor-based irrigation can also reduce total quantities of nitrates and phosphates in effluent [6,7]. Furthermore, when used to schedule irrigation, soil moisture sensors reduce crop losses due to disease and increase profit for ornamental plant producers [4].
When using ET-based irrigation scheduling, a reference ET (ET 0 ) and the crop coefficient (K C ) must be known [15]. ET 0 is often available from local agricultural weather reports. While K C values have been determined for a wide variety of field-grown crops, K C values have been estimated for only a few dozen out of the thousands of woody ornamental taxa grown today [6,13,16,17]. The determination of K C for more taxa of container-grown is necessary for ET-based models to be useful in irrigation scheduling. To reduce the need to determine K C for every container taxa grown today, Yeager [18] suggests using indicator species to group plants having similar water needs into the same irrigation blocks. However, there is little quantitative data regarding container plant water use to allow selection of indicator plants or to group plants by water needs.
The objectives of this project were to compare the effects of three irrigation treatments based on plant daily water use (DWU) to a control daily irrigation rate of 19 mm d −1 for woody ornamental shrubs produced in containers. Plant growth index (GI) and dry weight, DWU, K C , water use efficiency (WUE), substrate leachate electrical conductivity (EC) and pH, and foliar nutrient concentrations were determined to compare the 3 DWU-based irrigation treatments with the control.

Site
The Michigan State University Horticulture Teaching and Research Center, Holt, Mich., USA is located at latitude 42.67 • N, longitude −84.48 • W, and elevation of 264 m. Plants were grown on a site representative of a typical production nursery. Containers were placed on a level surface of limestone gravel underlain at a 15 cm depth with woven polypropylene permeable landscape fabric to reduce weed emergence and subsidence of the gravel. There were 12 irrigation zones with dimensions 4.9 m N to S × 7.3 m E to W and spaced 0.3 m apart. An on-site Michigan Enviro-weather station [19] was used to monitor environmental conditions including calculation of ET 0 with the modified Penman equation [15].

Irrigation System
Irrigation was activated in each production bed by 1.9 cm diameter 24 V alternating current solenoid valves. Irrigation nozzles (Pro-Spray ® , Hunter Industries Incorporated; San Marcos, CA, USA) were mounted on 1.3 cm diameter by 0.66 m high risers. The nozzles were spaced 2.44 m apart along the perimeter of each irrigation zone with all water directed inward. Four 90 • nozzles were positioned on the corners of the irrigation zone, two 180 • nozzles were positioned between the corner nozzles on each E-W perimeter, one 180 • nozzle was placed on both the N and S edges of irrigation zone, and two 360 • nozzles were positioned along the centerline of the irrigation zone. Each nozzle had a 2.44 m radius of throw to provide 100% nozzle-to-nozzle overlap.
Irrigation system distribution uniformity (DU) and application rate in each treatment replicate were determined in 2009 and 2010 using 16 rain gauges randomly interspersed throughout the irrigation zone and allowed to collect water for 20 min as described in Dudek and Fernandez [20]. The average application rate and DU was 47.3 mm·hr −1 and 74.8% in 2009 and 46.4 mm·hr −1 and 76.8% in 2010. were obtained from a commercial nursery in 5.7 × 5.7 cm plug containers on 1 August 2008. They were planted in 10.2 L containers with an 85 pine bark: 15 peat moss (vol:vol) substrate between 2 September to 9 September 2008 for use in the 2009 study. Cuttings from the same nursery of Hydrangea arborescens L. 'Abetwo', Hydrangea paniculata 'Limelight', Rhus aromatica Aiton 'Gro-Low', Spiraea fritschiana C.K. Schneid. 'Wilma', Syringa meyeri C.K. Schneid. 'Palibin', Syringa xhyacinthiflora Rehd. 'Evangeline', Viburnum dentatum L. 'Ralph Senior', and Weigela florida 'Alexandra' were potted as described above between 15 September to 18 September 2009 for use in the 2010 study. All cultural practices in both years except irrigation were identical for all treatments. On 9 June 2009 and 22 June 2010, each container was top-dressed with 54 g of 19N-2.6P-10K controlled release fertilizer with micronutrients (5-6-month release at 26.7 • C or 21.1 • C, Polyon ® Reactive Layers Coating, Harrell's Inc., Lakeland, FL). Weeds were removed by hand pulling as necessary. Plants were overwintered each winter in a minimally-heated (−2.2 • C minimum temperature) quonset house covered with 4 mil overwintering film permitting 30% light transmission. In May of each year, the film was removed and plants were irrigated to container capacity as needed until beginning the treatments.

Experimental Design
A control and three irrigation treatments were replicated three times and assigned to the 12 irrigation zones in a completely randomized design. The control was application of 19 mm·d −1 , and the treatments were (1) irrigation applied to replace 100% daily water use (100DWU) each day, (2) applications alternating 100% DWU with 75% DWU in a 2-day cycle (100-75DWU), and (3) a 3-day application cycle replacing 100% DWU on the first day and 75% DWU on the second and third days (100-75-75DWU). Each treatment replicate contained six subreplicates of each of the eight shrub species for a total of 48 experimental plants in each treatment replicate. Experimental plants were randomized in six rows of eight at the center of the irrigation zone and inset at least 1.2 m from the edge with guard plants surrounding the perimeter to reduce edge effects. Guard plants consisted of several species having similar growth rates to the experimental plants and were arranged in the same sequence in each treatment replicate.

Daily Water Use and Irrigation Scheduling, 2009
Substrate volumetric water content (θ) was measured in 2009 for every plant using time domain reflectometry (TDR) soil moisture sensors (ThetaProbe Type mL2x, Delta-T Devices Ltd., Cambridge, UK) connected to a handheld reader (ThetaMeter Hand-Held Readout Unit Type HH1, Delta-T Devices Ltd., Cambridge, UK). For each container, the soil moisture sensor was inserted vertically into the substrate halfway between the center and the outer wall of the container to a depth of 6 cm in three locations 120 • apart. Volumetric water contents were calibrated for organic substrate using an equation developed by Warsaw et al. [13] for the same substrate. On 11 June 2009, at initiation of the study, irrigation was applied until the substrate for all plants exceeded container capacity. Gravimetric water was allowed to drain for 30 min.
Daily water use for each plant was calculated as: DWU = (θ i − θ f ) × container substrate volume where: DWU = daily water use in L θ i = initial volumetric water content (%) after gravitational water drained θ f = final volumetric water content (%) after 24 h container substrate volume = 9.7 L Control and DWU-based treatments were programmed into a time-based controller (Rain Bird ESP-12LX Plus, Rain Bird Corporation, Azusa, CA, USA) for the period until the next measurement of DWU. New DWU were obtained approximately every 21 d. Overhead irrigation was applied daily from 11 June 2009 (Day 1) to 14 October 2009 (Day 126) beginning at 0700 HR based on the DWU of the lowest water user(s). Any taxa requiring more than 50 mL (0.89 mm) over the base irrigation level received the balance by hand watering.

Daily Water Use and Irrigation Scheduling, 2010
In 2010, irrigation treatments were similar except that time capacitance soil moisture sensors (model 10HS, Decagon Devices, Inc., Pullman, WA) replaced the TDR sensor to provide continuous θ sampling and real-time irrigation control. A total of 96 sensors, one per species in each replicate, were connected in single-ended configuration to a datalogger system (AM16/32B multiplexer and CR3000 datalogger, Campbell Scientific, Inc., Logan, UT, USA). The datalogger recorded θ for each sensor at 15 min intervals from 7 June to 31 October 2010. A relay controller (SDM-CD16AC, Campbell Scientific, Inc., Logan, UT, USA) was connected to the datalogger to control the irrigation solenoid valves for each treatment replicate.
On 20 June 2010, each of the 96 sensors was individually calibrated in situ to the substrate moisture content. Irrigation was applied to bring the substrate to container capacity, and each container was weighed after drainage ceased using a PM 30 electric balance (Mettler-Toledo, Inc. Columbus, OH, USA). Five subsequent weights were taken during a 2-d dry-down period and recorded with coinciding sensor output. These data were plotted using Microsoft Excel (2007) and the trend line feature was used to obtain a single best-fit quadratic equation. Calibrations were verified using the PROC REG function in SAS (SAS Institute; Cary, NC, USA) before inclusion in the datalogger program. The calibration equations were then incorporated in the datalogger programming using Version 2.7.0.16 of CRBasic Editor (Campbell Scientific, Inc., 2006, Logan, UT, USA). Daily Water Use was calculated based on the daily change in θ and container substrate volume as in the previous year.
Irrigation run time was then calculated for each taxon in the irrigation zone by: where AR is the irrigation application rate as determined above. The program used the highest run time from the 8 plant taxa multiplied by the appropriate treatment fraction (100% or 75%, depending on treatment cycle) to apply irrigation for the proper duration.

Plant Response
Crop coefficients (K C ) were determined for each taxon receiving 100DWU (well-watered and no limitations on crop growth or evapotranspiration) using the formula K C = ET A /ET 0 [15], where ET A is actual crop evapotranspiration (measured as DWU) and ET 0 is reference evapotranspiration obtained from the on-site weather station. An adjusted K C (K Cadj ) was determined using the same equation for the plants under potentially growth and/or ET limiting conditions [15], e.g., the control and deficit irrigation treatments. Plants were classified as low (K C < 2), moderate (2 ≤ K C < 3), or high (K C ≥ 3) water users as described by Warsaw et al. [13].
Monthly growth index (GI) was calculated as the average of plant height (H) from the container rim to the highest point of the plant and plant widths along the north-south (W NS ) and the east-west axis (W EW ) axis [GI = (H + W NS + W EW )/3]. Prior to the first measurement, all taxa were pruned to a uniform size. Plant shoot dry weight for Rhus aromatica 'Gro-Low', Viburnum dentatum 'Ralph Senior', and Weigela florida 'Alexandra' was measured at the end of the experiment in 2010 using three plants from each treatment replicate. The stem was cut at the substrate surface and the entire top was bagged, oven (Grieve Corp. model SC400, Round Lake, IL, USA) dried at 80 • C until no change in dry weight was detected, and weighed.
The pour-through extraction method [21] was used to determine electrical conductivity (EC) and pH using a portable EC meter and a portable pH meter (Horiba Cardy Twin EC and Horiba Cardy Twin pH, Spectrum Technologies, Inc., Plainfield, IL, USA). Leachate was collected 30 to 60 min after irrigation for each taxon using the same plants each time 3 times throughout 2009 and 6 times throughout 2010.
Foliar samples for nutrient analysis were collected mid-season and late-season from five taxa in 2009 and six taxa in 2010. Sixty recently fully expanded leaves were collected for each treatment replicate for each taxon (approximately 10 leaves per subreplicate). Percent (dry weight) N, P, K, Ca, Mg, Na, S, and concentration of Fe, Zn, Mn, Cu, B, and Al were determined. The Dumas combustion procedure (AOAC 968.06) [22] was used to determine foliar N. Other minerals were dissolved from organic material using open vessel microwave digestion (SW846-3050B) with mineral analysis determined using inductively coupled argon plasma (ICAP analysis (AOAC 985.01) [22].

Statistical Analysis
Analysis of variance was conducted for irrigation volume, DWU, K C , GI, dry weight, substrate EC and pH, and foliar nutrient content using the PROC GLM procedure of SAS (SAS Version 9.1; SAS Institute, Cary, NC) and data normality were checked using the PROC UNIVARIATE procedure. DWU data showed several outliers due to a variety of factors ranging from sensors becoming dislodged in the substrate to rodent damage of sensor cables. Consequently, any DWU values in excess of 3000 mL were excluded from the analyses for DWU, K C , and WUE. When significant (α ≤ 0.05) differences among treatments or taxa were indicated, Tukey's Honestly Significant test was used to separate means. On 31 August 2010 (day 70) in the last replicate to run each day (a 100-75-75DWU replicate), a fault caused the farm irrigation pump to shut off prematurely for 4 days resulting in no irrigation for the replicate. Therefore, data from that replicate was excluded from analysis of DWU and K C throughout the entire season and GI, EC, pH, and foliar nutrient concentration from day 70 onward. H. paniculata 'Limelight' required the most irrigation and I. virginica 'Morton' the least in 2009. Compared to their respective controls, irrigation was reduced for H. paniculata 'Limelight' by 7%, 19%, and 23% for 100DWU, 100-75DWU, and 100-75-75DWU, respectively, and by 48%, 55%, and 57% for I. virginica 'Morton' for 100DWU, 100-75DWU, and 100-75-75DWU, respectively, with reductions for other taxa in between (Table 1). In 2010, irrigation was based on the highest DWU among taxa within each treatment, therefore, there are no differences in irrigation due to taxa, only treatment (Table 1). Compared to the control, total irrigation applications in 2010 were 19% and 50% greater for 100DWU and 100-75DWU while 100-75-75DWU received 18% less.

Irrigation Volume, Daily Water Use, and Plant Growth
Since the 75DWU irrigation rates in 2009 were based on the 100DWU irrigation rate it is possible to have different measured seasonal DWU (Table 2) than total amount of irrigation applied (Table 1) for 100-75DWU and 100-75-75DWU. Average DWU was highest in 2009 for 100DWU H. paniculata 'Limelight' and lowest for 100DWU A. arbutifolia 'Brilliantissima' ( Table 2). In 2009, only C. sericea 'Farrow' and I. virginica 'Morton' had differences within species due to treatment, where 100DWU was lower than control. In 2010, the highest DWU occurred for 6 of the 8 taxa in 100DWU or 100-75DWU treatments. Both Hydrangea species typically had the highest DWU among taxa within a treatment. The lowest water use taxa included S. meyeri 'Palibin' and V. dentatum 'Ralph Senior' in the control and 100-75 DWU, and S. xhyacinthiflora 'Evangeline' in 100DWU, 100-75DWU and 100-75-75DWU.     Since the 75DWU irrigation rates in 2009 were based on the 100DWU irrigation rate it is possible to have different measured seasonal DWU (Table 2) than total amount of irrigation applied (Table 1) for 100-75DWU and 100-75-75DWU. Average DWU was highest in 2009 for 100DWU H. paniculata 'Limelight' and lowest for 100DWU A. arbutifolia 'Brilliantissima' ( Table 2). In 2009, only C. sericea 'Farrow' and I. virginica 'Morton' had differences within species due to treatment, where 100DWU was lower than control. In 2010, the highest DWU occurred for 6 of the 8 taxa in 100DWU or 100-75DWU treatments. Both Hydrangea species typically had the highest DWU among taxa within a treatment. The lowest water use taxa included S. meyeri 'Palibin' and V. dentatum 'Ralph Senior' in the control and 100-75 DWU, and S. xhyacinthiflora 'Evangeline' in 100DWU, 100-75DWU and 100-75-75DWU.    For both years, DWU generally increased from June to mid-August, declined in September, and plateaued late September through October (Figures 3 and 4). In 2009 (Figure 3), DWU peaked on day 52 for all taxa and DWU exceeded the control irrigation rate for C. sericea 'Farrow' (20 mm), H. paniculata 'Limelight' (28 mm), P. opulifolius 'Seward' (25 mm), and W. florida 'Alexandra' (20 mm). Therefore, from day 52 until the next measurement of DWU (day 81), these taxa were receiving more irrigation than the control. No other taxa had DWU at or above the control on any other sampling day except W. florida 'Alexandra' on day 76. In 2010 (Figure 4), DWU peaked on Day 55 for H. arborescens 'Abetwo' (36 mm), H. paniculata 'Limelight' (43 mm), R. aromatica 'Gro-low' (32 mm), and W. florida 'Alexandra' (23 mm). For Hydrangea arborescens 'Abetwo', H. paniculata 'Limelight', and R. aromatica 'Gro-low', DWU peaked above the control irrigation rate for 19, 52, and 27 days, respectively. In contrast, many days preceding or following peak days had DWU below the control.

Crop Coefficients
Crop coefficients (KC) varied by month across the growing season (Tables 3 and 4). Differences in KC and KCadj demonstrate differences in water use efficiency due to over-or under-watering. KC or KCadj peaked on day 74 in 2009 (Table 3). Similarly, in 2010, the highest KC or KCadj was observed on days 78 to 81 depending on taxa (Table 4). When differences in KC or KCadj occurred in 2009, they were usually lower for one of the DWU treatments than the control although in a few instances KCadj was lower for the control than one of the DWU treatments (Table 3). However, there was no consistent pattern in differences. From July through October 2010, the highest KCadj was found for the 100-75DWU treatments in the high water use taxa, H. arborescens 'Abetwo', H. paniculata 'Limelight', and R. aromatica 'Gro-Low' (Table 4), which also had the highest season average DWU for these taxa ( Table 2). In contrast, low and moderate water use species usually had the lowest KCadj in 100-75DWU and highest KC or KCadj in the control or 100DWU, both of these treatments received less irrigation than 100-75DWU (Table 2). KC or KCadj was generally higher for plants receiving more water among high water users and decreased when low water users receive too much water. Using the classification system proposed by Warsaw et al. [13], A. arbutifolia 'Brilliantissima', C. sericea 'Farrow', 100DWU = 100% daily water use (DWU) replacement each day; 100-75DWU = 2-day cycle alternating 100% DWU and 75% DWU; and 100-75-75DWU = 3 day cycle with 100% DWU replacement the first day then 2 days 75% DWU replacement. Each day was analyzed separately (Tukey's test, α = 0.05, n = 18). Means followed by the same letters are not different. NS = not significant. Right y-axis indicates DWU where the entire bars represent DWU (mm) averaged from all treatments (n = 12); shaded portions of bars represent daily K C (DWU: ET 0 , n = 12). Overhead irrigation scheduling based on highest DWU of the 8 taxa in each treatment replicate each day. y Different letters denote significant differences between treatments in order from top to bottom: Control, 100DWU, 100-75DWU, 100-75-75DWU with A associated with the greatest mean value and subsequent letters associated with lower mean values.
In both 2009 and 2010, plant GI steadily increased in June and July, slowed between August and September, and reached a plateau in September to October as new growth began hardening in preparation for winter (Figures 3 and 4). There were no differences in GI in 2009 except for H. paniculata 'Limelight' where control was smaller than 100DWU from day 42 on and smaller than 100-75-75DWU on days 42 and 74, however, these differences were fairly small (Figure 3). No growth differences were observed on any measurement date throughout 2010 except the final date where H. paniculata 'Limelight' had smaller GI for 100-75-75DWU than other treatments, again the differences were minor (Figure 4). There were no differences in shoot dry weight for the three taxa evaluated in 2010 (data not shown).

Crop Coefficients
Crop coefficients (K C ) varied by month across the growing season (Tables 3 and 4). Differences in K C and K Cadj demonstrate differences in water use efficiency due to over-or under-watering. K C or K Cadj peaked on day 74 in 2009 (Table 3). Similarly, in 2010, the highest K C or K Cadj was observed on days 78 to 81 depending on taxa (Table 4). When differences in K C or K Cadj occurred in 2009, they were usually lower for one of the DWU treatments than the control although in a few instances K Cadj was lower for the control than one of the DWU treatments (Table 3). However, there was no consistent pattern in differences. From July through October 2010, the highest K Cadj was found for the 100-75DWU treatments in the high water use taxa, H. arborescens 'Abetwo', H. paniculata 'Limelight', and R. aromatica 'Gro-Low' (Table 4), which also had the highest season average DWU for these taxa ( Table 2). In contrast, low and moderate water use species usually had the lowest K Cadj in 100-75DWU and highest K C or K Cadj in the control or 100DWU, both of these treatments received less irrigation than 100-75DWU (Table 2). K C or K Cadj was generally higher for plants receiving more water among high water users and decreased when low water users receive too much water. Using
In 2009, the differences in substrate leachate pH due to treatment were less than 0.5 units and occurred only for C. sericea and W. florida on day 57 and I. virginica on days 57 and 120 (data not shown). Substrate leachate pH for all taxa and treatments in 2009 was 6.76 ± 0.03, 6.76 ± 0.05, and 7.18 ± 0.04 (mean ± SE) on days 16, 57, and 120, respectively. There were some minor differences in pH due to treatment early in 2010 but after day 59 the differences in pH were less than 0.5 units and only for S. meyeri (data not shown). Substrate leachate pH for all taxa and treatments in 2010 was 6.19 ± 0.05, 6.57 ± 0.07, 7.19 ± 0.07, 7.18 ± 0.05, 7.21± 0.04, and 7.59 ± 0.06 (mean ± SE) on days 2, 17, 34, 59, 79, and 111, respectively.

Foliar Analysis
In 2009, foliar P and/or K concentrations were higher for at least one of the DWU treatments compared to the control for all taxa except W. florida 'Alexandra' on day 90 (Table 5). Additionally, W. florida 'Alexandra' had higher foliar K for 100DWU and 100-75-75DWU than control on day 63 and P. opulifolius 'Seward' had higher foliar P and K concentration for all DWU treatments than the control on day 63. In 2010, the only differences in foliar analysis were lower foliar N concentrations in the 100-75DWU than 100DWU for H. paniculata 'Limelight' on day 36 and lower for 100-75DWU than all other treatments for V. dentatum 'Ralph Senior' on day 64 (Table 6). Foliar concentrations of other nutrients were general similar among treatments in both years (data not shown). Table 5. Foliar analysis (% dry wt.) sampled on days 58 and 85 of five taxa grown in 10.2 L containers and subject to four irrigation treatments from 11 June (Day 1) to 14 October 2009 (Day 126). Control = 19 mm·application −1 ; 100DWU = 100% daily water use (DWU) replacement each day; 100-75DWU = 2-day cycle alternating 100% DWU and 75% DWU; and 100-75-75DWU = 3-day cycle with 100% DWU replacement the first day then 2 days 75% DWU replacement. Overhead irrigation scheduling based on lowest DWU of the 8 taxa during each measurement period with remaining water requirement supplied by hand each day in DWU treatments as necessary. Recommendations nutrient ranges as % dry weight for woody ornamental plants are 2-4.5 for N, 0.2-0.6 for P, and 1.5-3.5 for K [23].

Discussion
Irrigating based on DWU by taxa ensures that plants are receiving sufficient irrigation for growth without over-or under-irrigating. In 2009, the seasonal volume of irrigation applied was less for all DWU-based treatments compared to the controls, suggesting most taxa in the control treatment were over-irrigated. When intermittent sampling techniques are used to estimate DWU, as in 2009, under-or overestimation could result in applying too little or too much water between sampling dates. However, scheduling plants with a wide range of water use needs based on the taxa with the highest water requirement in real time, as done in 2010, resulted in greater water use and over-irrigation of a majority of the taxa. In 2010, less water was used over the season for the control and 100-75-75DWU treatments. A. arbutifolia 'Brilliantissima', C. sericea 'Farrow' and H. paniculata 'Limelight' were under-irrigated for the 100DWU on days 16, 47, 20, respectively, while the other taxa were over-irrigated for the majority of the days. The two deficit irrigation treatments resulted in similar under-or over-irrigation for the same taxa. Yet despite the under-or over-irrigation as a result of sampling technique, there were very few and only minor differences in plant growth. These results and other studies demonstrate that container-grown plants are tolerant of these levels of deficit irrigation [6,7,13,24] indicating elasticity in growth response to irrigation.
While differences in growth among treatments were minimal for both years, the amount of water applied in 2009 was 7 to 57% less when irrigation was determined by taxa. Since irrigation applications were based on the highest DWU in 2010, the low and moderate water use species were over-watered for most days in 2010, particularly mid-season when K C or K Cadj peaked for high users. This emphasizes the importance of grouping plants with similar water use classifications, as suggested by Burger et al. [24] and Warsaw et al. [13], into irrigation zones that can be scheduled more precisely.
Crop coefficients have been shown to increase as plants grow [16], and they are also affected by changes in weather or season [16,17]. Calendar month serves as a convenient interval to reevaluate K C for seasonal changes due to plant growth and environmental conditions [17]. In both 2009 and 2010, most taxa had the lowest K C or K Cadj in June or July when GI was low before plant canopies had covered the substrate surface. As plants neared maximum GI, K C or K Cadj was generally highest from August to October. However, while K C or K Cadj of high-water users tended to have a large increase between June and August, the increase was less pronounced in low water use taxa. In a study of 12 hardy ornamental shrubs, plant size did not influence K C early in the growing season but correlated with K C when plant size had increased by July [25]. Schuch and Burger [17] found that K C of container-grown shrubs was relatively consistent across warm and cool phases of a production cycle for low water-use taxa but K C varied by as much as 3.7 between months for taxa that were high water users. Similarly, Irmak [26] showed that K C of Viburnum odoratissimum Ker.-Gawl was higher in fall than summer. In the current study, DWU decreased for all plants from late August to October while K C showed little change. While there are limitations in accurately scheduling irrigation from ET 0 using K C , K C is still valuable for grouping ornamental plants by relative water needs. This study reveals that higher water use plants exhibit greater seasonal changes in K C as related to periods of active growth and plant size than do lower water users.
Since the control plants received the most water of any irrigation treatment in 2009, difference in foliar analysis could be attributable to either leaching nutrients from the substrate, nutrient dilution within the plant, or a combination of both [27]. This may also have been the case for H. paniculata 'Limelight' 100-75 DWU treatment in 2010 which received the most water of all treatments. The lower nutrient content for the second date in each year could be due to the time of year when nutrients may be mobilizing out of leaves and the CRF is releasing at a slower rate due to cooler temperatures and less solar radiation incident upon the black containers. Since the later samples in 2009 and 2010 were taken only 91 and 64 days after application of a 5-to 6-month CRF, it is unlikely that the fertilizer was exhausted.
Contrasting the two scheduling methods used in 2009 and 2010, intermittent DWU quantification risks over-or under-estimation of crop water needs for the interval between measurements whereas real-time estimation compensates for changes in DWU at daily resolution. Additionally, real-time irrigation scheduling using DWU have shortened the production of Gardenia jasminoides by as much 64% at one production nursery in Georgia by increasing its rate of growth [28]. Other incentives for real-time scheduling include shortened crop production windows that free up space for new crops sooner, reduced losses from disease, and increased overall crop profitability [4].
Because K C values normalize for environmental variables [15], cross-season comparisons can be made for the same taxa. Two taxa were in common for both years. H. paniculata 'Limelight' was a high user in both 2009 and 2010 whereas W. florida 'Alexandra' was classified as a high-water user in 2009, but a low user in 2010. Using the same location and experimental design as the current study, Warsaw et al. [13] classified C. sericea 'Farrow', S. fritschiana 'Wilma' and W. florida 'Alexandra' as high users with K C = 3.4, 3.6 and 3.6, respectively, similar to this study. Several taxa in this and other studies [6,7,13,17,26] have seasonal K C above 4.0. As suggested by Warsaw et al. [13], as additional K C values are determined the water use classifications may require adjustment. We propose the addition of a "very high" category with K C ≥ 4.0 as the criterion, making the high category 3.0 ≤ K C < 4.0. H. paniculata 'Limelight', P. opulifolius 'Seward', and W. florida 'Alexandra' (2009) would be very high users under this classification. Seasonal K C for the 10 new taxa in this study add to the growing number of published K C values for container-grown woody ornamentals [6,7,13,17,26] further helping to establish indicator species for low, moderate, and high water use classifications that will aid in grouping plants into irrigation blocks based on similar water needs.
In a survey of nursery growers about the perceived benefits and limitations of wireless sensor networks, nearly half cited concerns that the systems may prove unreliable or not control irrigation correctly [29]. Likely, they worry that such complications would negatively impact plant growth and profitability. In a study of 24 ornamental shrub taxa, Warsaw et al. [13] showed only one plant in a deficit DWU treatment that grew less than the plants receiving 100% of their DWU. Their findings and those from the current study strongly suggest that a buffer exists in many ornamental plant species against light to moderate moisture deficits before growth reductions occur. Therefore, if minor malfunctions of sensing equipment or operator error should occur, losses in growth would likely be minimal or nonexistent if problems are corrected relatively quickly.
Grower-friendly systems are now being tested on commercial operations that permit multiple sensor inputs in a wireless network [30,31]. Used alone, these systems provide growers with additional data to make more informed scheduling decisions [28]. They also permit full automation of the irrigation scheduling process and have shown to reduce irrigation volume applied while increasing growth of ornamental trees in pot-in-pot production compared to "experience"-based irrigation [3]. Although widespread adoption of these systems may take several years, monitoring of every ornamental crop produced at most nurseries will not likely be economically feasible in the near future. Therefore, knowledge of relative water use classifications, derived by K C or other means, will be an essential part of effective sensor integration into existing irrigation infrastructure to help growers maximize water use savings and achieve greater crop profitability.

Conclusions
Scheduling irrigation based on plant daily water use can be used to reduce the amount of water needed for production of container-grown woody ornamentals with little or no effect on plant growth. When irrigation was applied based on daily water use for each taxon, including deficit irrigation treatments, the volume of irrigation required during the growing season was reduced between 7% and 57%. Reduced irrigation resulted in higher foliar %N and %P at times, most likely due to reduced leaching of nutrients out of containers. Scheduling irrigation based on the taxon with the highest water use (H. paniculata 'Limelight') resulted in over-irrigation for other taxa in the same irrigation zone, although with no practical effect on plant growth. H. paniculata 'Limelight' was also the only taxa for which there were irrigation treatments on growth, with slightly lower GI for control in 2009 and slightly lower GI for 100-75-75 in 2010. The limited effect of deficit irrigation or over-irrigation on plant growth indicates a high level of elasticity in plant response to irrigation and should be taken into consideration when discussing irrigation scheduling with practitioners. The knowledge of taxa-specific K C values will aid in grouping plants in irrigation zones based on water use to further increase water use efficiency.