Linking Remote Sensing and Geodiversity and Their Traits Relevant to Biodiversity—Part I: Soil Characteristics

In the face of rapid global change it is imperative to preserve geodiversity for the overall conservation of biodiversity. Geodiversity is important for understanding complex biogeochemical and physical processes and is directly and indirectly linked to biodiversity on all scales of ecosystem organization. Despite the great importance of geodiversity, there is a lack of suitable monitoring methods. Compared to conventional in-situ techniques, remote sensing (RS) techniques provide a pathway towards cost-effective, increasingly more available, comprehensive, and repeatable, as well as standardized monitoring of continuous geodiversity on the local to global scale. This paper gives an overview of the state-of-the-art approaches for monitoring soil characteristics and soil moisture with unmanned aerial vehicles (UAV) and airand spaceborne remote sensing techniques. Initially, the definitions for geodiversity along with its five essential characteristics are provided, with an explanation for the latter. Then, the approaches of spectral traits (ST) and spectral trait variations (STV) to record geodiversity using RS are defined. LiDAR (light detection and ranging), thermal and microwave sensors, multispectral, and hyperspectral RS technologies to monitor soil characteristics and soil moisture are also presented. Furthermore, the paper discusses current and future satellite-borne sensors and missions as well as existing data products. Due to the prospects and limitations of the characteristics of different RS sensors, only specific geotraits and geodiversity characteristics can be recorded. The paper provides an overview of those geotraits.

• Discuss approaches to monitor geodiversity and its traits (geotraits) with RS, • Define geodiversity and its characteristics, • Explain the concepts of spectral traits (ST) and the spectral trait variation (STV) approach applicable for monitoring issues, • Present the state-of-the-art technologies and capabilities of monitoring geodiversity and traits remotely, including: Soil characteristics (mineralogical characterization, pedology, and soil moisture) with different RS sensors, and • Provide a concise overview of those geo-traits that can be monitored using RS.

Understanding Geodiversity
Gray [61] defined "geodiversity" as the diversity of soil, geological, and geomorphological characteristics and the processes that lead to these characteristics. Other definitions of geodiversity integrate elements and characteristics of the lithosphere, the atmosphere, the hydrosphere, and the cryosphere, as well as their processes and interactions within and between the geo-components that are directly and indirectly related to biodiversity [62][63][64].
Geodiversity in this article is defined as the range and variability of geo-components and their intraspecific and interspecific interactions on all levels of organization of their geo-components. Geodiversity comprises components of the atmospheric, the terrestrial, the marine and aquatic ecosystems, and the ecological complexes to which they belong. Geodiversity is described by five characteristics that appear on all levels of organization and interact with each other. These are (see also (I) Geo-genesis diversity -GGD (which is described by the geo-genesis concept -GGC) represents the diversity of the length of evolutionary pathways, linked to a given set of geo-taxa. Therefore, geo-taxa sets that maximize the accumulation of geo-functional diversity are identified. (II) Geo-taxonomic diversity -GTD (which is described by the geo-taxonomic concept, GTaxC) -is the diversity of geo-components that differ from a taxonomic perspective. (III) Geo-structural diversity -GSD (which is described by the geo-structural concept, GSC) -is the diversity of composition or configuration of 2D to 4D geo-components. (IV) Geo-functional diversity -GFD (which is described by the geo-functional concept, GFC) -is the diversity of geo-functions and processes as well as their intra-and inter-specific interactions. (V) Geo-trait diversity -GTD (which is described by the geo-trait concept, GTC) -represents the diversity of biogeochemical, bio-/geo-optical, chemical, physical, morphological, structural, textural, or functional characteristics of geo-components that affect, interact with, or are influenced by the geo-genesis diversity, the geo-taxonomic diversity, the geo-structural diversity, or the geo-functional diversity.
diversity of geo-functions and processes as well as their intra-and inter-specific interactions. (V) Geo-trait diversity -GTD (which is described by the geo-trait concept, GTC) -represents the diversity of biogeochemical, bio-/geo-optical, chemical, physical, morphological, structural, textural, or functional characteristics of geo-components that affect, interact with, or are influenced by the geo-genesis diversity, the geo-taxonomic diversity, the geo-structural diversity, or the geo-functional diversity.

Figure 1.
Five characteristics of soil diversity: Soil diversity as part of geodiversity can be described under five characteristics: Soil-genesis diversity, soil-taxonomic diversity, soil-structural diversity, soil-functional diversity, and soil-trait diversity. All of the characteristics involve different levels of soil organization from elements, minerals, molecules, pedons, soil communities, polypedons, up to pedoshere. Different characteristics of the processes (the extent, process intensity, process consistency, resilience, and their characteristics) influence the resilience of soil and ecosystem health (modified after Lausch et al. [57]).

Approach for Monitoring Geodiversity by RS
There are two main methods to monitor geodiversity, (i) in-situ or field-based monitoring and (ii) RS. The in-situ approach refers to the direct quantitative and qualitative observation of the environmental spheres (pedosphere, lithosphere, atmosphere, hydrosphere, and cryosphere) either by direct measurements or by laboratory analysis of samples taken directly (destructive) in the environment. By contrast, RS approaches enable a non-destructive monitoring of the geocharacteristics without direct contact. The distance between the sensor and the object can range from a few millimeters to thousands of kilometers, enabling the coverage of diverse scales. In the RS case, the sensors are mounted or integrated onto platforms and can be used at many different scales. RS can be used in the laboratory (e.g., spectro-radiometers), in the field on the ground (e.g., Gamma Ray spectrometry and GPR-ground-penetrating RADAR) to sense features of the Earth's subsurface, or be hand-held or tower-mounted spectro-radiometers or thermal IR sensors (close-range RS techniques). A large range of sensors is airborne (mounted on UAV, in microlights, gyrocopters, or Figure 1. Five characteristics of soil diversity: Soil diversity as part of geodiversity can be described under five characteristics: Soil-genesis diversity, soil-taxonomic diversity, soil-structural diversity, soil-functional diversity, and soil-trait diversity. All of the characteristics involve different levels of soil organization from elements, minerals, molecules, pedons, soil communities, polypedons, up to pedoshere. Different characteristics of the processes (the extent, process intensity, process consistency, resilience, and their characteristics) influence the resilience of soil and ecosystem health (modified after Lausch et al. [57]).

Approach for Monitoring Geodiversity by RS
There are two main methods to monitor geodiversity, (i) in-situ or field-based monitoring and (ii) RS. The in-situ approach refers to the direct quantitative and qualitative observation of the environmental spheres (pedosphere, lithosphere, atmosphere, hydrosphere, and cryosphere) either by direct measurements or by laboratory analysis of samples taken directly (destructive) in the environment. By contrast, RS approaches enable a non-destructive monitoring of the geo-characteristics without direct contact. The distance between the sensor and the object can range from a few millimeters to thousands of kilometers, enabling the coverage of diverse scales. In the RS case, the sensors are mounted or integrated onto platforms and can be used at many different scales. RS can be used in the laboratory (e.g., spectro-radiometers), in the field on the ground (e.g., Gamma Ray spectrometry and GPR-ground-penetrating RADAR) to sense features of the Earth's subsurface, or be hand-held or tower-mounted spectro-radiometers or thermal IR sensors (close-range RS techniques). A large range of sensors is airborne (mounted on UAV, in microlights, gyrocopters, or airplanes), or spaceborne (mounted on satellites, space shuttles). The scope of this review paper focuses on the state-of the-art in monitoring geodiversity and traits using airborne and spaceborne RS sensors and approaches.
RS is capable of monitoring some geotraits and geotrait variations based on the principles of image spectroscopy across the electromagnetic spectrum from ultraviolet light to microwaves. Geo-traits can be directly or indirectly recorded using RS techniques in the time and space domains. Yet, in contrast to in-situ observations, RS approaches are not able to record all geotraits and geotrait variations of the Remote Sens. 2019, 11, 2356 7 of 51 entities in geodiversity [57] (Figure 2). The geotraits that can be monitored using RS techniques are therefore called spectral traits (ST) and the changes to the spectral traits are referred to as spectral trait variations (STV). Consequently, the related RS approaches are referred to as the remote sensing-spectral traits and -spectral trait variations-concept (RS-SV/STV-C), respectively ( Figure 2).
focuses on the state-of the-art in monitoring geodiversity and traits using airborne and spaceborne RS sensors and approaches.
RS is capable of monitoring some geotraits and geotrait variations based on the principles of image spectroscopy across the electromagnetic spectrum from ultraviolet light to microwaves. Geotraits can be directly or indirectly recorded using RS techniques in the time and space domains. Yet, in contrast to in-situ observations, RS approaches are not able to record all geotraits and geotrait variations of the entities in geodiversity [57] (Figure 2). The geotraits that can be monitored using RS techniques are therefore called spectral traits (ST) and the changes to the spectral traits are referred to as spectral trait variations (STV). Consequently, the related RS approaches are referred to as the remote sensing-spectral traits and -spectral trait variations-concept (RS-SV/STV-C), respectively ( Figure 2).

Figure 2.
In-situ and remote sensing approaches to monitor geodiversity and their characteristics by remote sensing and in-situ approaches (modified after Lausch et al. [57]).

Trends in Air-and Spaceborne RS for Assessing Soil Characteristics
This section provides an overview of the state-of-the-art for assessing and monitoring geodiversity and traits using airborne (UAV, airplanes) and spaceborne (satellite) RS sensors and approaches ( Figure 3). We present different technologies such as RADAR (radio detection and ranging) LiDAR, thermal, multispectral, and hyperspectral sensors that are deployed to record geodiversity and traits. We discuss current and future satellite-borne sensors and missions as well as existing data products that allow the respective geo-compartments to be monitored. In-situ and remote sensing approaches to monitor geodiversity and their characteristics by remote sensing and in-situ approaches (modified after Lausch et al. [57]).

Trends in Air-and Spaceborne RS for Assessing Soil Characteristics
This section provides an overview of the state-of-the-art for assessing and monitoring geodiversity and traits using airborne (UAV, airplanes) and spaceborne (satellite) RS sensors and approaches ( Figure 3). We present different technologies such as RADAR (radio detection and ranging) LiDAR, thermal, multispectral, and hyperspectral sensors that are deployed to record geodiversity and traits. We discuss current and future satellite-borne sensors and missions as well as existing data products that allow the respective geo-compartments to be monitored.

Mineralogical Characterization by RS
Biodiversity is strongly linked to the prevailing geological and soil characteristics, since bedrock and unconsolidated rock often determines the prevailing pedological and morphological conditions, which not least influence the distribution of animal and plants species, populations, and communities

Mineralogical Characterization by RS
Biodiversity is strongly linked to the prevailing geological and soil characteristics, since bedrock and unconsolidated rock often determines the prevailing pedological and morphological conditions, which not least influence the distribution of animal and plants species, populations, and communities [6,65]. Therefore, mineralogy and soil properties play important roles in determining the ecology of an area through their contribution to the formation of soils and their influence on relief. Both factors can influence local climatic conditions, water availability, and thus the vegetation type and structure. In addition, geology will determine different landforms such as limestone karst, cliffs, and escarpments, which offer a range of different habitats and thus are key to the distribution of flora and fauna.
Space-and air-borne multi-and hyperspectral sensor data can significantly contribute to an area-wide understanding of land cover patterns and vegetation distribution and their changes, which can ultimately alter biodiversity on all spatial scales [48]. Comprehensive overviews of how image spectroscopy has aided the characterization and mapping of soil mineral composition are provided by Mulder et al. [66] and Wulf et al. [49]. Key characteristic surface mineralogical properties, which can be derived from optical RS data in the visible (VIS, λ = 0.4-0.7 µm), near infrared (NIR, λ = 0.7-1.0 µm), and short-wave thermal infrared (SWIR, λ = 1.0-2.5, 5-14 µm) are clay, sand, carbonate, silicate, sulphate, or iron content (see Table 1). They can be mapped using imaging spectroscopy, for example, with the airborne visible/infrared imaging spectrometer (AVIRIS; [49,67,68]). To access the characteristic absorption features, hyperspectral data are required, which limits the applicability of RS for this issue. EO-1 Hyperion (= Earth Observing-1 Hyperion; [69]) has been the only operational spaceborne imaging spectrometer to date that covers the full spectral range, including the short-wave infrared region for λ > 2 µm. Hyperion data were used, for example, to map iron-bearing minerals on tailings, dams, and areas affected by mining in South Africa (Figure 4; [70]). The surface cover of a platinum mine tailings facility was characterized with the United States geological survey (USGS) material identification and characterization algorithm (MICA; [71]) that analyzes the characteristic mineral absorption (Figure 4a). Complementary, the characteristic absorption feature of iron-bearing pyroxenes around λ = 900 nm was deployed to trace these minerals over a wider area by applying the three-point band depth index "iron feature depth" (IFD) to the multispectral data of Landsat-8 OLI [72]. Figure 4 shows the results of this approach, illustrating the close match to the hyperspectrally mapped iron bearing surface cover types shown in Figure 4a. Further capabilities to map iron were opened with the launch of the Sentinel-2 satellites [73], since the multispectral instrument (MSI) covers important iron absorption features [74]. Another example for the use of Hyperion data is the detection and quantification of different salt types that is combined with the detection of stable and dynamic areas at the surface of a Namibian salt pan on the basis of a 30-year-long Landsat time series [75]. The combination of multispectral time-series for monitoring dynamic processes with hyperspectral observations for soil and sediment characterization enabled the extraction of new knowledge on the salt pan crust development.
Currently, there is no spaceborne hyperspectral TIR sensor available. With the future surface biology and geology (SBG) mission, the mapping of more minerals will be enabled, due to the larger number of bands in the VIS-NIR-SWIR-TIR region and the broad swath-width, which will allow for more levels of information from complementary spectral regions in a synergistic fashion. Silicate minerals, for instance, demonstrate a distinct emittance minimum caused by fundamental Si-O stretching vibrations occurring near λ = 10 µm. Quartz and feldspar have emittance minima at shorter wavelengths (λ = 9.3 and 10 µm, respectively) than sheet silicates such as muscovite (λ = 10.3 µm) or chain silicates such as the amphibole minerals (λ = 10.7 µm; [81]). Carbonates have features associated with CO 3 internal vibrations both at λ = 11.4 and 14.3 µm due to C-O bending modes [82]. Sulfate minerals on the other hand have an intense feature near to λ = 8.7 µm caused by fundamental stretching motions [83]. The thermal range of the spectrum has been demonstrated to be very important for an improved determination of sand, clay, and organic carbon content in soils [84,85]. In addition to mineral mapping, the SBG mission should provide important information for soil textural and mineralogical characterization, for example, for sandy soils that are common in semi-arid landscapes.
Currently, several TIR hyperspectral airborne sensors are deployed for geological and soil mapping, including the spatially enhanced broadband array spectrograph system (SEBASS; [86,87]), the airborne hyperspectral scanner (AHS; [88]), AisaOWL [89][90][91], the thermal airborne spectrographic imagery (TASI; [92]), and the hyperspectral camera HyperCam [93]. In summary, hyperspectral RS allows for the discrimination of similar minerals with spectrally adjacent absorption features. Calcite-dolomite ratios, for example, were mapped in the SWIR using the GER 63-channel imaging spectrometer data (GERIS; [94] and by SEBASS in the TIR [95]. This type of information can thus be used to monitor and quantify the surface mineralogy that has an impact on biodiversity, from local to global scales.
Geologists have been using RS data since the introduction of RS technology to describe the status as well as the processes of the geology, geo-chemistry, and mineralization of an area, which are the basis of geological mapping, for structural interpretations, and for mineral resource mapping [96,97]. Due to the scope and manifold applications of RS for geological investigations, reference is only made to central work on this subject [78].

Pedology
Soil is the uppermost, weathered layer of the Earth's crust. It is a complex matrix at the interface of the lithosphere, biosphere, hydrosphere, and atmosphere governed by different site-specific characteristics. Soil provides the basis for plant growth, which among others is controlled by the physical, chemical, and biological properties of the soil. Soil-related factors like nutrient availability, carbon content, or soil structure affect biodiversity on the one hand, but biodiversity also determines

Pedology
Soil is the uppermost, weathered layer of the Earth's crust. It is a complex matrix at the interface of the lithosphere, biosphere, hydrosphere, and atmosphere governed by different site-specific characteristics. Soil provides the basis for plant growth, which among others is controlled by the physical, chemical, and biological properties of the soil. Soil-related factors like nutrient availability, carbon content, or soil structure affect biodiversity on the one hand, but biodiversity also determines the spatial soil heterogeneity on the other hand [48,99]. As a consequence, there are strong interactions between biodiversity and soil characteristics.
Soil traits can be monitored by RS approaches either directly (from bare soil, (Figure 4, Figure 5, and Table 1) or indirectly, i.e., through vegetation as a sensor or proxy of soil characteristics [48,100]). Thus, direct or indirect indicators derived from remotely sensed satellite or airborne data have been proven to be important for the prediction of soil variables, classes, or processes [101]. They are used in different digital soil mapping approaches, which combine soil-related factors or which are based on environmental similarities or correlations [102].
Following comprehensive reviews of remotely sensed variables for digital soil mapping [48,[103][104][105], remotely sensed indicators can be distinguished according to their spatio-temporal resolution or the respective sensor properties, which can refer to active or passive systems. Active systems like RADAR, LiDAR, or SONAR (sound navigation and ranging) are mainly used to detect surface properties or to estimate soil moisture content (see also chapter 4.3; [106]). Passive systems relate to reflectances and emitted radiation using the VIS-NIR-SWIR, the far-infrared, and the microwave region. Direct reflectances of bare soils enable the prediction of top-soil variables like soil texture, soil organic carbon, iron content, or heavy metals in plants as well as soil salinity or carbonates, the classification of soil types, or even hydraulic properties [107,108]. Since soils are mostly covered by vegetation, spatio-temporal variations in vegetation indices can also help to predict soil properties [109].
Imaging spectroscopy for mapping soil characteristics in the VIS-NIR-SWIR domain emerged at the beginning of the 21st century [110,111]. It builds upon many years of extensive research in soil spectroscopy under laboratory conditions (e.g., [111],) which revealed quantitative and qualitative relationships between the spectral signal and the chemical and physical properties of soils [112]. Castaldi et al. [113] recently showed that narrowband, hyperspectral imagers provide significantly higher potential for the quantitative estimation of soil variables compared to multispectral sensors because broadband instruments cannot resolve diagnostic spectral features of the soil spectrum. On the other hand, recent works show the potential of the Sentinel-2 sensor for soil organic carbon mapping [99,113]. For both laboratory and imaging spectroscopy, mostly non-parametric regression algorithms are applied [114]. Focusing on single mineral absorption features has been proven to be less successful due to the complex nature of soil [115]. Although automatic methods such as the HYSOMA (hyperspectral soil mapper) algorithms [116] are based on the direct analyses of the spectral signal, they proved to be more generic and have the potential to be transferable to the regional-global scale [117,118].
Compared to in-situ approaches, there are several items that make the remote spectroscopy of soils challenging. Firstly, optical RS captures only the properties of the uppermost centimeters of the top-soil and cannot provide information about the entire soil body. Secondly, atmospheric effects and sensor constraints [119], soil moisture contents [120], soil roughness, or soil surface coverage by vegetation or plant residues [117] can interfere with the spectral signal [121].
Vegetation cover is a major challenge in RS aided soil trait estimation in many parts of the world. Ouerghemmi et al. [122] reported that prediction errors increased progressively as vegetation cover exceeded 5% to 10%. The majority of hyperspectral studies therefore focus on bare soils accepting that a noticeable amount of cultivated area cannot be mapped instantly. The associated limitations can be reduced, either by the multi-temporal stacking of soil maps [123], pixel compositing [124], or by applying advanced algorithms to correct for the impact of vegetation [125] and soil moisture [126]. Beyer et al. [127] for example, suggested calculating a residual soil signature to reduce the influence of non-soil materials. Another strategy for areas with permanent vegetation may be to infer information on the underlying soil by exploring the spectral signal of the vegetation itself [128,129]. In this indirect approach, vegetation traits are used as indicators for the status or changes of soil traits [100].
Irrespective of the limitations of imaging spectroscopy, its potential to provide quantitative and qualitative soil information is demonstrated by several studies: Ben-Dor et al. [130] were some of the first to apply hyperspectral image data from the DAIS 7915 airborne scanner for the mapping of soil characteristics of clayey soils in Israel. They obtained reliable predictions for soil moisture, soil salinity (electrical conductivity), soil saturated moisture, and organic matter content, disclosing the spatial distribution of soil properties in the study area. Selige et al. [131] estimated the percentage of soil organic carbon, total nitrogen, sand, and clay of agricultural fields on a test site in East Germany from image data of the Australian HyMap sensor. Vohland et al. [132] used HyMap data to estimate soil organic carbon and, as a sub-fraction, microbial biomass-carbon for agricultural soils. The soil organic carbon content has also been explored in several other studies, for example by Stevens et al. [118] on Central European croplands in Luxembourg using the airborne hyperspectral scanner AHS-160 or by Castaldi et al. [113], who analyzed data from the airborne prism experiment (APEX) acquired in Belgium and Luxembourg. Deploying the Aisa/DUAL system, Kanning et al. [133] estimated soil organic carbon (SOC) [134] and the soil particle fraction sand, silt, and clay. Furthermore, Paz-Kagan et al. [135] developed a spectral soil quality index (SSQI) using airborne imaging spectroscopy. Selige et al. [131] and Vohland et al. [132] showed that nitrogen (an essential nutrient for plant growth) can be estimated by imaging spectroscopy. Further studies on nitrogen have been conducted under standardized conditions in the laboratory [136]. Imaging spectroscopy can also provide some insights into the pedogenesis processes, land degradation, and erosion processes by mapping the iron content as shown by [137], or by direct mineralogical mapping in association with erosion and deposition stages [138]. Applications on soil crust mapping were also successful for physical crust [139] and biological crust discrimination in the laboratory and the field [140], although at the imagery scale these applications are still challenging and scarce.
In terms of multi-spectral imagery provided by satellite sensors, a multitude of approaches have been successfully applied using sets of environmental covariates over recent decades [48]. With the increased spatio-temporal availability of remotely-sensed imagery and other auxiliary data such as digital elevation models, "digital soil mapping has shifted from a research phase into operational use" [102]. Since RS imagery is affected by different atmospheric conditions and soil moisture variations on image acquisition dates, the transferability of digital soil mapping solutions remains a key challenge [141]. To this end, the derivation of indicators based on the analysis of multi-(three or more images) or hyper-temporal imagery (e.g., images for one or many years; [105]) may be useful. Multior hyper-temporal image data can be obtained from satellite image archives [142] or be generated by applying data fusion algorithms [143]. Maynard and Levi [105] for example, were able to show that hyper-temporal time series of a vegetation index based on Landsat imagery enable typical and temporally stable spectral fingerprints to be derived, which significantly increased the prediction accuracy of soil texture. Especially in agricultural regions, multi-and hyper-temporal imagery enables bare soil areas to be detected at different time steps. As a result, regional [144] and even national bare soil composites [145,146] can be created and used for soil mapping parameterization.
Blasch et al. [147] applied multi-temporal RapidEye composites to predict spatial variations in soil organic matter (SOM). Multi-temporal Landsat image composites have also been used by Dematte et al. [148] in order to perform soil texture classification. Since soil-related indicators based on multiand hyper-temporal time series or composites are less affected by atmospheric conditions and soil moisture variations, they can be considered as a key for the automatic and operational derivation of standardized soil mapping products. Standardization means that all steps of geodata processing are reproducible, harmonized protocols are applied, and that the soil mapping products are evaluated by accuracy metrics [149]. This concerns both digital soil mapping parameterization and the provision of scale-specific harmonized and representative training samples [150,151].
on multi-and hyper-temporal time series or composites are less affected by atmospheric conditions and soil moisture variations, they can be considered as a key for the automatic and operational derivation of standardized soil mapping products. Standardization means that all steps of geodata processing are reproducible, harmonized protocols are applied, and that the soil mapping products are evaluated by accuracy metrics [149]. This concerns both digital soil mapping parameterization and the provision of scale-specific harmonized and representative training samples [150,151].  The RS-aided derivation of soil characteristics and soil traits shown in Table 1 and Figure 6 shows the enormous number of current and future space-based RS missions and satellites for monitoring soil characteristics and traits with information about the mission status, according to the CEOS database (Committee on Earth Observation Satellites).    Sensor is used on the RS platform: UAV 1 -unmanned aerial vehicles (UAV) ; airborne 2 -airborne RS platform ; spaceborne 3 -spaceborne RS platform.
See Chabrillat et al. [121] for a recent review of the potential of upcoming spaceborne hyperspectral imagery for global soil mapping and monitoring. Table 2 provides an overview of the recent and future spaceborne hyperspectral missions with their sensor characteristics deriving soil characteristics and traits.  Table 3 provides an overview of current and future important missions of the Copernicus program -Sentinel satellites monitoring geodiversity and their traits developed by ESA. For more information about the sensor characteristics of the Sentinel Satellites, see also Malenovský et al. [204].   [205]. Table 4 gives a selection of RS-aided data products of soil characteristics.   Table 4 gives a selection of RS-aided data products of soil characteristics.

Soil Moisture by RS
Soil moisture is part of the hydrological cycle, even if it only accounts for 0.001% of the global water storage, it still has a crucial effect on compositional, structural, and functional biodiversity [101,214,215]. Volume distribution and temporal dynamics of moisture within the soil are key drivers of biodiversity. Part of the soil water (the so-called plant available soil water) is accessible to plants for metabolic processes. The temporal and spatial dynamics of soil moisture reflecting the interplay between evaporation, transpiration, infiltration, and the replenishment of groundwater are imperative to understand water movements in the soil and the water cycle at local to global scales [216].
A reliable and adequate description of the spatial and temporal heterogeneity of soil moisture is one of the most critical challenges in the monitoring and modelling of water and energy fluxes, nutrient transport, and matter turnover within soil landscape systems. In order to generate reliable predictions about these processes from models, sufficient knowledge needs to be gained about the soil heterogeneity and moisture patterns at larger scales [217]. Furthermore, soil moisture is a key variable controlling biosphere-pedosphere/process-pattern interactions, hydrological, and biological processes, not to mention climate and other ecosystem processes including the dynamics in biodiversity [218].
Vereecken et al. [101] found that soil moisture was the most important driver, accounting for 65% of the variation in ecosystem multi-functionality. Moeslund et al. [219] found that soil moisture drives plant diversity patterns within grasslands. They calculated relative ecological indicator values for soil moisture and found pronounced differences in the responses of grassland types to grazing depending on the moisture regimes with an increasing effect of grazing from the dry towards the wet grassland types [219]. Especially in extreme ecosystems, e.g., the dry valleys of Antarctica, spatial soil moisture and soil carbon patterns dominate the distribution of soil organisms [220]. Established patterns of soil biological communities due to changes in soil moisture and temperature also vary over seasonal or larger time scales and alter the resource supply for the growth of plants. Droughts, for example, change the relationship between biodiversity and ecosystem functioning [221]. Moreover, projected increasing climate-induced drought severity suggests that changing tree and forest biogeography could substantially lag habitat shifts that are already underway [222]. One interesting example is that of nematodes (roundworms) that can enter anhydrobiosis and become inactive when the soil is dry. Therefore, their activity is closely related to levels of soil moisture, and the proportion of anhydrobiotic nematodes increases with the distance from stream sediments to drier soils [223].
Soil moisture measurement techniques have been recently reviewed by Babaeian et al. [50]. In general, there are two different approaches that use RS techniques to understand soil characteristics, their hydrological traits and eventually the spatial and temporal dynamics of soil moisture: (i) Direct measurements for areas where the soil has no or limited vegetation (using active and passive microwave sensors as well as optical RS data, Figure 7) and (ii) indirect measurements where vegetation covers the soil, using vegetation traits as a proxy of the soil and soil moisture traits. Optical RS data are preferred for indirect measurements of soil moisture patterns. In general, spaceborne RS of soil moisture is very advanced at global scales [51] with high accuracy [224] and a large variety of applications [225]. For local scales, adequate spatio-temporal resolutions are less common and tend to be the domain of airborne observations or proximalsensing, like cosmic ray neutron sensing [226] or ground penetrating RADAR [227,228].

Soil Moisture Characteristics using Active and Passive Microwave RS Approaches
Surface soil moisture estimation using microwave RS is based on the different dielectric properties (ε) of dry soil (ε~4) and liquid water (ε~80) [229]. However, extracting the part of the microwave signal that actually originates from the soil water is a challenging task. Not only sensor characteristics such as polarization, incidence angle, wavelength, and spatial as well as temporal resolution, but also surface characteristics such as vegetation cover with its shape, orientation, as well as water content and soil surface roughness have a distinct influence on the microwave signal that is received.
Both active and passive microwave systems are sensitive to the complex dielectric constant of the soil rather than to soil moisture directly. Here, soil dielectric mixing models have been developed for their linkage, e.g., by [230][231][232]. All models rely on the separate contribution of bound water and free water to compute the complex dielectric constant of the soil. The empirical approach by [230] has the advantage that it does not require information on soil texture, although it only appears to be satisfactory for coarse-textured soils. The semi-empirical approach by [231] is valid for a large range of microwave frequencies (1-18GHz), but is restricted to a certain range of soil textures. The Mironov approach [232] on the other hand is based on the refractive mixing model for moist soils. It is validated for a large range of soil textures and therefore suitable for global-scale applications (cf. SMOS [225] ( Figure 7a) and SMAP [233] satellite missions).

Active Microwave Sensors (RADAR, Scatterometers)
Active microwave sensors (RADAR, scatterometers) send microwaves using their antenna devices and are therefore independent of illumination and weather [234]. Due to the active transmission, the strength of the signal is distinctively higher than that of passive microwave sensors (radiometers), which depend on the weak natural emission of the radiation from Earth. Hence, the recorded microwave signal is not directly prone to system noise or exposed to external radiation sources/interferences due to the strength of the active signal compared to passive devices. Moreover, the higher radiation budget of the recorded scattering at the active sensors also allows a shorter integration time and therefore a distinctively higher spatial resolution in terms of meters (in case of synthetic aperture RADAR, SAR) compared to the kilometer range for passive sensors. However, the calibration of the active sensors is much more complex with a transmitting and a receiving antenna part in contrast to passive devices.
Active sensors pick up the backscattered microwave radiation, normally measured in normalized backscattering coefficients or sigma noughts σ 0 (normalized RADAR cross sections), returning from interaction with media on Earth [235]. For soil moisture estimation from natural surfaces, the signal recorded at the sensor is a mixture of scattering contributions. For bare soil surfaces, it is not only the dielectric constant (soil moisture), but also the soil roughness that influences the scattering of the microwaves depending on their strength [229]. The scattering scenario becomes more complicated with the occurrence of vegetation cover, where plant geometry, density, and moisture are influential factors on the scattering of the microwaves [236]. It is mainly a function of the wavelength as to how deep the microwaves can penetrate natural media. Hence, for soils with vegetation cover, longer wavelengths, such as L-band (23 cm), are preferred. This also enables soil moisture to be retrieved under vegetation cover [237] -at least until a certain density and wetness of the vegetation cover. In order to extract from the recorded backscattering signal, only the scattering component triggered by the moisture of the soil, decomposition methods are essential to invert soil moisture from the corresponding part of the signal. Polarimetry offers an exploitable observation space for the decomposition of RADAR measurements to invert for soil moisture [238][239][240].
Moreover, there are a plethora of algorithms for estimating soil moisture from active microwave RS, and a non-exhaustive overview is provided below [241][242][243]. The algorithms can mainly be divided into empirical (including machine-learning) [244][245][246][247][248][249][250], semi-empirical [251][252][253][254], and physical-based [229,255,256] retrieval techniques [237]. While empirical and semi-empirical algorithms can only be performed successfully in environments, where their empirical relationships were established, physical-based methodologies are generally valid and do not depend on test site characteristics. The inclusion of vegetation scattering comes with the selection of an appropriate model. Here, a range from more simple [257][258][259] to sophisticated [260,261] scattering models can be applied. However, the possibility of inversion decreases with the degree of complexity of the model due to an increase in variables for vegetation cover with a constant number of observations. Hence, the available observation space (multi-temporal, multi-angular, multi-frequency, multi-polarimetric) determines the complexity and performance of the inversion algorithm for soil moisture (under vegetation cover).
Beyond intensity-and polarimetry-based backscatter as well as machine-learning methods, another active microwave methodology to retrieve soil moisture dynamics in space and time is differential SAR interferometry (DInSAR). Ref. Morrison et al. [262][263][264] found significant dependence of the interferometric phase on changes in soil moisture. It was not possible to explain this relation by swelling soils and changes in penetration depth. The behavior of the phase points towards changes in volume scattering within the soil, which might also explain moisture-related temporal decorrelation [264].
At the global scale, a prominent active sensor system to estimate soil moisture is the advanced scatterometer on MetOp (ASCAT) [265,266]. Backscatter measurements at six different azimuth angles are used to calculate soil moisture by the change detection algorithm after Naeimi et al. [267]. The angular information is used to characterize the vegetation contribution and its temporal variability to be eliminated before soil moisture inversion. In contrast to this 12.5 km posting product, a 1 km product is available from Sentinel-1 [268] or a combined ASCAT-Sentinel-1 approach [269]. It has to be noted that the data are expressed in relative units (degree of saturation).

Passive Microwave Sensors
Passive microwave sensors (radiometers) record naturally emitted microwave radiation, usually expressed in brightness temperatures (Tb), i.e., the product of emissivity (e) and the physical temperature of the target (T). This implies that the physical temperature of the target needs to be known for soil moisture retrieval. The emissivity of a smooth surface can be predicted by the Fresnel reflection equations. Accordingly, it depends on the incidence angle and the complex dielectric constant of a soil. Rough soils behave differently, because roughness decreases the reflectivity and thus increases emissivity. In theory, with an increased surface area, rough targets can emit more thermal energy. In field applications, the effective soil roughness is probably more related to the distribution of water in the topsoil rather than a pure geometric soil surface roughness as the latter can only occur when the soil is very wet. In addition to the general problematic consideration of the incidence angles within one radiometer footprint, this issue affects the soil moisture retrieval, especially in mountainous areas and needs to be accounted for in the retrieval methodology.
Another factor causing attenuation of the microwave emission is vegetation. Moreover, vegetation adds its contribution to the land surface emission signal [270]. To simulate these effects, a simple approach referred to as the τ-ω model [271], was developed. The τ-ω model is a zero-order solution of the radiative transfer equation [272] and uses only two variables for canopy characterization -the vegetation optical depth (VOD) τ and the single-scattering albedo ω. While the first one parameterizes the attenuation effects, the latter one describes the scattering effects within the canopy [273]. The optical depth is often linearly related to vegetation water content (VWC), [274] and can to a certain extent be derived from leaf area index or multispectral vegetation indices [275]. At L-band, soil moisture retrieval is only possible up to VWC of 5 kg/m 2 [276]. In addition, to the sole and direct relationships of plant biodiversity with soil moisture patterns, vegetation is additionally involved in soil moisture retrieval. This additional component needs to be separated when soil moisture and biodiversity dependencies should be investigated exclusively.
There are several approaches for soil moisture inversion and most of them are based on the vegetation attenuation concept of Mo et al. [271] e.g., the normalized polarization difference (NPD) algorithm, the single-channel algorithm (SCA), the L-band microwave emission of the biosphere (L-MEB) model, the community microwave emission model (CMEM), the land parameter retrieval model (LPRM), and the University of Montana (UMT) approach [277][278][279]. It is worth noting that spaceborne brightness temperature products are typically only valid for the top-of-atmosphere (except for the SMAP products). To estimate surface level brightness temperatures before soil moisture inversion, atmosphere attenuation, upward atmosphere emissivity, and the polarization rotation according to the Faraday theory need to be considered.
Global scale soil moisture monitoring is provided by the Japan Aerospace Exploration Agency (JAXA) Global Change Observation Mission-1st Water (GCOM-W1) satellite that hosts the AMSR2 sensor [280]. Together with its precursor AMSR-E, the time series, with a gap of a few months, expands from mid-2002 until present. Further systems are the soil moisture and ocean salinity (SMOS) [281] and soil moisture active and passive (SMAP) missions [233,282], started in 2009 and 2015, respectively.

Combining Active and Passive Microwave Sensors
For the long-term analysis of surface soil moisture within the water and energy feedbacks of the climate system, large time series were generated and developed. The ESA Climate Change Initiative (CCI) combines active and passive microwave observations to obtain a consistent time series from 1978 until mid-2018 [283]. According to their suitability for soil moisture retrieval, C-band scatterometers (ERS-1/2 scatterometer, ASCAT) and multi-frequency radiometers (SMMR, SSM/I, TMI, AMSR-E, Windsat) were merged at the level of retrieved surface soil moisture data (Level 2) to avoid problems arising from different sensor specifications.
With the launch of NASA's missions Aquarius and SMAP, both active and passive microwave remote-sensing observations are combined to improve the spatial resolution [284]. The combination extracts the relative advantages of the two sensing techniques, as there is a tradeoff between resolution and soil moisture sensing sensitivity between active and passive microwave measurements [285]. Fusion methods include temporal change detection methods [286], Bayesian merging approaches [287], statistical disaggregation [285,288,289], and physics-based covariation algorithms [290][291][292]. Other methods retrieve vegetation variables from active microwave measurements for the utilization of passive microwave soil moisture inversion [293,294]. Unfortunately, the RADAR on the SMAP satellite went out of service on 7 July 2015. Therefore, combined active-passive microwave data were only recorded over the first months of the mission (April-July 2015). Due to the failure of SMAP's L-band RADRA, the substitution with ESA Copernicus Sentinel-1 s C-band RADAR is in final preparation [295] and a first 3 km soil moisture product was released in November 2017 [295]. Due to Sentinel-1 s varying acquisition geometry and the reduced amount of coincident active-passive overpasses, the standard statistical (time-series) downscaling approach using passive brightness temperatures with active RADAR backscatter is no longer possible and a physics-based downscaling was developed by [295,296] for a dual-frequency downscaling.
Moreover, soil moisture sensing is also a topic of discussion for airborne microwave RS working on resolutions that are valid for habitat research and precision farming and capable of retrieving soil moisture patterns with a high spatial resolution (<100 m). On airborne platforms joint active-passive remote-sensing instrument constellations include PLMR (polarimetric L-band multibeam radiometer) [178,297,298] (Figure 7c) and F-SAR sensors [299,300] (Figure 7d), also referred to as multi-sensor platforms.
With these multi-sensor platforms, [301] analyzed different active-passive fusion methods and their advantages and drawbacks for a central European region. In Australia, the SMAPEx campaigns (e.g., [302]) provided first combined observations of PLMR and PLIS (polarimetric L-band imaging synthetic aperture RADAR). In North America, the PALS (passive and active L-and S-band sensor; [303]) recorded soil moisture from RADAR and radiometer to analyze soil moisture heterogeneity across scales in several campaigns. Table 5 provides an overview of recent satellite missions and airborne systems for soil moisture estimation. With these sensors, soil moisture retrieval is only possible for the top few cm of the soil surface, namely the surface soil moisture. Root zone soil moisture, which to a larger extent effects the biodiversity of a habitat rather than the water content of the top layer, can be estimated by additional methods. Examples are: Direct retrieval by longer wavelengths such as P-band [304], surface soil moisture assimilation into a hydrological model [305][306][307][308][309][310], or data-driven methods such as neural networks [311] to improve root zone soil moisture estimates. Moreover, indirect methods use the plants as "sensors" of root-zone properties. Wilson [312] can therefore be used to gain knowledge about root zone soil conditions (see also Chapter 4.3.5) and Rudolph [313], for example, presented the link between crop-status patterns in large-scale multispectral satellite imagery with multi-receiver electromagnetic induction (EMI) hydro-geophysical data. and Rudolph [313], for example, presented the link between crop-status patterns in large-scale multispectral satellite imagery with multi-receiver electromagnetic induction (EMI) hydrogeophysical data.

Direct and Indirect Measurements by Optical and Thermal Sensors
To obtain and assess the direct approaches to assess spatial and multitemporal surface soil moisture data, in addition to microwave RS techniques, there are also various optical sensors such as MODIS [315], Landsat [316], hyperspectral RS sensors (HyMap, [317]), as well as thermal infrared sensors (Landsat, Sentinel-3, or SEVIRI [318], Table 2). Since soil moisture is subject to a very high spatial-temporal variability, the suitability of optical and thermal sensors to derive soil moisture related information very much depends on the spatial, spectral, and temporal resolution of the RS

Direct and Indirect Measurements by Optical and Thermal Sensors
To obtain and assess the direct approaches to assess spatial and multitemporal surface soil moisture data, in addition to microwave RS techniques, there are also various optical sensors such as MODIS [315], Landsat [316], hyperspectral RS sensors (HyMap, [317]), as well as thermal infrared sensors (Landsat, Sentinel-3, or SEVIRI [318], Table 2). Since soil moisture is subject to a very high spatial-temporal variability, the suitability of optical and thermal sensors to derive soil moisture related information very much depends on the spatial, spectral, and temporal resolution of the RS sensors. Furthermore, soil moisture is a very dynamic parameter along the soil profile, and passive and active RS sensors can sense soil moisture at different depths. In addition, to the sensor characteristics, an extensive acquisition of in-situ data on soil moisture is also required. Such in-situ measurements (previously conducted manually using close-range sensors) are often insufficient due to the tremendous spatial and temporal variability of RS data. To gain access to in-situ data with a high temporal and spatial resolution as well as to soil moisture data from different locations with differentiated land-use-land-cover and soil characteristics, the development and implementation of distributed soil moisture sensor networks is imperative to achieve an improved calibration and validation of airand spaceborne RS data. Some promising preliminary approaches have already been made in this respect [319].
Soil moisture is difficult to determine in most European regions using the direct approach with active and passive microwave and optical sensors, because the soil is mostly covered with vegetation that varies in height, density, and plant species composition over the entire year [100]. Therefore, vegetation and its biochemical, morphological, physiological, or functional characteristics of plants or spectral plant traits are used as a proxy for determining soil moisture patterns and soil characteristics in space and time [100] (see also Figure 8).
Remote Sens. 2019, 11, x FOR PEER REVIEW 24 of 53 that varies in height, density, and plant species composition over the entire year [100]. Therefore, vegetation and its biochemical, morphological, physiological, or functional characteristics of plants or spectral plant traits are used as a proxy for determining soil moisture patterns and soil characteristics in space and time [100] (see also Figure 8).

Airborne Geophysical Sensors of Natural Radiation-Gamma and Cosmic-Ray Neutron Sensors
Natural sources of radioactivity or cosmogenic radiation are increasingly used in environmental sciences for the spatial exploration of soil properties. While the space-borne detection of gamma and albedo-neutron radiation from satellites became a standard method to map the soil water distribution on Mars [320], the Earth's atmosphere allows for the detection of terrestrial radiation only with car-borne or low-flying airborne vehicles. Nevertheless, the much deeper soil penetration and the effortless installation are the main advantages of gamma and neutron sensing compared to optical techniques.
Sources of gamma radiation are radioactive isotopes in the ground, such as potassium-40, thorium-232, and their daughters in the decay chain. Their signal is attenuated by any material, but particularly by the water molecule. The gamma rays of the corresponding energy window can travel up to 30 cm in the ground and 25 m in the air [321]. Hence, gamma radiation can be a proxy for the average soil water content in the upper root zone [227]. However, the distribution of potassium and thorium in the soil is highly variable, and the signal attenuation depends on soil chemistry. The corresponding variability dominates over the relatively subtle dependence on soil moisture [322]. Therefore, spatial estimation of soil texture is the main application of gamma-ray surveys [323][324][325], while the estimation of soil or snow water relies on known reference data (i.e., background radiation) of the study region [326,327].
Cosmic-ray neutrons are part of the omnipresent background radiation on Earth. In contrast to electro-magnetic signals, neutrons do not interact with the electric fields of atoms, allowing them to penetrate deeply into materials [328]. In the soil, neutrons collide with atomic nuclei and reflect back to the atmosphere. Since neutrons are extraordinarily sensitive to the lightweight nucleus of the hydrogen atom, the reflected (or albedo) component of neutrons above ground depends inversely on the soil water content [329]. Consequently, neutrons are also sensitive to other sources of hydrogen, such as biomass and snow [330,331]. However, the target geometry does not play a role at all, such as terrain roughness and or leaf orientation, which can be an issue for electro-magnetic/optical remote sensing.
The neutrons penetrate the soil down to 15 cm (for wet) or 70 cm (for dry soils) and thereby sample the highly relevant "root zone" [328]. In air, neutrons can travel hundreds of meters before detection and thereby act as a proxy for the average water content within 10-20 hectares. This so-called footprint area increases with increasing terrestrial altitude and also with the detector height above ground. Due to their random-walk nature, neutrons enter the detector mostly isotropically, where the collisions with the detector gas induce countable electrical pulses [332]. Due to the isotropic nature and the fact that neutrons are almost insensitive to most materials other than hydrogen, the detector can be mounted effortlessly on or in a vehicle irrespective of configuration, orientation, viewing angle, or window.
In the last couple of years, soil moisture measurements with cosmic-ray neutrons have been conducted using stationary sensors or car-borne sensors ("rovers"), which are, however, limited to accessible terrain [240,333,334]. Very recent developments by Schrön [335] pioneered the application of airborne neutron sensing. First campaigns made use of a gyrocopter and in-situ data in areas of various land use types including agricultural fields, urban areas, forests, flood plains, and lakes (Figure 9b). The study indicated that neutrons are sensitive to soil water variability in heights of up to 200 m above ground. Both gamma and neutron methods rely on a high signal-to-noise ratio, which increases with detector volume and decreases with height above ground. Hence, ongoing developments are aiming for airborne technologies with high payload and low flying altitude. The proof of concept indicated a high potential of airborne neutron sensing, which could become a valuable addition -or even an alternative -to conventional remote-sensing methods. Moreover, cosmic-ray neutron data can also be used to ground-truth remote-sensing products [336,337], or in synergy with airborne PolSAR to correct the cosmic-ray soil moisture product for the influence of biomass [240]. developments are aiming for airborne technologies with high payload and low flying altitude. The proof of concept indicated a high potential of airborne neutron sensing, which could become a valuable addition -or even an alternative -to conventional remote-sensing methods. Moreover, cosmic-ray neutron data can also be used to ground-truth remote-sensing products [336,337], or in synergy with airborne PolSAR to correct the cosmic-ray soil moisture product for the influence of biomass [240].
The RS-aided derivation of geotraits of soil moisture is shown in Table 5. Figure 10 shows the enormous number of current and future space-based RS missions and satellites for monitoring soil moisture with information about the mission status, according to the CEOS database [205]. * SMAP RADAR stopped operation on 7 July 2015. Figure 10. Current and future RS missions instruments for monitoring soil moisture with information about the mission status, extracted from the CEOS database [205].
A selection of RS-aided data products for deriving soil moisture by RS is shown in Table 6.  Figure 10. Current and future RS missions instruments for monitoring soil moisture with information about the mission status, extracted from the CEOS database [205].
A selection of RS-aided data products for deriving soil moisture by RS is shown in Table 6. Land surface temperature (LST) is one of the most important state variables representing the coupled interaction of the surface energy and water balance and represents a valuable source of information for ecological and hydrological modeling from the local to the global scale [367][368][369]. The knowledge of LST provides crucial understanding of spatio-temporal variations of the surface equilibrium state [370] and is helpful in exploring and modelling plant-environment interactions [371]. LST is highly influenced by the radiative, thermal, and hydraulic properties of the soil-plant-atmosphere system and has therefore been recognized as one of the high-priority variables of the International Geosphere and Biosphere Program (IGBP) [372].
LST is used in various research contexts such as urban ecology like monitoring plant and human health during heat waves [373], for the description of the hydrological cycle, in climate research, or in studies of vegetation dynamics [374,375]. Furthermore, LST is often used to estimate evapotranspiration [369,[376][377][378], which is a variable that is highly controlled by atmospheric conditions, but also by stomata conductance, the plant available, the soil moisture, and processes of the surface-subsurface interactions [379]. In this sense, the monitoring of LST with high spatial and temporal resolution can provide valuable information about the water and energy exchange between the soil-plant-atmosphere continuum as well as related photosynthetic activities of the vegetation. Differences in the spatio-temporal behavior of LST can therefore be related to different plant species distributions, to differences related to the local energy, water, or nutrient conditions [380], and can even be used to improve the classification of soil texture data [181]. LST is also strongly influenced by the patterns and heterogeneity of vegetation and land surface characteristics such as soil, topography, and vegetation [381]. Therefore, the recording of surface characteristics and their heterogeneity by using RS is important for being able to adequately describe, explain, and predict LST distribution.
While LST is easily measured by thermometers at the point scale, RS thermal infrared data (TIR) are needed to derive LST routinely at high temporal and spatial resolutions over large spatial extents. However, the derivation of LST from TIR data is a difficult task for the following reasons: The radiance measurements on board the satellites not only depend on LST, but also surface emissivity and atmospheric conditions [382]. Therefore, besides cloud detection and radiometric calibration, corrections for emissivity and atmospheric effects have to be carried out. A large number of studies have addressed these issues and many of them are described in the review by Li et al. [370].
To quantify the landscape surface energy balance, there are various in-situ measurements of surface fluxes at the canopy level. Such in-situ measurements are very valuable, but they are only representative for small areas. It is therefore difficult and costly to investigate the detailed spatial pattern of energy fluxes over entire areas. TIR data can therefore be used to derive the LST of different surfaces. Hitherto, a range of airborne and satellite sensors were developed to record TIR image data i.e., Landsat TM/ETM+, MODIS, ASTER, and new satellites such as the HyspIRI that are under development Given a large number of influences on LST, airborne platforms [21] and UAV [383,384] are in use for the retrieval of LST. RS platforms and sensors currently providing TIR data differ in the spatial, spectral, and temporal resolution of LST data and are the only way to measure LST from the local to the global scale with high spatial and temporal resolution (see also Figure 11 and Table 7 The RS-aided derivation of traits of LST and related traits is shown in Table 7. Figure 12 shows the enormous number of current and future space-based RS missions and satellites for monitoring LST with information about the mission status, according to the CEOS database [205].    1 RGB/TIR 7.5-13 µm 1-5 cm [377] Sensor is used on the RS platform: UAV 1 -unmanned aerial vehicles (UAV) ; airborne 2 -airborne RS platform ; spaceborne 3 -spaceborne RS platform.
The RS-aided derivation of traits of LST and related traits is shown in Table 7. Figure 12 shows the enormous number of current and future space-based RS missions and satellites for monitoring LST with information about the mission status, according to the CEOS database [205]. Remote Sens. 2019, 11, x FOR PEER REVIEW 31 of 53 Figure 12. Current and future spaceborne RS missions and satellites for monitoring land surface temperature with information about the mission status, information by the CEOS database [205].
A selection of RS-aided data products for deriving LST from RS is shown in Table 8.

Conclusions and Further Requirements in Monitoring Geodiversity
In order to understand the complexity, processes, disturbances, and resilience of biodiversity, it is imperative to gain a deep understanding of the status, stress-induced changes, disturbances, and resource limitations for geodiversity and traits as well as their interactions and feedbacks with aboveand below-ground biodiversity.
Geodiversity and its five essential characteristics, a novel concept for the first time defined in this paper, and the definitions of traits and trait variations were introduced. Geodiversity and its A selection of RS-aided data products for deriving LST from RS is shown in Table 8.

Conclusions and Further Requirements in Monitoring Geodiversity
In order to understand the complexity, processes, disturbances, and resilience of biodiversity, it is imperative to gain a deep understanding of the status, stress-induced changes, disturbances, and resource limitations for geodiversity and traits as well as their interactions and feedbacks with above-and below-ground biodiversity.
Geodiversity and its five essential characteristics, a novel concept for the first time defined in this paper, and the definitions of traits and trait variations were introduced. Geodiversity and its traits (geotraits) can be recorded by in-situ and RS-techniques. In-situ techniques are accurate, largely point-based, but are more time-consuming and can only be repeated with considerable personnel and financial means. RS techniques on the other hand are a cost-effective alternative that are becoming increasingly more accessible, comprehensive, and repeatable, while providing the opportunity for a standardized recording of continuous geodiversity and trait variables. Geo-traits exist on all spatio-temporal scales and can thus be monitored by RS sensors on different platforms. This paper presents the state-of-the-art in monitoring geodiversity and its traits using air-and spaceborne RS of soil characteristics including soil moisture. RADAR, LiDAR, thermal sensors, multispectral, hyperspectral, and microwave RS technologies that record soil characteristics including soil moisture are presented. Furthermore, the paper discusses future satellites and existing data products that are suitable for monitoring geodiversity and its traits.
As a physical-based system, RS can monitor geodiversity and its traits (depending on the RS characteristics and composition and configuration of traits [1,3]) by using the spectral traits (ST) as well as the spectral traits variations (STV) approach. Consequently, the RS approach is then referred to as the remote sensing-spectral traits/spectral trait variations-concept-RS-ST/STV-C.
Unlike in-situ techniques, RS approaches can only record specific geo-traits/trait combinations and trait variations due to the different RS characteristics (spectral, radiometric, geometric, directional, and temporal). This paper provides an overview of those traits and trait variations that can be recorded by air-and spaceborne RS techniques (see Figure 13).
Remote Sens. 2019, 11, x FOR PEER REVIEW 32 of 53 traits (geotraits) can be recorded by in-situ and RS-techniques. In-situ techniques are accurate, largely point-based, but are more time-consuming and can only be repeated with considerable personnel and financial means. RS techniques on the other hand are a cost-effective alternative that are becoming increasingly more accessible, comprehensive, and repeatable, while providing the opportunity for a standardized recording of continuous geodiversity and trait variables. Geo-traits exist on all spatio-temporal scales and can thus be monitored by RS sensors on different platforms. This paper presents the state-of-the-art in monitoring geodiversity and its traits using air-and spaceborne RS of soil characteristics including soil moisture. RADAR, LiDAR, thermal sensors, multispectral, hyperspectral, and microwave RS technologies that record soil characteristics including soil moisture are presented. Furthermore, the paper discusses future satellites and existing data products that are suitable for monitoring geodiversity and its traits.
As a physical-based system, RS can monitor geodiversity and its traits (depending on the RS characteristics and composition and configuration of traits [1,3]) by using the spectral traits (ST) as well as the spectral traits variations (STV) approach. Consequently, the RS approach is then referred to as the remote sensing-spectral traits/spectral trait variations-concept-RS-ST/STV-C.
Unlike in-situ techniques, RS approaches can only record specific geo-traits/trait combinations and trait variations due to the different RS characteristics (spectral, radiometric, geometric, directional, and temporal). This paper provides an overview of those traits and trait variations that can be recorded by air-and spaceborne RS techniques (see Figure 13). There are limitations to monitoring soil characteristics and soil moisture, because they are influenced by the growth of vegetation. In the paper we illustrate how indirect techniques can be used on vegetation as a sensor, proxy, and indicator to monitor the status, stress, or resource limitations of soil characteristics and soil moisture.
No single monitoring technique, RS sensor, RS-approach, sensor platform, scale, or model approach is sufficient on its own to monitor and model the complexity of biodiversity, the abiotic There are limitations to monitoring soil characteristics and soil moisture, because they are influenced by the growth of vegetation. In the paper we illustrate how indirect techniques can be used on vegetation as a sensor, proxy, and indicator to monitor the status, stress, or resource limitations of soil characteristics and soil moisture.
No single monitoring technique, RS sensor, RS-approach, sensor platform, scale, or model approach is sufficient on its own to monitor and model the complexity of biodiversity, the abiotic systems, and the interactions between abiotic and biotic processes and functions in order to assess the