Varieties Matter When Planning for Poverty Alleviation and Ecological Restoration in Karst Fruit Cultivation Areas

: Landscapes are increasingly being managed to meet multiple objectives and balance social, economic, and environmental goals. The Honghe Hani and Yi Autonomous Prefecture in the Yunnan faulted Basin represent examples of areas with a range of concerns, including balancing farming with poverty alleviation, economic development, and ecological protection. Here, we set out four future land-use scenarios to quantify and map the values of key evidence-based, policy-relevant ecosystem services in three typical counties in this area. We found that implementing existing fruit-growing plans may increase the expected total value of ecosystem services due to increased acreage, but it does not actually increase the total value of ecosystem services per unit area. For example, total ecosystem service provision in Jianshui County decreased by 3% by 2023 under the current trend after the expansion of the planting area was subtracted. However, planting fruit trees with better ecological beneﬁts in areas of greater degradation risk, can increase the delivery of ecosystem services by 10–20%. Thus, variety coordination based on detailed study of the geographical contexts and ecological-economic performance of different types of fruit cultivation can produce ecological-economic improvements beyond simple expansion and better inform county-scale policy, planning, and management interventions.


Introduction
The faulted Yunnan Basin is a major rocky desertified area in China formed by violent unbalanced crustal movements during the Tertiary period [1,2]. The annual erosion modulus of soil can reach 98,100 t/km 2 ·a, which is much higher than the average 500 t/km 2 ·a for much of China [3,4]. These unique geographical conditions lead to a simple community structure that is vulnerable to degradation under human disturbance, which in turn leads to excessive population growth and economic poverty [5]. The population density of most karst counties in Yunnan Province is much higher than in other provinces, generally 120-250 people/km 2 [6]. By 2017, there were 2.79 million poor people in Yunnan, accounting for 9.1% of China's poorest population. These areas often consist of undeveloped barren hillsides suitable for fruit cultivation. In addition, fruit trees provide ecological functions, including soil and water conservation, which can help promote the development of ecological tourism, leisure tourism, and other industries [7,8].
Given these opportunities, China's central and local governments issued a series of documents to support the development of fruit farming and eliminate rural poverty; these include "some opinions on giving priority to the development of agriculture and rural areas and doing a good job in the work of agriculture, rural areas, and farmers" and the "targeted poverty alleviation plan of Yunnan Province's Characteristic Industries" (http://www.gov.cn; http://www.yn.gov.cn/, accessed on 30 September 2020). Between 2008 and 2018, the annual average fruit planting area and output of Yunnan displayed rapid and stable growth at 7.6% and 9.9%, respectively (http://data.stats.gov.cn, accessed on 30 September 2020), with "one main producing area and four sub-areas" becoming the main production area along with Honghe Prefecture [9]. The per capita net income of rural residents in karst areas also increased by 79.9%, which was higher than the national mean of 54.4% for the same period [10].
Numerous studies have been conducted on the overall impacts of industrial poverty alleviation and various ecological projects (e.g., afforestation and grass planting; forest and grassland protection) [11][12][13]. However, those efforts focused mainly on long-term trends or macro-scale processes using remote sensing images and models, while less attention has been paid to specific modes of planting at the town scale [12][13][14][15]. Furthermore, existing studies have demonstrated that different varieties of fruit have different economic and ecological values [16][17][18], which makes it possible to improve the total output of regional ecosystem services through the coordinated cultivation of multiple varieties of fruit. However, to the best of our knowledge, there have not been many studies on this. To address this knowledge gap, this study used scenario-based simulation and a geographical information system (GIS) framework to explore changes in ecosystem service delivery associated with the development of fruit planting in Yunnan Province, and the transformational effects of different production modes at the county scale. In doing so, we explored the impacts of the coordinated planting of fruit varieties on regional-scale economic and ecological benefits.

Research Area and Fruit Production Modes
Honghe Hani and Yi Autonomous Prefecture, Yunnan Province, China, is a typical faulted karst basin where 17.58% of the population (587,000 people) live below the poverty line and 45.04% (2345.08 km 2 ) of the country is undergoing desertification [19]. In the last 20 years, the fruit-growing industry has begun to develop in this area with the support of government poverty alleviation policies and the implementation of a three-phase desertification control project.
This study focused on three typical demonstration areas subject to desertification in Honghe Hani and Yi Autonomous Prefecture: Jianshui County, Luxi County, and Mengzi City ( Figure 1). Based on the Statistical Yearbook (2015-2018) and a 2016 preliminary field survey, two typical planting modes were selected for each site (Table A2). Orange and pomegranate cultivation were selected for Jianshui County, which has an elevation ranging from 230-2515 m, average annual temperature of 19.4 • C, average annual rainfall of 1045.5 mm, and an annual average of 2047.9 h of sunshine. Pear and peach cultivation were selected for Luxi County, which has an elevation ranging from 820-2459 m, average annual temperature of 16.0 • C, average annual rainfall of 1028.1 mm, and an annual average of 1951.1 h of sunshine. Finally, apple and marigold cultivation were selected for Mengzi City, which has an elevation ranging from 397-2494 m, average annual temperature of 19.0 • C, annual average rainfall of 1377.1 mm, and an annual average of 2228.3 h of sunshine (Table A1). To obtain the data required, a face-to-face questionnaire survey was conducted with farmers from July-September in 2017 and 2018, and field samples were collected in 2017 and tested in 2018 [20].

Geospatial Data
Slope data was calculated from a 30 m × 30 m resolution digital elevation model (DEM) obtained from the Geoscientific Data Cloud website (http://www.gscloud.cn, accessed on 30 September 2020). Land-use data for 2015, at the same resolution, were derived from Landsat-8 images [21]. Monthly temperature and rainfall data were obtained from the National Meteorological Data Center and interpolated from 2012-2016 records. To evaluate the maximum area suitable for planting in each mode, climatic and geographical factors were classified, as follows. The "Technical Regulations for Investigation of Land Use Status" and "Technical Regulations for Determining the Gradation of Cultivated Land Using DEM (for trial implementation)" guidelines were used to divide slope gradients into the following four grades: <6 • , 6-15 • , 15-25 • , and >25 • . The growth characteristics of the modes of cultivation studied were used to divide the annual average rainfall into the following categories: <750 mm, 750-1000 mm, 1000-1250 mm, and >1250 mm. The annual average temperature was divided into the following categories: <15 • C, 15-20 • C, and >20 • C. We aimed to focus on the economic and ecological benefits brought about by changes in planting modes; hence, land-use data were reclassified into eight categories including rice fields, other farmland, bare farmland, and forests. All GIS layers were generated as 30 m × 30 m raster images.

Ecosystem Service Data
Previous studies have found that climate factors and altitude can significantly change the ripening time of fruits planted in Yunnan, but have little effect on other characteristics [22]. Therefore, we used pattern-based ecosystem service data from previous studies to represent the entire study area (Table A4). Briefly, we categorized the provision of fruits and flowers into the following ecosystem services as per the Millennium Ecosystem Assessment framework and evaluation criteria for karst regions: general provisioning services, reducing topsoil loss, maintaining soil fertility, and photosynthetic carbon fixation as a regulating service, photosynthetic oxygen release and nutrient retention as supporting services, and education as a cultural service [20] (Tables A3 and A4). In general, pomegranate cultivation should be prioritized where economic benefits are sought and apple cultivation should be promoted where ecological benefits are needed. To eliminate the impact of the differences in the composition of and calculation methods for different ecosystem services, relative ecosystem services were calculated based on the following formulas.
In Equation (1), Relative ES is the value of one specific type of ecosystem service (ES) after standardization, ES i represents the actual total amount of ecosystem service i (e.g., maintaining soil fertility) in one quadrat, ES i−max is the maximum value of the actual total amount of ecosystem service i in all land-use patterns quadrats, and ES i−min denotes the minimum value under the same condition.
To equalize the weights of the contributions of the four types of ecosystem services to total ecosystem services (TES), the latter was calculated using Equation (2). The modified index eliminated the impact of differences in the number of types of service in each category.
In Equation (2), Relative ES p is the standardized value of provisioning services.

Scenario Setting
Planting area data for each planting mode in 2016, 2017, and 2018 were obtained from the three county governments. The planting areas for each mode in 2023 were used as the baseline scenario (i.e., the result of development according to current trends). In the ecological priority scenario, the planting mode with the greatest ecological benefits is promoted preferentially; in the economic priority scenario, the planting mode with the greatest economic benefits is promoted. A land classification map of the three counties was obtained at the same time to help identify areas suitable for planting and those at high risk of degradation; this was done by superposing topography, soil erosion, land-use, and climate datasets ( Figure 2, Tables A5-A7). The scenario analysis was conducted based on the transformation of farmland into various planting modes, as per the relevant national land-use policies and regulations (e.g., Law of the People's Republic of China on Land Administration; Forest Law of the People's Republic of China), and is summarized in Table 1.

Scenarios Description
Baseline scenario Existing policies and regulations are not adjusted. No major technological advances occur for any of the planting modes.

Economic priority scenario
Planting structure strongly promotes higher economic output. All new planting areas and suitable areas obtained from reducing areas planted in other modes are planted with crops with high economic service value.

Compromise scenario
Planting structure is transformed to high modes of economic output while limiting ecological damage. Crops with high economic value are planted in areas where the risk of erosion is low. Areas where the risk of erosion is high are planted with crops of high ecological value.

Ecological priority scenario
Planting structure strongly promotes high ecological gains. All new planting areas and suitable areas obtained from reducing areas planted in other modes are planted with crops with high ecological service value.
To better measure the changes in ecosystem services resulting from land-use changes, the equal-area change rate (EACR) was calculated based on Equation To better measure the changes in ecosystem services resulting from land-use changes, the equal-area change rate (EACR) was calculated based on Equation

Scenarios Description
Baseline scenario Existing policies and regulations are not adjusted. No major technological advances occur for any of the planting modes.

Economic priority scenario
Planting structure strongly promotes higher economic output. All new planting areas and suitable areas obtained from reducing areas planted in other modes are planted with crops with high economic service value.

Compromise scenario
Planting structure is transformed to high modes of economic output while limiting ecological damage. Crops with high economic value are planted in areas where the risk of erosion is low. Areas where the risk of erosion is high are planted with crops of high ecological value.

Ecological priority scenario
Planting structure strongly promotes high ecological gains. All new planting areas and suitable areas obtained from reducing areas planted in other modes are planted with crops with high ecological service value.

Temperature, Rainfall, Slope, Soil Erosion, and Land-Use Patterns
Overall, 39.12% of the study area had a slope gradient between 6 and 15°, 24.28% had a gradient <6°, and 24.64% ranged from 15° and 25°. Steep areas (>25°) accounted for 11.96% of the total area of the three counties, although there was some variability between them. The terrain in Luxi County was relatively flat, with 6° and 6-15° slopes accounting for 74.73% of its land area and slopes >25° covering only 7.92% of the county.
Most soil erosion across the entire study area was classified as 'slight' (59.11%) or 'mild' (29.05%). The area of 'moderate' erosion accounted for 10.99% of the total, whereas only 0.58% of the land was classified as experiencing 'intense' erosion. In Luxi County, areas of 'slight' and 'mild' erosion accounted for 89.94% of the land area, with the remaining areas experiencing 'moderate' erosion (11.47%). In Jianshui County, erosion levels in 98.32% of the land was classified as either 'slight' (52.17%), 'mild' (34.83%), or 'moderate' (11.32%), with 1.68%, mainly in the central area, undergoing 'intense' erosion. In Mengzi City, 73.75% of the land was classified as experiencing 'mild' erosion; combined with areas of 'slight' and 'moderate' erosion, this accounted for 99.97% of the total. Areas of 'moderate' erosion were concentrated mainly in the central-eastern area of the city (Figure 3d).
The land-use types in the study area were mainly grassland (48.79%) and shrubland (35.77%); forests only accounted for 1.70% ( Figure 4). All farmland types accounted for 7.49% of the total area. The proportion of grassland in Luxi County was greater than the overall average at 55.72% and were mainly distributed in the east and south. Shrubland accounted for 23.66%; some scattered forests accounted for 1.69%; and farmland, which was concentrated mainly in the northwestern and central regions, accounted for 13.57%.
Grassland (46.97%), shrubland (39.57%), and forests (16.74%) accounted for the majority of the land area in Jianshui County. Farmland in this area was mainly concentrated in its northwest and central regions and accounted for 5.69% of its land area; this was lower than the overall average for the study area. Similarly, grassland, shrubland, and forests accounted for 46.75%, 38.46%, and 1.75% of the area in Mengzi City. Farmland accounted for 5.78% of its area and was concentrated mainly in its northwestern corner (Figure 4). counting for 35.91%, 15-25° slopes accounting for 26.95%, and > 25° slopes accounting 12.57%, the last mainly in the northwest (Figure 3c).
Most soil erosion across the entire study area was classified as 'slight' (59.11%) or 'mild' (29.05%). The area of 'moderate' erosion accounted for 10.99% of the total, whereas only 0.58% of the land was classified as experiencing 'intense' erosion. In Luxi County, areas of 'slight' and 'mild' erosion accounted for 89.94% of the land area, with the remaining areas experiencing 'moderate' erosion (11.47%). In Jianshui County, erosion levels in 98.32% of the land was classified as either 'slight' (52.17%), 'mild' (34.83%), or 'moderate' (11.32%), with 1.68%, mainly in the central area, undergoing 'intense' erosion. In Mengzi City, 73.75% of the land was classified as experiencing 'mild' erosion; combined with areas of 'slight' and 'moderate' erosion, this accounted for 99.97% of the total. Areas of 'moderate' erosion were concentrated mainly in the central-eastern area of the city (Figure 3d).  A comparison of the rainfall and temperature classifications with the growth characteristics of each cultivation mode demonstrates that each of the counties could support the planting of all six fruit species considered; this is consistent with the field survey. Overall, land in Luxi County was divided into 14 categories (Table A5), of which land with a slope < 6 • , 'slight' erosion, a mean annual temperature of 15-20 • C, and mean annual rainfall of 1000-1250 mm accounted for the largest area. The same was found for Mengzi City, although this was divided into 15 different categories (Table A7). Land in Jianshui County was more variable, and was thus classified into 37 different categories (Table A6). Here, areas with a slope < 6 • , 'slight' erosion, mean annual temperature of 15-20 • C, and mean annual rainfall of 750-1000 mm accounted for the largest area. Finally, land at high  (Tables A5-A7). accounted for 23.66%; some scattered forests accounted for 1.69%; and farmland, which was concentrated mainly in the northwestern and central regions, accounted for 13.57%. Grassland (46.97%), shrubland (39.57%), and forests (16.74%) accounted for the majority of the land area in Jianshui County. Farmland in this area was mainly concentrated in its northwest and central regions and accounted for 5.69% of its land area; this was lower than the overall average for the study area. Similarly, grassland, shrubland, and forests accounted for 46.75%, 38.46%, and 1.75% of the area in Mengzi City. Farmland accounted for 5.78% of its area and was concentrated mainly in its northwestern corner ( Figure  4). A comparison of the rainfall and temperature classifications with the growth characteristics of each cultivation mode demonstrates that each of the counties could support the planting of all six fruit species considered; this is consistent with the field survey. Overall, land in Luxi County was divided into 14 categories (Table A5), of which land with a slope < 6°, 'slight' erosion, a mean annual temperature of 15-20 °C, and mean annual rainfall of 1000-1250 mm accounted for the largest area. The same was found for

Change in Planting Area under Different Scenarios
According to the statistics for 2016-2018 obtained from the Luxi County government, the marigold planting area decreased by 31% each year; based on this trend, the estimated planting area would be 87.03 ha by 2023. The apple planting area, calculated using the same principles but with a positive trend, is expected to reach 5094.02 ha by 2023. Across all crop types considered, an additional 7374.98 ha of land is expected to be newly planted by 2023 compared to 2018 (Table 2). In terms of the different scenarios, pomegranate cultivation was promoted in the economic priority scenario, increasing in planting area by 7854.62 ha. Under the compromise scenario, apples would be planted on 2300.58 ha of land at high risk of erosion and pomegranates would be planted in other areas. In the ecological priority scenario, all new growing areas would be planted with apple trees.
In Jianshu County, the pear planting area decreased between 2016 and 2018 at an average rate of 4% each year, with a projected 2023 planting area of 523.44 ha ( Table 3). The planting areas for oranges, peaches, and pomegranates increased, with the orange cultivation area projected to increase the most to reach 10,764.80 ha by 2023. In general, compared to 2018, a total of 6736.38 ha is projected to be newly planted by 2023 (Table 3). Under the different scenarios, pomegranates account for 100% of this increase, whereas pear cultivation would be reduced by 128.89 ha. In the compromise scenario, 4698.9 ha of land at high risk of erosion would be planted with apples (for ecological gain) while the remaining areas are planted with pomegranates (for economic gain). In the ecological priority scenario, apple cultivation is prioritized in 6685.27 ha (Table 3). In Mengzi City, the marigold and apple planting areas greatly decreased between 2016 and 2018 at average annual rates of 17% and 13%, respectively. Based on these trends, the marigold and apple planting areas are projected to be 402.26 ha and 1876.52 ha, respectively, by 2023. The pear planting area also slightly decreased during this period, whereas the peach planting area increased the most and is projected to reach 12,688.14 ha by 2023. Across all crop types, an area of 13,852.24 ha is projected to be newly planted by 2023, compared to 2018. Pomegranate planting was included in all areas in Mengzi City under the economic priority scenario, with reductions in marigold, pear, and apple planting areas, totaling 14,480.94 ha. In the compromise scenario, 489.33 ha of land at high risk of erosion is planted with apples and the remaining 13,911.65 ha is planted with pomegranates. In the ecological priority scenario, the areas made available from reductions in marigold, pear, and apple cultivation are replanted with apple trees (Table 4).

Changes in Ecosystem Service Provision
According to the current development trends in Luxi County, the provisioning services provided by the six cultivation modes are expected to increase by 2023 at a rate of 4% (Figure 5a). The most significant increase is projected to be in regulating services, with an increase of 7%, followed by supporting services at 6%. Overall, TES in Luxi County is projected to increase by 6% by 2023. Under the economic priority scenario, provisioning services in Luxi County increase by 246% by 2023, compared to 2018. Supporting services and cultural services also increase by 2% and 10%, respectively. However, regulating services are projected to decrease by 7% by 2023. An overall increase of 15% in total ecosystem service provision is projected under the economic priority scenario. Under the compromise scenario, the increase in provisioning services is reduced and the EACR is 179%; however, the reduction in regulating services also decreased, with an EACR of −2%, whereas there was a doubling of growth in supporting services. Overall, there was an approximate 14% increase in the growth rate of TES provision, which is in line with the economic priority scenario. In the ecological priority scenario, the EACR of the provisioning service is 17%. However, the growth rate of regulating services and supporting services increased, with EACRs of 11% and 9%, respectively. Overall, total ecosystem service provision increased by 11% under this scenario (Table 5). Jianshui County, like Luxi County, is projected to experience an increase in the total provision of ecosystem services by 2023, based on current trends (Figure 5b). However, after excluding the change caused by the increase in the planting area, several ecosystem services are projected to decrease, with supporting services displaying the greatest decline of −15%. The EACR of supporting services is 4%, and the declines in regulating and cultural services are both 1%; the overall decline across all service types is 3%. In the economic priority scenario, the growth rate of provisioning services is significant, with an EACR of 169%. Supporting services and cultural services also display significant growth at 11% and 13%, respectively. In contrast, regulating services display a 7% decline, resulting in an overall decrease of 23%. Under the compromise scenario, all ecosystem services demonstrated significant increases; the rate of increase of provisioning services was the highest, with an EACR of 47%. Supporting services also increased significantly, by 19%, and the growth rates of both regulating and cultural services was 13%. Overall, under the compromise scenario, total ecosystem service delivery is projected to increase by 18% by 2023, which is much higher than the 3% reduction expected under the current trend.
Finally, under the ecological priority scenario, regulating and supporting services increase to 23% while provisioning services are reduced by 10%, although this is lower than the 15% decline expected under current trends. Overall, total ecosystem service delivery increases by 16% under this scenario (Table 5).  Jianshui County, like Luxi County, is projected to experience an increase in the total provision of ecosystem services by 2023, based on current trends (Figure 5b). However, after excluding the change caused by the increase in the planting area, several ecosystem services are projected to decrease, with supporting services displaying the greatest de- According to the trend for 2016-2018, various ecosystem services in Mengzi City are also projected to increase by 2023 (Figure 5c). Excluding the area factor, provisioning services displayed a notable downward trend at a 28% reduction; cultural services demonstrated a decline of 8%. At the same time, regulating services and supporting services displayed significant growth rates of 11% and 8%, respectively. As a result, a slight decline of 3% is projected overall. In the economic priority scenario, the growth of provisioning services is significant, with an EACR of 58%. There are also slight increases, of 2% and 6%, respectively, in supporting and cultural services. However, regulating services decrease by 10%, resulting in overall growth of 10%. In the compromise scenario, the increase in provisioning services decreases slightly to 55%. The decline in regulating services is also slightly reduced at 8%. Overall, there are no notable differences under the economic priority scenario, which has an EACR of 10%. In the ecological priority scenario, there is a significant decline in provisioning services (by 32%), whereas regulating and supporting services increase by 21% and 12%, respectively, yielding an overall increase of 4% under this scenario (Table 5).
Considering all areas together, all ecosystem service types are expected to increase by 2023 (Figure 5d), with an overall increase of 78% accounting for area effects. Provisioning services increase the least at 50%, while regulating and supporting services increase by 87% and 84%, respectively. In the economic priority scenario, provisioning services have a significant increase at 287%, supporting and cultural services increase by 82% and 93%, respectively, and regulating services increase by 61%. This equates to an overall increase in ecosystem services provision of 103%. In the compromise scenario, the increase in provisioning is slightly dampened at 226%, and regulating and supporting services display rates of change of 72% and 86%, respectively. Overall, ecosystem service provision is slightly reduced relative to the economic priority scenario with an EACR of 100%. In the ecological priority scenario, the change in provisioning services is consistent with the current trend at 50%; however, growth in regulating and supporting services is significantly enhanced at 105% and 98%, respectively. This equates to an overall increase in ecosystem services provision of 93% for this scenario (Table 5).

Discussion
The complex relationships between land-use and ecosystem services make it difficult for stakeholders to obtain and interpret key information on ecosystem service provision in different future land-use planning contexts [23]. Many studies have noted that land-use largely determines the transformation of regional ecosystem services, while the increase or decrease of ecosystem services will simultaneously cause changes in land-use modes [24,25]. In reality, the mutual transformation of different land-use modes is often accompanied by the expansion of specific land-use types. Unlike previous studies that have focused on multi-category land-use types (e.g., grassland, forests, and farming areas) [26,27], we focused on the rapid spread of planting in rural areas aimed at generating high economic benefits.
Our results demonstrate that, based on current planting trends in Luxi County, Jianshui County, and Mengzi City, areas of fruit cultivation and associated profits will increase in all cases, in line with national policy on poverty alleviation and desertification control. However, this hindered the study of regional changes in ecosystem service provision caused by land-use changes. The expansion in the planting area meant that total ecosystem service provisions for Luxi County, Jianshui County, and Mengzi City were projected to increase significantly; however, accounting for the increase in the growing area, a 3% decline is projected for Jianshui County and Mengzi City (Table 5). For example, all types of services are expected to decline in Jianshui County, while provisioning services in Mengzi City could decrease by as much as 28%, contrary to expectations. Across the entire study area, TES provision will increase by 78% by 2023, based on current development trends, indicating that complementary and diverse land-use modes can limit ongoing decline; in other words, there are tradeoffs in spatial ecosystem services [28]. In addition, fine-scale land-use management interventions can be used to maximize ecosystem service gains [29].
In our forecasts, the economic priority scenario met expectations to provide the largest increase in provisioning services while also achieving the highest overall TES. In contrast, provisioning services declined in Jianshui County and Mengzi City under the ecological priority scenario. Furthermore, provisioning services increased by 17% under the ecological priority scenario in Luxi County, where there was an overall increase of 11% in all services; this was 4% lower than in the economic priority scenario. Declines in supporting, regulating, and cultural services in Luxi County are dampened, consistent with two key conclusions in previous research. The first of these was that there are tradeoffs between provisioning services and other types of services [20,30]. The second is that a reasonable spatial arrangement of land uses can ease these tradeoffs, and that outcomes can be improved by employing sophisticated land management methods at smaller scales, depending on the context [31,32].
According to the ecological priority scenario, the provision of supporting services increased by 50% across the entire research area; this is consistent with current trends. Ecosystem disservices (e.g., fertilizer, water, and pesticide consumption) can also change simultaneously with adjustments in the variety of crops planted [33,34]. Our results indicate that irrigation costs will increase to CNY 9.13 million in 2023 and CNY 14.46 million under the economic priority scenario (Tables A9 and A10). On the contrary, the costs of irrigation, pesticides, and water consumption are >10% lower under the ecological priority scenario than under current trends. In addition, the portion of land planted with apple trees will be greatly increased, which will assist in the preservation of soil nutrients (Table A8). This demonstrates that the current development pattern is not considered optimal. Overall, the planting trend for development in Jianshui County should be adjusted, as the compromise scenario offers an effective method of reversing the current declines in ecosystem service provision while also stimulating growth. Luxi County and Mengzi City should plan development according to the compromise scenario; however, the provision of regulation services needs to be further improved in these areas.
The inability to collect sufficient information to build accurate fruit-planting maps, due to a lack of satellite data and local-scale species-specific information, was a major limitation of our study [35,36]. It is difficult to accurately evaluate long-term trends in ecosystem service provision in the fruit planting industry, as it is highly market dependent. Many external factors (e.g., changes in varieties and international market influences) affect the provisioning services associated with fruit cultivation, whereas regulation services and supporting services (including carbon fixation and oxygen release) are comparatively stable [37]. Equally, however, the ecosystem services and disservices provided by orchards that are longstanding elements of a landscape will interact with the surrounding landscape. Our study measured this only in terms of crop water requirements and irrigation cost to reflect water consumption by orchards; however, this does not consider the water exchange with the surrounding area [38]. Therefore, improved long-term statistics are needed for specific fruit varieties to more accurately predict their benefits. In addition, fruit trees-a crucial element in alleviating poverty among fruit growers-play a non-substitutable role in leisure agriculture and will provide farmers with more benefits via their ornamental value. For example, a flowering time spanning February-May and a picking period from July-October can be created by planting pear, cherry, and pomegranate orchards in adjacent areas; this would support a six-month "golden tourist season" [39]. However, tourism values are also affected by multiple interacting factors, making the benefits of developing specific fruit-farming practices difficult to determine.

Conclusions
Changes in ecosystem service provision that are the result of land-use change have received widespread attention; however, previous research has largely focused on largescale and cross-industry changes. Using a scenario-based analysis combined with a GIS framework, we found that economic income remains the primary consideration for the promotion (or self-selection by farmers) of particular fruit trees in three counties studied in China. However, by purposefully adjusting and promoting the cultivation of specific fruits, ecosystem service provision could be improved by 10-20% overall (Tables 2-5). This implies that fine-scale land-use policy, planning, and management interventions could reconcile tradeoffs in ecosystem services and support ecologically and economically coordinated development.      Note: The unit for "Irrigation", "Pesticide" and, "Fertilizer" is "CHY Million". The unit for "Water" is "Million m 3 ".