Comparison of Wood Quality of Douglas Fir and Spruce from Afforested Agricultural Land and Permanent Forest Land in the Czech Republic

This study compares the quality of wood from two distinct sites in the Czech Republic—from former afforested agricultural land and forest land. We compared the properties of Norway spruce wood (Picea abies Karst.) and Scots pine wood (Pinus sylvestris L.), the most important domestic tree species, to Douglas fir (Pseudotsuga menziesii (Mirbel) Franco), a North American tree species and a potential substitute for the domestic spruce. Wood density, modulus of elasticity (MOE), modulus of rupture (MOR) and impact bending strength were the properties tested that were used for comparing tree species. Without taking into consideration the site, the highest density values from the tested tree species were obtained for Douglas fir (0.568 g·cm−3), followed by the pine (0.508 g·cm−3) and the spruce (0.463 g·cm−3). The Douglas fir also dominated in the remaining assessed properties, whilst the influence of site was not confirmed, with the exception of MOE and MOR, and only for the Douglas fir wood, wherein higher values were obtained for forest land. In terms assessed Douglas fir properties, it exceeds the domestic softwoods and represents a possible suitable replacement for them. The site only plays a role in terms of the Douglas fir, and only for certain properties.


Introduction
As in other European countries, the afforested area is increasing steadily in the Czech Republic.It has increased from approximately 25% to 34% at present since midway through the 18th century.After medieval deforestation, large areas were reforested again for production purposes.The most extensive afforestation effort took place after the Second World War (1939)(1940)(1941)(1942)(1943)(1944)(1945), when large areas were afforested in regions with former German settlement [1], and the area of forest lands continued to increase during the remainder of the 20th century, reaching its peak in 1991 after a huge transformation of the farm sector [2].
Douglasfir is the most popular and utilized introduced tree species in Europe, including in the Czech Republic, and it has the highest production potential among the native and introduced tree species [3].It very often represents planted and cultivated species, especially in France, Germany, Italy and the United Kingdom [4].In the Czech Republic, the area is only represented by some 5800 ha, increasing by approximately 100 ha per year [5,6].Its role is particularly assumed as a substitute for the declining Norway spruce, which is endangered by climatic changes at lower altitudes [7], and this species, native on some 11-15% of the Czech territory, shows many health problems, yet is at the same time the key species of Mid-European forestry [8,9].The supply of Norway spruce timber will likely decrease in the near future [10].Afforestation of agricultural lands constitutes a serious change in soil dynamics, including soil structure, organic matter and nutrient dynamics [11][12][13][14].Compared to Norway spruce, Douglas fir shows less negative effects on the forest soil [15,16] and herb vegetation layer [17], and thus Douglas fir can represent a very convenient substitute for declining spruce, at least as a part of its non-native range in commercial forests [18].Douglas fir also appears promising under local conditions in terms of wood quality [19].
There are very few works that have evaluated the properties of wood from former agricultural land in the Czech Republic and other European countries.Most of the works deal with tree species growth, volume production or health status, and at present there is only one study from the Czech Republic dealing with spruce wood quality from afforested agricultural land [20].Within the wider region, Polish studies are available on wood properties of pine from agricultural land [21,22].In Sweden, Johansson [23] dealt with the impact of these specific sites on the quality of spruce wood.
The aim of this study is to evaluate the quality of Douglas fir wood growing on a site formerly used as agricultural land.Based on wood density, impact bending strength, bending strength and MOE, the work assesses the extent to which such habitat types are suitable for the production of quality wood and, in particular, assesses the suitability of Douglas fir for substitution of spruce wood, the original and currently the most widely used coniferous species in the Czech Republic on wooded former agricultural land.The variability of the properties in relation to the position in the stem and the influence of the density on the strength characteristics were also evaluated.

Materials
For the purpose of this study, 2 different sites in the Central Bohemia area (Czech Republic) were selected.The first site, Krymlov (49.9460656N, 14.9257086 E), represents forested agricultural land (AL).The growth is a mixture of spruce, Scots pine, white birch and Douglas fir at a site characterized by an altitude of 430 m above sea level, an annual average temperature of 7.5 • C and 600 mm of annual precipitation.The soil type is characterized as Gleyed Luvisol to Luvic Gley.Under these conditions, the tree species reached a standing volume in the following order: pine, spruce, birch and Douglas fir 352, 349, 157 and 438 m 3 •ha −1 , and Douglas fir thus clearly represented the most productive tree species from all tested species [24].
The second study plot, Amerika (50.0072625N, 14.8537100 E), represents reference growth on permanent forest land (FL), i.e., the forest continuity has never been interrupted at the site.This is a stand with Norway spruce and Douglas fir.The site is relatively acidic, characterized by an altitude 420 m above sea level, a mean annual temperature of 8.5 • C and precipitation of 550-650 mm per year.The soil is a Luvisol.The standing volume was 507 m 3 •ha −1 for spruce and 579 m 3 •ha −1 for Douglas fir [15].The age of all felled trees at both sites was around 60 years.
A total of four sample trees of Douglas fir, four samples trees of spruce and four samples trees of pine trees were taken.An equivalent number of Douglas fir and spruce samples were taken on the former afforested agricultural land for comparison with the neighbouring reference forest land.Unfortunately, it was not possible to find forest soil in the adjoining afforested agricultural land with a mixture of Douglas fir-spruce-pine; however, we considered it a shame not to take advantage of the possibility of assessing the quality of the pine wood from the agricultural land, and it was at least partially included in the evaluation.
The basic forest characteristics of the specimens, i.e., the range of heights and enumeration diameter according to tree species and site, are shown in Table 1.

Methods
Sections (120 cm in length) were taken from each sample tree, representing the basal part of the stem and 20%, 40% and 60% of the tree height, marked as vertical positions (VP) 1, 2, 3 and 4 in respective order.The sections were cut using a band saw in order to obtain the central plank.The central plank was then the starting material for the production of test specimens for the appropriate tests and assessments of the influence of the horizontal position (HP), where position 1 denotes the part closest to the centre of the stem, whilst the position with the highest number indicates the part near the bark.The length of the test specimens differed depending on the test, but the cross-section area of 20 × 20 mm (radial × tangential) was the same for all tests.
For the purpose of evaluating the investigated tree species and the impact of site, we tested the following physical and mechanical wood properties.For physical properties, wood density was evaluated on specimens at 12% moisture content after conditioning at 20 • C and 65% relative humidity, and a total of 3695 specimens measuring 20 × 20 × 30 mm were used to determine the density according to the Czech national standard [25].Density (in g•cm −3 ) was calculated according to the following formula: where ρ 12 is the density at a wood moisture content of 12%, m 12 is the mass of the test samples (g) at a wood moisture of 12%, and a 12 , b 12 , and l 12 are the dimensions of the test samples (cm) at a wood moisture of 12%.
For mechanical properties, modulus of elasticity in bending (MOE), modulus of rupture (MOR) and impact bending strength were evaluated.The tests for mechanical properties were evaluated on specimens at 12% of moisture content after conditioning at 20 • C and 65% relative humidity according to the Czech national standard [26].Specimens measuring 20 × 20 × 300 mm were used for all of the specified tests.
MOE was determined and evaluated according to the Czech national standard [27].A total of 1139 specimens were used.MOE (in MPa) was calculated according to the following formula: where ∆F is the difference between the forces at maximum and minimum load limits (N), l 0 is the distance between the supports (mm), b and h are the width and height (mm), respectively, of the test samples, and ∆y is the test sample deflection (mm) in the area of pure bending, equal to the difference between the bending values corresponding to maximum and minimum load limits.MOR was determined and evaluated according to the Czech national standard [28].A total of 1139 specimens were used.MOR (in MPa) was calculated according to the following formula: where F max is the maximum load (N), l 0 is the distance between the two supports (mm), and b and h are the respective width and height (mm) of the test samples.
Impact bending strength was determined and evaluated according to the Czech national standard [29].A total of 2005 specimens were used, and impact bending strength (in J•cm −2 ) was calculated according to the following formula: where W is the power consumed for the wood rupture (J), and b and h are the test sample dimensions (cm) in the radial and tangential directions respectively.Statistically significant differences in the assessed wood properties between tree species within one site, and differences between the same tree species depending on the site, or the influence of other factors such as vertical position and horizontal position in the stem, were assessed using ANOVA and subsequently, Tukey's multiple-range test.A significance level α = 0.05 was used as the default value.A linear regression model was used to assess the effect of density on the strength characteristics.
Due to the fact that pine was only represented on agricultural land and was missing from the reference site, the values obtained were those used only for comparison with other tree species within sites and for comparison with literature.Pine was not used to evaluate the impact of sites.

Density
The highest wood density value was exhibited by Douglas fir for both of the assessed sites, followed by the pine, while the lowest density value was obtained by the spruce for both sites.The differences between individual species within sites were statistically significant, but the difference is also apparent from the point of view of their use in the woodworking industry, especially between the spruce and the Douglas fir.According to our results, the origin of a site does not affect density (Figure 1).No statistically significant difference was proven for any of the tree species (see Table 2).

Stiffness
Similar to density, the highest MOE value was obtained for Douglas fir wood, again at both sites.
Unlike density, the second highest value was obtained by the spruce, and the lowest value by the pine.In terms of processing and utilization, pine wood is the least flexible; however, no statistically

Stiffness
Similar to density, the highest MOE value was obtained for Douglas fir wood, again at both sites.Unlike density, the second highest value was obtained by the spruce, and the lowest value by the pine.In terms of processing and utilization, pine wood is the least flexible; however, no statistically significant difference was found between spruce and pine, and Douglas fir clearly differs from both species.In the case of MOE, the sites had an impact, but only in the case of the Douglas fir (Figure 2).The Douglas fir achieves a higher MOE value on forest soil (see Table 3).

Bending Strength
The Douglas fir wood once again dominates in terms of bending strength at both sites.On the former agricultural land, pine obtained the second highest value.However, this value is not statistically significant compared to the spruce that obtained the lowest value.The influence of the site on bending strength is conclusive for Douglas fir wood, whereas for the spruce, the influence of the site was not confirmed (Figure 3) and the values are practically identical (see Table 4).
Table 4. Bending strength (in MPa) of individual tree species according to the evaluated site.

Bending Strength
The Douglas fir wood once again dominates in terms of bending strength at both sites.On the former agricultural land, pine obtained the second highest value.However, this value is not statistically significant compared to the spruce that obtained the lowest value.The influence of the site on bending strength is conclusive for Douglas fir wood, whereas for the spruce, the influence of the site was not confirmed (Figure 3) and the values are practically identical (see Table 4).

Impact Bending Strength
In terms of impact bending strength, similar to all other tested properties, Douglas fir wood obtained the highest values regardless of site.The value represents almost twice the value for spruce and pine and is statistically significant, and Douglas fir wood therefore better resists dynamic loads than domestic tree species.The values for spruce and pine differ only slightly and are essentially identical in terms of statistics (see Table 5).The impact of sites on Douglas fir or the spruce was not confirmed (Figure 4).Another factor that greatly contributes to the variability of the evaluated properties, regardless of tree species or site, is the position in the stem.As expected, the influence of the horizontal position

Impact Bending Strength
In terms of impact bending strength, similar to all other tested properties, Douglas fir wood obtained the highest values regardless of site.The value represents almost twice the value for spruce and pine and is statistically significant, and Douglas fir wood therefore better resists dynamic loads than domestic tree species.The values for spruce and pine differ only slightly and are essentially identical in terms of statistics (see Table 5).The impact of sites on Douglas fir or the spruce was not confirmed (Figure 4).Another factor that greatly contributes to the variability of the evaluated properties, regardless of tree species or site, is the position in the stem.As expected, the influence of the horizontal position was clearly demonstrated, where the lowest values are obtained in the centre of the stem, and the values increase in the direction to the bark.This is particularly striking in the Douglas fir and the pine.Figure 5 shows the course of density depending on the position in the stem according to the tree species.There is a similar layout trend for MOE, MOR and impact bending strength, while the influence of the vertical position proved to be ambiguous and it was not possible to trace any characteristic course.
Since density is often understood as a basic indicator of mechanical properties of wood, we were interested in how much the density value correlates with the strength and elastic properties of wood.Regarding the evaluated properties, the tightest correlation is between density and bending strength, as shown in Figure 6 for spruce wood; a similar trend was found for other species as well, and the tightest dependency ratio pertains to the pine (R 2 = 0.57), followed by the spruce (R 2 = 0.55) and the Douglas fir (R 2 = 0.53) (see Table 6, bold text).as shown in Figure 6 for spruce wood; a similar trend was found for other species as well, and the tightest dependency ratio pertains to the pine (R 2 = 0.57), followed by the spruce (R 2 = 0.55) and the Douglas fir (R 2 = 0.53) (see Table 6, bold text).       of forest growth (analogy of nutrient delivery to agricultural crops) on wood properties, where it is clearly demonstrated that higher nutrient intake corresponds to a decrease in density and strength of the spruce, pine or the Douglas fir [34][35][36].However, this may not be the norm, as proven by Bartoš [20] for spruce wood in the Czech Republic, and wood from such sites need not be regarded as less valuable.This is confirmed by our results.It should be noted that some authors obtained even better wood properties on former agricultural land [37].The Douglas fir is clearly justified as a potential substitute for spruce wood because the Douglas fir dominates in all rated properties.The difference is particularly obvious in the density, where the difference is not only statistically significant, but also significant in terms of the utilization of the wood, and the site does not play a role here.Similar is the striking difference in the impact bending strength, both in terms of statistics and usability of wood.The difference between the Douglas fir and spruce is not as pronounced in MOE and MOR, mainly due to the differences that occurred with the Douglas fir depending on the site; nevertheless, the Douglas fir wood still achieves higher values for these properties.
The increasing trend of density, and hence strength characteristics, in the direction from the centre of the stem to the cambium, is characteristic of conifers due to the presence of juvenile wood [38,39].This was observed for the Douglas fir [40], for spruce wood [41], for pines [42] and other conifers [43].
Density affects impact bending strength the least.In the case of the Douglas fir and the spruce, it can be stated that on forest soil the dependence of strength and elastic characteristics on density is greater.The more significant influence of density on strength characteristics was also confirmed by Jelonek [37] (R 2 = 0.53).

Conclusions
Douglas fir is a promising tree species for Czech forestry.With the growing area of forests, the issue of the quality of wood originating from afforested former farmland is the focus.The aim of this study is to evaluate the wood properties of Douglas fir originating from such sites in contrast to native spruce and to some extent, pine.Although wood originating from former farm land is often regarded as inferior, this was not confirmed in most cases of our research, and our results are as follows.
In all of the tested properties, the Douglas fir had the highest values and clearly exceeds the evaluated domestic conifers.In term of density, it exceeds the spruce by more than 100 kg•m −3 , and the value impact bending strength is also almost double.
The origin of the site plays no role at all in the case of the spruce.In the case of the Douglas fir, an impact was only observed for MOE and MOR, where higher values were obtained on forest soil.
Based on a comparison with literature, sites provide wood of adequate quality in terms of density for all tree species and impact bending strength for the Douglas fir.For the remaining properties, the values are lower than those reported in the literature.
The position in the stem proved to be another factor that significantly influences the variability of wood properties, as the values increase with increasing distance from the centre of the stem.These trends are primarily striking for the Douglas fir and pine.The influence of the vertical position in the stem was not confirmed.
The dependence of the measured mechanical properties on density is the highest for bending strength.The highest value, R 2 = 0.57, was obtained for the pine.

Figure 1 .
Figure 1.Impact of site on wood density.

Figure 1 .
Figure 1.Impact of site on wood density.

Figure 2 .
Figure 2. Impact of site on modulus of elasticity.

Figure 2 .
Figure 2. Impact of site on modulus of elasticity.

Figure 3 .
Figure 3. Impact of site on modulus of rupture.

Figure 3 .
Figure 3. of site on modulus of rupture.

Figure 4 .
Figure 4. Impact of site on impact bending strength.

Figure 4 . 14 Figure 5 .
Figure 4. Impact of site on impact bending strength.

Figure 6 .
Figure 6.Dependence of bending strength of spruce wood on density.

Figure 6 .
Figure 6.Dependence of bending strength of spruce wood on density.

Figure 6 .
Figure 6.Dependence of bending strength of spruce wood on density.

Table 1 .
Height and diameter of the sample trees.

Table 2 .
Wood density (in g•cm −3 ) of individual tree species according to the evaluated sites.

Table 3 .
Modulus of elasticity (in MPa) of individual tree species according to the evaluated sites.

Table 4 .
Bending strength (in MPa) of individual tree species according to the evaluated site.

Table 5 .
Impact bending strength (in J.cm -2 ) of individual tree species according to the evaluated site.

Table 5 .
Impact bending strength (in J•cm −2 ) of individual tree species according to the evaluated site.