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Sleswig-Holstein geest Schleswig-holsteinische Geestschlge e. Gdansk Lowland Danziger-Niederungsschlge f. British Lowland in Germany (Galloway, Ayrshire, Shorthorn) 2. Short-horned breeds Bos taurus brachyceros a. Schwyzer Schwyzer-Schlag b. Vorarlberger Vorarlberger-Schlge c. Alguer Alguer-Schlag d. Tyrol Grey Ober-Innthaler-Schlag e. Mrzthal Mrzthaler-Schlag f. Murboden Murbodner-Schlag g. Elling Ellinger-Schlag 3. Broad-headed breeds Bos taurus frontosus a. Bernese Berner-Schlag b. Fribourg Freiburger Schlag c. Ansbach-Triesdorf Ansbach-Triesdorfer Schlag d. Glan Glan Schlag e. Scheinfeld Scheinfelder-Schlag f. Limpurg and other breeds of Schwbisch-Limpurger-Schlag und brigen Wrttemberg Wrttemberger Schlge g. Kuhland Kuhlnder Schlag 4. Short-headed breeds Bos taurus brachycephalus a. Hrens Eringer-Schlag b. Tux-Zillertal Zillertal-Duxer-Schlag c. Pustertaler Pusterthaler-Schlag d. Voigtland Voigtlnder-Schlag e. Egerland Egerlnder-Schlag f. Devon 5. Crossed landraces a. Tauern Tauern-Schlge 1. Mlltal Mllthaler-Schlag 2. Pinzgauer Pinzgauer-Schlag b. Norische Norische Schlge 1. Mariahof Mariahofer-Schlag 2. Lavanttal Lavantthaler-Schlag c. Harz Harzer-Schlag d. Kampeten Kampeten-Schlag Table S2. Classification according to Werner (1912) ADDIN RW.CITE{{37 Werner, A.H. 1892/a;}}[2] Breed names are translated according to Masons World Dictionary of Livestock Breeds. German names are added if they are fairly different from English translation. (A) Bovini Group I. Buffaloes Group II. Wisents, Bisontina 1. European wisent: Bison europaeus Owen. Syn.: Bos Bison H. Smith 2. American bison: Bison americanus Gmel Group III. Bibos cattle, Bibovina 1. Banteng (Sunda Ochse): Bos sondacus Mller and Schlegel, Syn.: Bos banteng Raffl. 2. Gayal: Bos frontalis Lambert, Syn.: B. Gavaeus Roulin, B. Sylhetanus, Bibo Subhaemachalus Hodgs. 3. Gaur: Bos gaurus Hamilton, Smith, Syn.: Bibos cavifrons Hodgs. 4. Yak (Grunzochse): Bos (Pophagus) grunniens Linnaeus, Syn.:B. Pophagus Pallas 5. Zebu: Bos Zebu. Syn.: B. indicus Linnaeus, B. sondaicus indicus a. Indian zebu: Bos Zebu indicus Linnaeus Large Indian zebu: Bos Zebu Indicus major Fitzinger Medium large Indian zebu: Bos Zebu indicus medicus Fitzinger Dwarf zebu: Bos Zebu indicus minor Fitzinger b. African zebu: Bos Zebu africanus Fitzinger Galla zebu: Bos Zebu africanus Sanga Fitzinger (= B. sondacus africanus, B. sondacus longicornis) Ethiopian zebu: Bos Zebu africanus aethiopicus Fitzinger Afrikander: Bos Zebu africanus hottentottus Fitzinger Group IV. Common cattle, Taurina 1. Aurochs (Urochse): Bos primigenius Bojanus 2. Domestic cattle: Bos taurus Linnaeus (B) Domestic cattle. A. Primigenius Bos taurus primigenius Rtimeyer 1. Breed group of steppe cattle Grey Steppe (Podolische Steppenrasse), B.t.pr. podolicus Kalmyk (Kalmcken) and Kazakh (Kirgisen), B.t.pr. nomas Donube Steppe (unterer Donau), B.t.pr. dacicus Balkan Mountain steppe-shorthorn, B.t.pr. montanus Hungarian (Siebenrgische) Steppe, B.t.pr. hungaricus Romagnola, B.t.pr. romanicus 2. Breed group of Lowland cattle Germanische Rasse, B.t.pr. germanicus Friesian, B.t.pr. germanicus var. frisius Unicolored Red-brown East-Friesian, (Schsische) B.t.pr. germanicus var. saxonicus Flemish Red, B.t.pr. germanicus var. flandricus Normande, B.t.pr. germanicus var. normannus [British] Anglo-Saxon breeds, B.t.pr. germanicus anglosaxonicus 3. Breed group of British cattle White Park, B.t.pr. ferus Highland, B.t.pr. scoticus 4. Breed group of Russian cattle Great Russian Land cattle, B.t.pr. sarmaticus 5. Breed group of Scandinavian Swedish Mountain, B.t.pr. suecicus Fitzinger Finnish Norwegian Mountain [Telemark], B.t.pr. norvegicus Fitzinger B. Broad-headed cattle Bos taurus frontosus Nilsson 1. Breed group of spotted lowland cattle Gotland [Smland-Gotland], B.t.front. goticus Simmental (Burgundisch) breeds, B.t. front. burgundicus 2. Breed group of unicolored valley cattle Yellow Franconian (Frnkisch), B.t.front. franconicus Fitzinger Austrian Blond (Norisch), B.t. front. noricus. Piemontese, B.t.front. piemontanus C. Long-headed cattle Bos taurus longifrons Owen 1. Breed group of Alpine cattle. B.t.long. alpestris A. Wagner Brown Mountain, B.t.long. alp.var. brunneus Grey Mountain, B.t.long. alp.var. griseus Gelbvieh, B.t.long. alp.var. flavus 2. Breed group of French long-headed cattle Parthnais (Venden)and related breeds, B.t.long. ligeriensis Sanson Gasconne, B.t.long. vasconiensis Channel Islands cattle [Jersey, Guernsey], B.t.long. isolanus 3. Breed group of long-headed cattle in southeastern-central Europe Polish Brown, B.t.long. polonicus Adametz Illyrian and Busha, B.t.long. illyricus Adametz D. Short-headed cattle Bos taurus brachycephalus Wilckens 1. Breed group of Iberian cattle Iberian, B.t.brach. ibericus North- African [Algerian Brown Atlas] cattle, B.t.brach. africanicus Italian island cattle [Sicily, Corsica] and Camargue, B.t.brach. isolanus Pyrenean breeds, B.t.brach. pyreneus Limousin and pre-Blonde dAquitaine breeds, B.t.brach. aquitanicus Sanson 2. Breed group of Celtic highland cattle Celtic [Kerry, Welsh and Bretonne P.N.], B.t.brach. celticus Longhorn, B.t. brach. licestriensis Fitzinger British [Devon, Sussex, Hereford], B.t.brach. britannicus Salers (Auvergne), B.t.brach. alverniensis Fitzinger [Italian] Grey Alpine, B.t. brach. Italicus Tauern or Bunte Tiroler [Tuxer, Zillertaler, Pustrertal, Herens, Vosges], B.t. brach. tauricus Salzburger [Pinzgauer, Pongauer, Lungauer, Mlltaler, East-Styrian Spotted] B.t.brach.salisburgensis German Red Highland, B.t.brach. teutonicus Red Blazed (whiteheaded) Middle German Red group E. Breeds of German colonies in Africa. Table S3. Survey (A) and classification (B) of cattle breeds according to Keller (1905) ADDIN RW.CITE{{41 Keller 1905/a;}}[3]. (A) Survey The European Stem 1. Steppe cattle or Podolian cattle (Bos primigenius podolicus) The many strains of Steppe cattle are of strong constitution with heavy long to medium long horns, high withers, retracted belly and sloping rump. The hair color is commonly light, whitish grey, ash grey or dark grey; the hair course, bristly, sometimes curled. The skin is firm. The Steppe breed make good working cattle, on the other hand, the udder is small and the milk production low. The Steppe breed are spread south-eastwards from the Alps over Hungary, Romania, in Turkey, Greece, and south Russia. In the east it spread into Australia and also the South-Siberian cattle are related. Westward this type is found in Italy, where it was introduced in the 6th century by the Langobardian King Agiluf and penetrated from Hungary. Some scientist classify the heavy-horned and strong build cattle of the Iberian Peninsula also as Steppe breed, although this is incorrect as these are of a completely different origin. 2. Lowland cattle (Bos primigenius hollandicus) In its most beautiful form we find it in the heavy polder cattle form Holland, furthermore in East Friesia, Schleswig-Holstein and Oldenburg. The primigenius-like head is long, but with rather short, forward curved horns; the neck long with a small dewlap, the withers low, the hips wide with extended pelvic bones. The thin skin is flexible, the hair fine and commonly shining. The coat color seems mostly pied in black or red. The work capacity and beef production are outstanding; however most lowland cattle are also exemplary concerning milk production. 3. Frontosus cattle (Bos primigenius frontosus) Presumably these were introduced from northern Europe, but in their complete development they are currently found in west and north Switzerland. In their breed characteristics, they are most distinct from the original stem form, as the form of the skull has a roof like forehead. The medium long horns are flattened, grow downwards, then to the sides with the tips upturned, though often also downwards and sideways. The dewlap is well developed. The body conformation is varied with enormous animals among the highly bred strains. The coat color is black- or red-pied (Fleckvieh). Most well known is the red flecked Simmental strain, which is extensively exported for breeding purposes. Thereafter the black-pied Freiburger [Fribourg] strain, of which the horn form and skull are primigenius-like, and which in recent times is pushed back. The Longhorn breed, which is widely spread in England, can also be regarded as belonging to the frontosus group. According to Rtimeyer the Frontosus breed is a culture product developed from the Primigenius breed. The Oriental Stem 1. Zebu breed (Bos sondaicus indicus) This breed is spread over southern Asia, but most frequently found and most characteristic in India. In a wide belt from the northeast Horn of Africa into the German East-African colonial territory the cattle are strong influenced by Indian blood, which can be explained by ancient commercial trade. The Indian zebu shows the most variation. Besides powerful strains dwarfed cattle are found, the body is somewhat low [on the legs]. The fat hump is often well developed; the thin dewlap hanging low from the neck. The conformation of the head is very characteristic: commonly of strong concave profile, short horns, at the best medium, pointing backward in line with the forehead profile, with the tips turned inward. In shorthorned forms these point upward and backward, thus forming a 45 degree angle with the axis of the skull. The ears are peculiar, commonly hanging down. The color of the hair shows nuances, in milky white, grey-white, yellow, red-brown and pied; in pied animals the splashes are not sharply drawn. 2. Sanga breed (Bos sondaicus africanus) An African form and most pure found in the Abyssinian Sanga, though also spread from the highlands of Habesha to the upper Nile and Lake Chad. The horns are larger then in the related Indian zebu, most common lyre-formed and not so strong pointing backward but more upward. The body is agile and high on the legs with a fat hump on the back; the tail is long. In the lowlands the coat color is whitish, grey white, reddish-brown or red-pied; in the highlands black is favored because the Abyssinians believe that this is warmer. As draught animal and beef producer these animals are of great importance; the milk production is low. 3. Giant horned breed (Bos sondaicus longicornis) It obvious represents a breed, which originated from common Sanga cattle. Though, it seems to be ancient as we find these longhorned cattle already depicted during the earliest Egyptian dynasties. Currently they are pushed back to Central Africa, and to be met with in the Lake District with the agricultural colonies of Abyssinian descent as Watusi cattle. However populations are also found in southern Abyssinia. On the whole these peculiar African breed is diminishing. Medium strong, unicolored mahogany to dark brown-pied; in southern Abyssinia also brindled specimens can be found. The horns are mighty and are more than one meter in length, 4050 cm in circumference at the bases; growing upward, bending backward and upward as in the Sanga; the tips bending backward, or more often inward; not uncommon the horns are closed [at the tips]. Scientifically the breed is not important. 4. Brachyceros breed (Bos sondaicus brachyceros) On the contrary to the previous breed one finds brachyceros cattle in the periphery of the area where oriental domestic cattle are endemic, which encompasses East Asia, then West Asia, North Africa and more important Europe. The breed seems ancient, as it is found in the times of the pharaohs in the Nile valley and has a remarkable similarity with European Braunvieh [Brown Mountain cattle]. It holds small, elegantly build breeds. The fat hump has completely vanished. The horns are small, upward growing and upward bend, the forehead is pushed in between the eye sockets and the muzzle is fine. The back part of the head bulges in a hump which slopes sharp downward; the corners grow only seldom into horn roots (as for instance by Sardinian cattle). The color is commonly dark. Very good examples of this type are Algerian and Moroccan cattle. In Europe they must be put in the cattle groups which almost all are rooted in the Torfrind [Neolithic village cattle]. In the central Alps we find Braunvieh, which in its most pure form is kept pure in the region around the Gotthard Massive, but has also spread in Vorarlberg, the Bavarian Alps and Tyrol Alps. The coat is unicolored and, varies from dark brown to light mouse grey. Characteristic for the breed is a dark muzzle with light muzzle band and a light colored eel stripe. In Eastern Europe Albanian cattle, Illyrian cattle and wide spread Polish Red, also found in northern Russia, have to be regarded as belonging to the brachyceros stem. Well-bred and with famous dairy qualities are the small island cattle or Jersey cattle. 5. Brachycephalus breed (Bos sondaicus brachycephalus) Closely related to the previous breed, and clearly developed from artificial breeding on European soil. In the Alps and especially in the north of the Alps their remains have turned up in great numbers. The short-headed type is characterized by its short head. The forehead is very wide between the eyes. The rounded horns are strong, often very heavy and lyre-formed, white with black tips, often pointed somewhat downwards. The color of the hair is blackish-brown with a light eel stripe, but may also be reddish brown or mahogany. As with Swiss Brown a mahogany muzzle band and eel stripe of the same color is seen in the darker variants. We have observed heavy built, heavy boned short-headed cattle, besides small, elegantly built forms. Important for these breeds is the insular spreading. It is to be found in the southern, long stretched valleys of Valais and is in Switzerland known by the name of Eringer [Hrens] breed. These small cattle often had white parts, but are currently commonly unicolored black or dark brown with reddish tinge with mahogany muzzle band and an eel stripe. Related are the Duxer [Tuxer] and Pustertal strain, also the Voigtland and Egerland strains, as well as the Devon from the English counties Devon, Sussex and Hereford. Heavy short headed cattle with mighty lyre-formed horns and extreme short heads are found on the Iberian Peninsula. 6. Akeratos breed (Bos sondaicus akeratos) These are completely without horns and apparently developed independently in several places in the Old World from shorthorned cattle. Outside Europe polled cattle already turned up in the times of the pharaohs of old Egypt. They are also found in Somalia, while in Unyoro in central Africa most cattle are polled and humpless. Polled cattle are presumed to have also been in the possession of the Scythians. Possibly they were spread to the north, as currently they are mainly spread over northern Europe, Scandinavia (Fjellras) [Swedish Mountain], Iceland, Scotland, England and Wales, in northern Russia and sparsely in Oldenburg. The coat color is commonly white; but yellowish-red, brown-red and black is also observed. (B) Classification Group name Latin name type description typical breed European Stem Bos primigenius Steppe cattle B.p. podolicus work type with heavy horns Grey Steppe cattle, Italian white cattle Lowland cattle B.p. hollandicus dairy cattle with long head Dutch-Friesian, NW German Frontosus cattle B.p. frontosus broad head, horns downward Simmental-Fleckvieh. Oriental Stem Bos sondaicus Zebu B.s. indicus humped, often hanging ears Indian, East-African zebus Sanga B.s. africanus upward directed lyre horns, slim, Galla, Dinka, Fulani zebus long-legged Giant horned B.s. longicornis giant horned sanga Watussi Brachyceros B.s. brachyceros shorthorned, unicolored, Swiss Brown, Busha, Polish Red, dark muzzle, light muzzle band Jersey, Brown Atlas. and eel-stripe Brachycephalus B.s. brachycephalus very broad forehead , Herens, Devon, Sussex, Hereford brown-black to red-brown Akeratos B.s. akeratos naturally polled European, African. Table S4. Classification according to Holecek Holleschowitz (1939) ADDIN RW.CITE{{113 Holecek Holleschowitz, C. 1939/a;}}[4]. I. Breed group of longhorn (aurochs) descent A. Lowland breeds (Die Rassen des Niederungsrindes) 1. East-Friesian 2. Jeverlnder 3. Wesermarsch 4. East-Prussian Lowland 5. Red-Pied Schleswig-Holstein 6. Red-Pied Lower Rhineland (Niederrheiner) 7. Red-Pied Westphalian 8. Dutch breeds: a. Meuse-Rhine-Yssel b. Dutch-Friesian c. Groningen Whiteheaded 9. French Lowland breeds: a. Flemish Red b. Normande c. Bordelais 10. Shorthorn 11. Ayrshire 12. Jutland Black-Pied 13. Swedish Lowland 14. Telemark 15. Trondheim B. Broad-headed Fleckvieh en Blondvieh (Gruppe der breitstirnigen Rassen) 1. Simmental 2. Fribourg (Freiburger) 3. Miesbacher (Oberbayrisches Alpenfleckvieh) 4. South-German Fleckvieh (Hhenfleckvieh) 5. Tyrol Spotted (Unterinntaler Fleckvieh) 6. East-Styrian Spotted (Oststeirisches Fleckvieh) 7. Ennstaler Spotted (Bergschecken) 8. Bernese red spotted (Berner-Rotschecken) a. Bonhyhdi b. Kravarsk (Kuhlnder) c. Hrbineck (Schnhengster) d. Hanna-Berne 9. French Simmental a. Montbliard b. Comtois (Tourache) c. Abondance (Chablaisienne) 10. Italian Simmental 11. Yellow Franconian: a. Franconian (Maintaler) b.Glan-Donnersberg c. Limpurg d. Lahn 12. Carinthian Blond 13. Murboden 14. Fmeline, Bressane 15. Charolais C. West-European breeds of aurochs descent 1. Pinzgauer 2. Salers (Auvergne) 3. Limousin 4. Garonne 5. Pyrenean: a. Bazadais b. Lourdes c. Gascon 6. Parthenais 7. Devon 8. Sussex-Rind 9. Hereford D. Primitive, land races of aurochs descent 1. Hungarian Grey 2. Romanian Steppe 3. Moldovian [Bessarabian] Grey 4. Ukrainian Grey 5. Italian White breeds a. Romagnola b. Maremmana c. Chianina d. Marchigiana e. Bolognese f. Italian [Apulian] Podolian 6. Andalusian 7. Barosso 8. Galician Blond 9. [Scottish] Highland II. Breed group of shorthorned cattle A. European shorthorn breeds 1. Jersey 2. Guernsey 3. Kerry 4. Bretonnte Pie-Noir 5. Red Danish 6. Angler 7. Middle German Red a. Bavarian Red b. Vogelsberg c. Vogtland Red d. Waldeck f. Harzer Red g. Odenwald h. Westphalian Red 8. Polish [Silesian] Red [Lowland] 9. Polish Red [Highland] B. Alpine shorthorn breeds 1. Swiss Brown 2. German Brown [Allguer] 3. Vorarlberg Brown [Montafon] 4. Tyrol Grey [Oberinttal] 5. Waldviertel 6. Slovakian Red C. Primitive, landrace shorthorns 1. Illyrian Shorthorns a. Polim Busha b. Busha and Illyrian Dwarf 2. Macedonian Busha 3. West-Macedonian 4. Greek Shorthorn 5. Rodope Shorthorn 6. North- African Shorthorn a. Tunesian Guelma b. Algerian Guelma c. Kabyle d. Moroccan Brown Atlas 7. Anatolian 8. Caucasian mountain cattle 9.Krim mountain cattle 10. Carpathian 11. Russian Landcattle III. Breed group of short-headed cattle 1. Tux-Zillertal 2. Hrens (Eringer) 3. Tarentaise 4. Cheb (Egerlnder). IV Breed group of polled cattle 1. Swedish Mountain (Fjllras) 2. Finnish 3. North- Russian Polled Table S5. Classification according to Amschler as described by Haring et al. (1961) ADDIN RW.CITE{{139 Haring, F. 1961/a;}}[5] Names of extinct breeds or types are in italic. Type Breeds Primigenius Hungarian Grey cattle Podolian Grey Steppe cattle Egyptian longhorn Apis bull English Longhorn Scottish Highland Brachyceros Tyrol Grey Jersey Guernsey Angler Red Danish Shorthorn Dutch-Friesian German Black-Pied German Red-Pied Double Purpose Red Highland (German and Polish) Primigenius influenced by brachyceros Simmental Fleckvieh Pinzgauer Murboden Carinthian Blond German Gelbvieh Telemark Limousin Tarentaise Hereford Brachyceros influenced by primigenius Swiss Brown Red-pied and Black-pied Lowland strains Ayrshire Swedish Red-and-White Kholmogory Yaroslavl Glan and Donnersberg Waldviertel Blond Aberdeen-Angus Breton Black-Pied Normande Table S6. Classification according to Sanson (1884) ADDIN RW.CITE{{69 Sanson, A. 1884/a;}}[6]. Longheaded (Dolichocphales) Lowlands strain (B.t. batavicus): Hollandaise [Dutch], Flamande [Flemish] Germanic strain (B.t. germanicus): Normande Irish strain (B.t. hibernicus): Bretonne, Froment du Lon, Jersiase [Jersey] (Normande influence) British strain (B.t. britanicus): none Alpine strain resembling the brachyceros (B.t. alpinus analogue Brachyceros): Schwitz [Swiss Brown], Tarentaise, Gasconne Aquitaine strain (B.t. aquitanicus): Blonde dAquitaine, Limousine, Blonde des Pyrnes (Lourdaise) Shortheaded (Brachycphales) Asian strain (B.t. asiaticus): Camargue Iberian strain (B.t. ibericus): Corse, Blonde des Pyrnes (Basquaise et Barnaise) Vendenne strain resembling the primigenius (B.t. liguriensis, analogue primigenius): Parthenaise, Aubrac. Auvergnate strain (B.t. arvernensis): Salers, Ferrandaise Jura strain (B.t. jurassicus) analogue B.frontosus): Pie-rouge de l Est, Montbliarde, Abondance, Charolaise Scottish strain (B.t. caledonensis): none. Crossbreds (French) Jersey, Maine-Anjou, Bazadaise, Mzenc, Villard-de-Lans, Vosgienne Table S7. Classification according to Diffloth (1914) ADDIN RW.CITE{{114 Diffloth, P. 1914/a;}}[7]. The breed groups are divided into short-headed and long-headed types as by Sanson ADDIN RW.CITE{{69 Sanson, A. 1884/a;}}[6]. 1. Des Pays Bas [LowLand] Black-pied, red-pied, blue-pied and unicolored red lowland type breeds of England, Holland, Germany, Denmark, France and Belgium (including the Shorthorns) 2. Germaniques [Germanic] Normande, red-pied Holstein 3. Irlandais [Irish] Bretonne, Bordelais, Kerry, Devon, Ayrshire, Jersey, Guernsey 4. Du basin de la Loire Parthenais, Marchois, Aubrac, Angls-Cevennes [Venden] 5. dAquitaine [Aquitanian] Limousin, Garonnais, Agenais, Lourdais, Pyrenean, Bazadais 6. Auvergnat [Auvergne] Salers, Ferrandais, Mzenc, Central German Red Highland 7. Jurassique [Jura] Swiss and French Simmental type breeds, the Charolais, East-French blond breeds, German and Austrian white-backed breeds, German Yellow and the Hereford 8. Des Alpes [Alpine] Grey and Brown breeds of Switzerland, Germany, Austria, France (including the Tarentaise and Gasconne), and Italy (including the Piemontese) 9. Ibrique [Iberian] Les Landes, Corsican, North-African Atlas breeds, Sicilian, Sardinian, Podolian and Italian Podolian, Spanish and Portuguese breeds 10. Asiatique [Asian] Camargue 11. Britannique or scythe Aberdeen-Angus, Galloway, Suffolk, [British or Scythian] polled Norfolk, Norwegian, Russian, Icelandic 12. cossais [Scottish] Highland, White Park Table S8. Group classification to cranium, according to Kaltenegger (1904) ADDIN RW.CITE{{79 Kaltenegger, F. 1904}}[8] 1. Long-headed type (syn. shorthorn) Bos taurus longifrons (= brachyceros) Montafon, Rendena 2. Flat-fore-headed type Bos taurus planifrons (= primigenius) Oberinntal, Lechtal, Etchtal, Wipptal, Murboden, Mrztal 3. Larg-headed type Bos taurus grandifrons (= frontosus) Maltein, Mariahof, Lavanttal, Pinzgauer, Mlltal, Ennstal 4. Broad-headed type (= .short-headed) Bos taurus latifrons (= brachycephalus) Zillertal, Tux, Pustertal Table S9. Classification system of Baron (1928) ADDIN RW.CITE{{140 Baron, R. 1928/a;}}[9], SystmeCoordonnes baroniennes. (1) Morphology (La plastique) (a) Profile (silhouet) Les variations du profil (silhouette) straight back and legs rectilignes crocked back and legs convexilignes hollow back and X type legs concavilignes (b) Types (proportions) Les variations des proportions medium (= non specific mdiolignes ( msomorphes) or dual-purpose) large and long (= dairy type) longilignes ( dolichomorphes) large and rounded (= beef type) brvilignes ( brachymorpes) (c) Size Les variations du format large grand format ( hypermtriques) medium (135 cm at the withers) format moyen ( eumtriques) small petit format (hypomtrique) miniature trs petit format (ellipomtrie) (2) Coloration (La phanroptique) of coat, muzzle and mucosa. (3) Development (L nergique): production type (corresponds with 1b). Table S10. Coordinates (A) and classification (B) according to Dechambre (1913) ADDIN RW.CITE{{80 Dechambre, P. 1913/a;}}[10] Names of extinct and ancestor breeds are in italic. (A)Coordinates according to Baron (Coordonnes baroniennes) Skull Frontline of the skull Concave (Front concave, proceros) Flat (Front plat, orthoceros) Convex with high poll (Front convexe chignon saillant, opisthoceros) Size of horn Short (Cornes courtes, brachyceros) Medium (Cornes moyennes, msoceros) Bend inward (crowned) (Corne en couronne) Sickle-formed (Corne en croissant) Bend sideward (Corne en roue, trochoceros) Long (Cornes longues, dolichoceros) Bend in a hook, upward (Corne en crochet) Lyre-formed (Corne en lyre, lyriceros) Long, upturned and twisted (Corne en tire-bouchon, strepsiceros) Proportion Size large [long, tall] (les formes longiligne [tir, lanc]) intermediate [average] (mdioligne [intermdiaire entre les deux prcdents]) small [stocky, stout] (brviligne [trapu, ramass]) Weight heavy [more than average] (hypermtrique) average [averaging 650 kg] (eumtrique) light [less than average] (ellipomtrique) Basic types (groupes plastique) Bos primigenius evolved into the rectilinear type (type rectiligne) Bos frontosus evolved into the convex type (type convexiligne) Bos brachyceros evolved into the concave type (type concaviligne) (B) Classification 1. Rectilinear breeds, flat scull Intermediate size Heavy Primitive form: Bos primigenius Average Vendenne, Parthenaise, Marachine Light Ayrshire, Kerry, Bretonne Pie Noir Small size Primitive form: Bos brachyceros Average Brown breeds of Switzerland and central Europe Swiss Brown, German and Austrian Brown, Allguer, Italian Brown, Pontremolese, Piemontese French blond and grey breeds Tarentaise, Marchoise, Aubrac, Cvennes, Gasconne Light Carhaix pie rouge, Corsie, Sarda Large size Average Large Grey Steppe breeds and derived breeds Steppe breeds of Asia and Russia, Podolian Steppe, Hungarian Steppe, Romagnola, Pugliese and Italian derivatives. 2. I. Concave formed breeds, horned Intermediate size Cotentine, Normande Heavy German red-pieds, Breitenburg, Niederrhein Average Wild White Park and Scotch Highlan. and light Small size Primitive form Bos longifrons Average Lowland breeds of N. and W. Europe: Jutland, Dutch-Friesian, German Black-Pied, Flamande, Danish Red, Angeln, Devon derived small Durham, Bordelaise Island breeds and from La Manche Jersey, Guernsey, Cotentine. Light Large size Primitive form Bos mauritanicus Average Spanish and Portuguese Brown breeds Maronesa, Barrosa, Lidia, Franqueira Camargue Light Brown Atlas 2.II. Polled breeds North Russia, Scandinavia and Island Zyriane, Swedish Mountain, Icelandic British breeds Galloway, Aberdeen-Angus, Red Poll, Sarlabot. West-African breeds. Brazilian Mocha [= Mocho nacional] 3. Convex formed breeds Intermediate size Average and medium-large German and C. European Blond breeds Yellow Franconian, Limpurger, Glan-Donnersberg, Mainland French Blond breeds: Fmeline, Mezenc, Villard de Lans, Limousine, (Garonnais), derived: Bazadaise Blond breeds of Spain and Portugal Light Pyrenean breeds Barnaise, Lourdaise, Landaise, Central-Pyrenean [= Casta] Average Salers, Ferrandaise Red Spanish Rubia Gallega, Asturiana, Minhota, Arouquesa. derived: Caracu. German Red Highlan. Light Harz Red, Voigtland Red Small size Primitive form Bos frontosus Light Swedish Red-Pied, Norfolk Average and medium large Hereford, Longhorn Pied breeds of central Europe and Germany Pinzgauer, Hinterwalder and derivates, Tux-Zillertaler Swiss pied breeds Simmental, Fribourg French breeds Mancelle, Comtoise, Montbliarde, Abondance,Vosgienne, Bressane, Morvandelle, derived: Charolais, Nivernais Table S11. Classification according to Chacrin and Dumont (1921) ADDIN RW.CITE{{81 Chacrin, E. 1921/a;}}[11] in the Larousse Agricole [Larousse Agricultural Encyclopedia]. Extinct breeds are in italic. Breeds Sub-breeds Group I: Flat (Droit)forehead and poll flat, horns growing in line with the poll. Vendene Parthenaise, Maraichine, Poitevine, Nantaise. Aubrac Laguiole, Angles, Montagne-Noire Swiss Brown Schwyz Tarentaise Gasconne Mirandaise Bretonne Pie Noir De Rennes, De Carhaix Breton Pie Rouge Des Ctes-du-Nord Froment du Lon Group II: Concaveforehead hollow, horns growing out in front of the poll. Hollandaise and derivates Black-Pied Dutch-Friesian, Meuse-Rhine-Yssel, Groningen Whiteheaded, Friesch, Bleue du Nord, etc. Flamande Artsienne, Picarde, Maroillaise, de Bergues Durham or Shorthorn Durham-Mancelle Normande Cotentine, Augeronne, Cauchoise Jersey Brown Atlas [Tunisian] Kef, [Algerian] Geulma, Moroccan Brown Atlas Group III: Convexbulging poll, horns flattened and growing out behind the poll. Limousine Garonnaise Agenaise, Prigourdine Bazadaise Villard-de-Lans Mzenc Fmeline Barnaise Basque, dUrt Lourdaise Casta d Aure, de Saint-Girons Salers Ferrandaise Pie rouge de l Est and Jurassique Simmental, Bernoise, Comtoise, Montbliarde, Abondance Vosgienne Charolaise Nivernaise Table S12. Group classification to region and color according to (A) Kaltenegger (1904) ADDIN RW.CITE{{79 Kaltenegger, Ferdinant 1904/a;}}[8] and (B) Mller (1957) ADDIN RW.CITE{{82 Mller, Wilhelm 1957/a;}}[12]. (A) Kaltenegger (1904) ADDIN RW.CITE{{79 Kaltenegger, Ferdinant 1904/a;}}[8] Western group: Brown Mountain (Braunvieh) Montafon, Rendena Grey Mountain (Grauvieh) Oberintal, Lechtall, Etschtal, Wipptal Central group: Red-pied (Rotbuntvieh) Zillertal, Pinzgauer, Mlltal, Ennstal Black-pied and Brown-pied (Schwarz- und Braunbuntvieh) Tux, Pustertal Eastern group: Blond (Blondvieh) Maltein, Mariahof, Lavantal Grey Mountain (Grauvieh) Murboden, Mrztal (B) Mller (1957) ADDIN RW.CITE{{82 Mller, Wilhelm 1957/a;}}[12] Region Type of cattle Western Rheatian zone western unicolored ones. Adjoining Norik zone central group of pieds Eastern Pannonic zone eastern group of unicolored breeds Table S13. Classification of Spanish cattle according to Snchez Belda (1981, 1984) ADDIN RW.CITE{{84 Sanchez Belda, A. 1981; 85 Sanchez Belda, A. 1984}}[13,14]. Group Type Presumed descent Tronco turdetano Bvido rubio turdetanao also rojo convexo Bos taurus turdetanus Red and brown breeds with pink nose and convex head profile in France, Pyrenees, Balearic Islands, north-western and southern Spain, Canary Islands; several British red breeds, German and Austrian yellow breeds. Tronco ibrico Bvido negro ortoide Bos taurus ibricus Brown and black, elegant cattle in central and South Spain. Tronco cntabrio Bvido Castao concavo cntabro Chestnut colored breeds with black nose in Cantabrian Mountains, northwestern Spain and northern Portugal, subdivided into Castaa concavo and Moreno del Noroeste ADDIN RW.CITE{{201 Garcia, M.L.G. 1981;202 Garcia, Jos Emilio Yanes 2000/pRef. in: ;}}[15,16] Tronco Castao ultraconvexo related to African cattle Murciana and Pajuna. Table S14. Classification according to Simon and Buchenauer (1993) ADDIN RW.CITE{{55 Simon, D.L. 1993/a;}}[17]. 1. Black-and-White Pattern group 1.1. Holstein-Friesian 1.2. Original Black-Pied group 1.3. Russian Black-Pied 1.4. Other Black-Pied 1.5. White Belted group 1.6. Color Sided 1.7. Whiteheaded 2. Black group 2.1. Iberian Black 2.2. English Black 3. Red Pattern group 3.1. Red-Pied group 3.2. Simmental group 3.3. Black Forest 3.4. Pinzgauer group 3.5. Hereford group 3.6. Ayrshire group 3.7. White lineback 4. Red groups 4.1. English Red 4.2. Shorthorn group 4.3. Central-European, Red Northern Central type, German Red Hill group, Baltic type 4.4. Scandinavian Red 4.5. Russian Red 4.6. Iberian Red 5. Brown group 5.1. Brown Swiss group 5.2. Alpine Brown 5.3. Brown Mountain 5.4. Iberian Brown 5.5. Balkan Brown 5,6. British Brown 6. Grey Cattle group 6.1. Grey Steppe/Podolian 6.2. Grey Mountain 7. Blue Cattle group 8. Blond Cattle 8.1. Blonde d Aquitaine group 8.2. Gelbvieh group 8.3. Limousin group 8.4. Channel Island Cattle group 8.5. Italian Blond 8.6. Iberian Blond 9. White Cattle 9.1. British White 9.2. Charolais group 9.3. Italian White 10. Multicolored group In this group are unimproved cattle breeds which vary in coat color. Relationships among these breeds are not documented Table S15. Classification according to Bougler (1998) ADDIN RW.CITE{{141 Bougler, J. 1998/a;}}[18]. Grey Steppe strain, found from the Russian steppe, through the Balkans and Italy as far as the Iberian Peninsula; in France represented only by the Gasconne. Brown strain, which occupy the southern flanks of the Alpine arch and the borders of the Mediterranean; Brown Mountain, Corse, Tarentaise, Aubrac, Parthenaise. Red-Pied Mountain strain, found on the flanks north of the Alpine arch; Simmental, Montbliarde, Abondance, Charolais. Red strain, which stretches from the North Sea to northern Russia: Flamande. North Sea strain, which today consists mainly dual-purpose and dairy breeds: PrimHolstein, Pie Rouge des Plaines, Normande, Maine-Anjou (renamed Rouge des Prs), Armoricaine. Breton strain: Bretonne Pie Noir, Froment du Lon, Jersey. Blond and Red strain, found from the Massif central to the southwest and further into Spain; Blonde dAquitaine, Limousin and Salers. Table S16. Classification according to Ramm (1901) ADDIN RW.CITE{{56 Ramm 1901/a;}}[19]. Group I: Breeds and strains from the Netherlands, Denmark and Germany Subgroups 13: Dutch, Danish, German Lowland strains a: dairy type b: dairy-beef type from the German low lands Subgroup 4: Swiss Mountain and adjoining German strains a: Simmental and Fleckvieh b: Brown Mountain Subgroup 5: white-backed landraces from the South-German highlands Subgroup 6: heavy, yellow and red highland strains from central Germany Subgroup 7: light build, unicolored red and red white-headed (blazed) breeds Group II: Breeds and strains from AustriaHungary Subgroup 1: mountain strains from the Austrian Alps a. Alpine grey b. Fleckvieh c. Blond and yellow Subgroup 2: landraces in Upper and Lower Austria, Bohemia, Mhren, Galizien, Krain and Kstland Subgroup 3:Hungarian breeds Group III: Strains from Russia Subgroup 1: Landraces from Russia, Poland, the Baltics and Finland Subgroup 2: South-Russian Steppe cattle Group IV: Breeds and strains from Sweden and Norway Subgroup 1: Swedish breeds Subgroup 2: Norwegian strains Group V: Breeds from Belgium and France Subgroup 1: Belgian breeds Subgroup 2: French breeds a. lowland breeds b. land and mountain breeds south of the Loire and upper Rhone c. breeds of central and eastern France d. breeds from Tunisia and Algeria Group VI: Breeds from Italy Subgroup 1: North-Italian breeds Subgroup 2: Central and South-Italian breeds Subgroup 3: Italian islands breeds Group VII: Breeds from Spain and Portugal Subgroup 1: Spanish breeds Subgroup 2: Portuguese breeds Group VIII: Breeds from Great Britain and America Subgroup 1: British breeds a. beef type strains b. dairy type strains Subgroup 2: American a. Texas cattle b. 3 imported breeds Table S17. Classification according to Zwaenepoel (1920) ADDIN RW.CITE{{90 Zwaenepoel, H. 1920/a;}}[20]. Names of extinct breeds are in italic. I. Dairy breeds of the northern French lowlands Rouge Flamande Normande Bretonne II. Cattle belonging to the Swiss type Pied breeds Comtoise, Tourache, Montbliarde, Gessiene, Abondance, Bressane, Vosgienne, Morvandelle, Mancelle Brown breeds Tarentaise, brown breeds of the Massif Central, Marchoise, Aubrac, Angls, Cvennes, Gasconne, Corse III. Cattle of the Midi (Pyrenees) belonging to Romagna (a) and Spanish cattle (b) a. Lourdaise b. Basque, Barnaise, Casta, Camargue IV. Cattle of the Centre A. Central group Charolais-Nivernais, Limousine, Fmeline, Mzenc, Villard de Lans, Ferrandaise B. Western group Parthnaise, Garonnaise, Bazadaise, Bordelaise V. Imported cattle and crossbreeds Durham, Durham-Mancelle [pre Maine-Anjou] etc., Jersey, Dutch-Friesian, Flamande- Hollandaise, Swiss Brown, Race du Hainaut [pre Belgian White-Blue] Table S18. Classification according to Hansen (1927) ADDIN RW.CITE{{93 Hansen, J. 1927/a;}}[21]. Group 1. Cattle strains in Germany, Holland, Denmark and Switzerland. I. Lowland strains A. Dutch strains B. German lowland cattle 1. Dairy and dairy-beef strains; 2. Beef strains C. Danish lowland cattle II. Mountain cattle in Switzerland and Germany 1. Fleckvieh; 2. Brown Mountain III Land cattle races in Germany 1. Unicolored yellow; 2. Unicolored red and red-brown; 3. Pied and white-headed Group 2: Cattle strains in Austria Group 3: ,, ,, ,, Hungary Group 4. ,, ,, ,, Czechoslovakia Group 5. ,, ,, ,, Poland Group 6. ,, ,, ,, Lithuania Group 7. ,, ,, ,, Latvia and Estonia Group 8. ,, ,, ,, Finland Group 9. ,, ,, ,, Russia Group 10 ,, ,, ,, Sweden and Norway Group 11 ,, ,, ,, Belgium and France Group 12 ,, ,, ,, Italy, Spain and Portugal Group 13 ,, ,, ,, Great Britain and Ireland Group 14 ,, ,, ,, North America Table S19. Classification according to French et al. (1966) ADDIN RW.CITE{{59 French, M.H. 1966/a;}}[22]. Scandinavian and North-European group Denmark: Red Danish, Danish Black-and-White, Danish Jersey, Shorthor. Finland: Finncattle, Finnish Ayrshire Iceland: Icelandic breed Norway: Norwegian Red, South- and Westland, Color-sided Trnder and Northland, Telemark, Dle, Mlselv, Norwegian Jersey Sweden: Swedish Red-and-White, Swedish Friesian, Swedish Polled, Swedish Jersey United Kingdom and Ireland Aberdeen Angus, Ayrshire, Blue Albion, North Devon, Dexter, British Friesian, Galloway, Belted Galloway, Old Gloucestershire, Guernsey, Hereford, Highland, Jersey, Kerry, Lincoln Red, Longhorn, Red Poll, Shetland, Shorthorn, South Devon, Sussex, Welsh Black, White Park North Sea and Baltic Littoral Netherlands: Dutch Friesian, Groningen White-headed, Meuse-Rhine-Ijssel Federal Republic of Germany: Black-and-White Lowland, Red-and-White Lowland, Angeln, Shorthorn, German Simmental, German Brown, German Yellow, German Red, Pinzgauer, Small Spotted Highland, Murnau-Werdenfels Poland: Black-and-White Lowland, Polish Red, Red-and-White Lowland Western Europe Belgium: Red West-Flemish, Central and Upland, Red-and-White East-Flemish, Red and White Campine, Black-and-White Herve, Friesian, Meuse-Rhine Ijssel Luxembourg. France: Normany, French Friesian, Eastern Red-and-White, Charolais, Limousin, Salers, Armorican, Maine-Anjou, Brittany Black-and-White, Gascony, Flemish, Pathenay, Brown Swiss, Aubrack, Blond Pyrenean, Garonne, Tarentaise Alpine Europe Switzerland: Simmental, Brown Swiss, Fribourg, Hrens Austria: Austrian Simmental, Austrian Yellow, Pinzgau, Austrian Brown, Grey Tirolean, Black-and-White Lowland Czechoslovakia: Czechoslovak Red-and-White, Slovakian Red-and-White, Pinzgau The Iberian Peninsula and Italy Portugal: Miranda or Ratinha, Turino, Barrosa, Aroucesa, Alentejo, Mrtola, Minho, Algarve, Brava Spain: Pyrenean or Basque, Tudanca, Asturian, Leonese, Retinta or Extremea, Avila, Berrenda, Black Andalusian, Salamanca or Morucho, Zamora, Extremadura, Cceres, Murcia or Levantine, Black-and-White Lowland, Brown Swiss, Simmental, Fighting bull Italy: Brown Swiss, Friesian, Red-and-White Valdostana, Black-and-White Valdostana, Rendena, Burlina, Romagna, Chiana, Marche, Maremma, Pugliese, Piedmont, Grey Alpine, Modena, Simmental, Reggio, Modica, Garfagnina, Pisa, Tarina, Modica-Sardinian, Sardinian The Balkans and Turkey Hungary: Hungarian Red-and-White, Simmental, Hungarian Brown, Hungarian Stepp. Romania: Romanian Bua, Romanian Steppe, Romanian Spotted, Maramures Brown, Transylvanian Pinzgau, Romanian Red Yugoslavia: Yugoslav Bua, Yugoslav Red-and-White, Pinzgau, Istrian, Brown Slovenian Bulgaria: Grey Iskur, Sofia Brown and Montafon crosses, Kula and Simmental crosses, Shorthorned Rhodope, Red Sadova and Ukrainian Steppe Albania Greece: Greek Shorthorn, Greek Steppe Turkey: Grey Steppe U.S.S.R. Kholmogor, Friesian, Oldenburg, Tagil, Yaroslavl, Ukrainian Red, Angeln, Red Danish, Estonian Red, Latvian Red, Suksun, Polish Red, Ukrainian White-head, Gorbatov Red, Yurino, Istoben, Ayrshire, East-Finnish, Simmental,Tambov Red, Bestuzhev, Ukrainian Grey, Hereford, Shorthorn, Aberdeen-Angus, Kalmyk Table S20. Classification according Doutresoulle (1947) ADDIN RW.CITE{{217 Doutresoulle, G. 1947}}[23]. Taurine type (a) derived from longhorned cattle: NDama and the breed of Chad (Kuri) (b) derived from shorthorned cattle: cattle of the Lagoons (West-African Shorthorn) Zebu type (after Curson and Epstein) (a) shorthorned zebu: zebus brachyceros (b) sanga zebu: true zebu Hamitic longhorn, with small, cervico-thoracic hump (c) West-African zebus with lyre-formed horns: NDama shorthorned zebu Table S21. Classification of West-African Livestock according to Mason (1951) ADDIN RW.CITE{{95 Mason, I.L 1951/a;}}[24]. Group I. Lake Chad cattle 1. Kuri 2. Kuri zebu crosses Group II. Small humpless cattle 1. Dwarf Shorthorn 2. NDama 3. Intermediate types a. Baoul (Ivory Coast) b. Gold Coast Group III. Humped humpless crosses 1. Djakor (Senegal) 2. Bambara or Mer (French Sudan) 3. Sanga (Gold Coast) 4. Borgu (Dahomey and Nigeria) 5. Biu (Nigeria) Group IV. Humped cattle (zebus) Subgroup A. Short-horned zebus 1. Maure 2. Tuareg 3. Azaouak 4. Shuwa 5. Fellata (Chad) 6. Sokoto Subgroup B. Medium-horned zebus 1. Diali 2. Adamawa Subgroup C. Lyre-horned zebus 1. Senegal Fulani 2. Sudanese Fulani 3. White Fulani (Nigeria) Subgroup D. Long-lyre-horned zebus 1. Red Bororo Table S22. Classification according to Joshi et al. (1957) ADDIN RW.CITE{{97 Joshi, N.R. 1957/a;}}[25]. Group I. Humpless or vestigially-humped cattle of the lower Nile valley and Mediterranean Africa. the cattle of Egypt, including the Damietta, Baladi, Saidi and Maryuti. Group II. Zebus of the sub Saharan zone with many points of similarity with the Indo-Pakistan zebus. This group may be subdivided into 1. Medium (a) and short-horned (b) zebus a. Adamawa zebu, Azaouak zebu, Maure zebu, Northern Sudan shorthorn zebu. b. Shuwa zebu, Sokoto zebu 2. Lyre-horned (a) and long-horned (b) zebus a. Fulani zebus (Nigerian , Senegal , Sudenase and White Fulani). b. MBororo Group III. Humpless, straight-backed cattle of West Africa a. NDama b. West-African Shorthorned cattle Group IV. Kuri cattle of Lake Chad, humpless and with characteristic bulbous horns Group V. Cattle of much of central and southern Africa from the flood plain of the Nile in the Sudan, through South-western Uganda, Rwanda and Burundi to the Rhodesias [Zimbabwe, Malawi], Bechuanaland [Botswana], Swaziland and Basutoland, characterized by large- or medium-sized lyre-shaped horns, small or vestigial humps and moderately sloping hindquarters a. Ankole cattle of Uganda, Ruanda-Urundi, eastern [Belgian] Congo and Tanganyika b. Barotse cattle of the western part of Northern Rhodesia [Zimbabwe] c. Basuto cattle d. Nguni cattle of Zululand and Swaziland e. Nilotic cattle of the southern Sudan f. Nioka cattle of the Eastern Province of the [Belgian] Congo g. Nganda cattle of Uganda h. Tonga cattle of the Southern Province of Northern Rhodesia [Zimbabwe] Group VI. Cattle types of East Africa, a large heterogeneous population composed of often ill-defined groups merging into one another and, in some cases, into types which have been listed in group V, but which all appear to be predominantly derived from zebu stocks similar to those of the Indo-Pakistan peninsula. a. Angoni cattle of the Eastern Province of Northern Rhodesia [Zimbabwe] b. Boran cattle of southern Ethiopia, Somalia and northern Kenya c. Bukedi zebu of Uganda d. Galla, Jiddu and Tuni cattle of Somalia e. Lugware cattle of the Belgian Congo and Uganda f. Nandi cattle of western Kenya g. Southern Sudan Hill zebu h. Tanganyika shorthorned zebu i. Toposa-Murle cattle of Southeastern Sudan Group VII. Africander [Afrikaner] cattle of southern Africa Group VIII. The Madagascar zebu Table S23. Classification of African cattle according to the Colonial Advisory Council of Agriculture, Animal Health and Forestry (1957) ADDIN RW.CITE{{99 Colonial Advisory Council of Agriculture Animal Health and Forestry 1957}}[26]. The cattle types of East, Central and South Africa Longhorn humpless Shorthorn humped Cervico-thoracic humped zebu (neck-humped) Thoracic humped zebu (chest-humped) Sanga The cattle types of West Africa Longhorn humpless Shorthorn humpless Cervico-thoracic humped zebu Thoracic humped zebu Sanga (either Cervico-thoracic nor thoracic-humped) The cattle of North Africa Shorthorn humpless Table S24. Classification of Indian and Pakistan cattle according to Joshi et al. (1953) ADDIN RW.CITE{{103 Joshi, N.R. 1953/a;}}[27]. Group I. The breeds classified in this Group are lyre-horned grey animals with wide foreheads, prominent orbital arches and a flat or dished-in profile. They are deep bodied, powerful draft animals. Breeds: Kankrej, Kenwariya (Kentkatha), Kherigarh, Malvi, Tharparkar (Thari). Group II. The breeds classified in this Group are short-horned, white or light grey in color with long coffin-shaped skulls. The face is slightly convex in profile. Breeds: Bachaur, Bhagnari, Gaolao, Hariana, Krishna Valley, Mewati, Nagori, Ongole, Rath. Group III. Breeds classified in this Group are more ponderous in build and have pendulous dewlaps and sheaths. They often have lateral and curled horns, and usually red or some shade of red color, being occasionally spotted: the best dairy breeds among zebus are found in this group. Breeds: Dangi, Deoni, Gir, Nimari, Red Sindhi, Sahiwal. Group IV. Breeds classified in this Group, are popularly termed Mysore cattle. They are characterized by prominent foreheads and long, pointed horns rising close together, and are, with few exceptions, poor milkers. Breeds: Amrit Mahal, Hallikar, Kangayam, Khillari. Group V. The breeds included in this Group are a heterogeneous mixture of distinct strains. They are found all over the Indo-Pakistan area and particularly in the Himalayas, in the hills of Baluchistan and in the rugged mountainous areas of North Pakistan. Breeds: Lohani, Ponwar, Siri. Group VI. The Dhanni breed is the only one included in this Group. It does not seem to fit into any of the other groups described, and therefore requires a separate classification. Table S25. Classification of Chinese cattle according to Epstein (1969) ADDIN RW.CITE{{49 Epstein, H. 1969/a;}}[28]. Mongolian Cattle Humpless Cattle of Northeast China (Manchuria) Pinchow and Sanho Dairy Cattle of China Peking Black-Pied or Peking Black-and-White Dairy Breed Humpless Dwarf Cattle of Tibet Cervico-Thoracic-Humped Cattle of Central China Chinchwan Cattle Nanyang Cattle Shantung Cattle Chowpei and Hwangpei Cattle of Hupeh Cattle of Szechwan, Hunan and Kweichow Zebu Cattle of South China Cattle of Taiwan Table S26. Classification of tropical cattle according to Maule (1990) ADDIN RW.CITE{{106 Maule, John P. 1990/a;}}[29]. Group I Zebu (Bos indicus; thoracic humped cattle) A. Indo Pakistan 1. Lyre-horned, grey 2. Shorthorned, white or grey coffin shaped skull 3. Lateral horned, red, red-and-white or black-and-white 4. Mysore breeds: grey, long backward pointed horns 5. Small hill cattle 6. Cattle of Sri Lanka B. African 1. North Sudan zebu 2. East-African shorthorned zebu 3. Madagascar zebu 4. West-African short- and medium-horned zebu 5. West-African lyre horned and long lyre-horned C. Brazilian D. Middle and Far East 1. Middle East 2. Southeast Asia 3. South China zebu 4. Taiwan zebu E. Brahman Group II Sanga (Cervico-thoracic humped cattle) A. Sudan and East Africa 1. Nilotic 2. Danakil 3. Ankole B. Southern African Longhorned 1. Setswana 2. Nguni 3. Africander C. Southern African Shorthorned 1. Mashona 2. Tonga 3. Basuto 4. Drakensberger D. Sanga zebu crosses 1. Ethiopian 2. East Africa Group III Humpless A. Humpless cattle of West and North Africa 1. West-African Longhorn 2. West-African Shorthorn 3. North-African Shorthorn 4. Ethiopian Shorthorn B. Middle East Shorthorn 1. Syria and Palestine 2. Cyprus 3. South-Anatolian Red 4. Kurdi C. Criollo of Latin America 1. South-American (except Brazil) 2. Brazilian 3. Central America 4. Caribbean D. Far East Group IV Humped humpless A. Zebu West-African Humpless 1. Zebu NDama / Dwarf 2. Zebu Kuri B. Zebu Brachyceros 1. Middle East 2. Far East C. Bos taurus Bos indicus 1. North America 2. Caribbean 3. South America 4. Asia 5. Africa 6. Australia Group V Bos bibos (domesticated species) A. Bos bibos javanicus and crosses 1. Banteng or Bali cattle 2. Banteng Ongole zebu B. Bos bibos frontalis 1. Mithan or gayal 2. Mithan Siri zebu Table S27. Classification and phylogeny according to Baker and Manwell (1980) ADDIN RW.CITE{{22 Baker,C M. 1980/a;}}[30] and Manwell and Baker (1980) ADDIN RW.CITE{{21 Manwell,C. 1980/a;}}[31]. (A) Classification Abbreviation on map I. European: Bos taurus L. A. North-European N 1. Scandinavian: Finnish, Icelandic, Norwegian Red, Swedish Mountain, Swedish Polled, Swedish Red Polled, Swedish Red-and-White 2. Celtic: Aberdeen Angus, Ayrshire, Belted Galloway, Dexter, Galloway, Highland, Kerry, Welsh Black, Wild White 3. Shorthorn: Beef Shorthorn, Dairy Shorthorn, Milking Shorthorn, Northern Dairy Shorthorn 4. English Lowland: Hereford, Lincoln Red, Longhorn, North Devon, Polled Hereford, Red Poll, South Devon, Sussex B. Pied Lowland L 1. Red-Pied Lowland: Campine, East Flanders, German Red-Pied, Meuse-Rhine-Yssel, Polish Red- and-White Lowland 2. Black-Pied Lowland: Belgian Black-Pied, Danish Black-Pied, Dutch Black-Pied, Estonian Black-Pied, Friesian, German Black-Pied, Gronigen White-headed, Kholmogor, Middle and Upper Belgium, Polish Black-and-White Lowland, Russian Black-Pied, Siberian Black-Pied, Swedish Lowland, Yaroslavl C. European Red brachyceros R 1. Lowland Red: Danish Red, Flamande, Latvian Brown, Red Angeln, West Flanders 2. East- and Central European Red: Bulgarian Red, German Red, Polish Red, White Russian Red D. Channel Island brachyceros C 1. Channel Island: Guernsey, Jersey 2. French: Brittany Shorthorn, Normande 3. Derived: Canadian E. Upland brachyceros U 1. Pinzgau type: Pinzgau 2. Spotted Mountain: a. Main group: Fleckvieh, Fribourg, Hinterwald, Hohenfleckvieh, Simmental, Vorderwald,Yellow Franconian b. Derived group: Aosta, Montbliarde, Red-Pied Friuli 3. Yellow Mountain: Austrian, German Yellow, Carinthian Blondvieh, German Yellow, Murboden, Waldviertel. 4. Brown Mountain: Austrian Brown, Brown Swiss, Bruna Alpina, German Brown, Murnau-Werdenfels, Rendena, Tarentaise 5. Grey Mountain: Tyrolean Grey 6. Hrens type: Hrens 7. Italian Mediterranean: Reggiana, Sarda 8. Balkan Mediterranean: Busha, Cika, Rhodope, Slovene Grey-Brown 9. Near-Eastern and North-African Mediterranean: Algerian Hill, Anatolian Black, Cyprus, Damascus, East-Turkish, Egyptian, Oksh, South-Anatolian Red F. Primigenius-brachyceros Mixed M 1. French: Charolais, Garonne, Limousin, Parthenaise 2. Italian: Agerola, Modica, Modica-Calabrian, Piedmont 3. Lowland Steppe: Bestuzhev, Red Steppe, Ukrainian Whitehead 4. Brown Mountain Steppe: Bulgarian Brown, Romanian Brown, Sofia Brown 5. Simmental Steppe: Slovakian Pied, Slovene Light Red Spotted 6. Eastern European: Ala-tau, Kostroma G. Primigenius P 1. Iberian primigenius: Alentejo, Criollo, Mixican Fighting, Texas Longhorn, Tinema 2. Italian podolic: Calabrian, Chianina, Marchigiana, Maremma, Modena, Romaga 3. Podolic: Grey Iskur, Grey Steppe, Hungarian Grey, Istrian, Romanian Steppe, Yugoslavian Steppe II. Turano-Mongolian: Kalmyk, Yakutian. III. East-Asian taurine types: Japanese Black, Japanese Brown, Japanese Shorthorn. taurine-indicine intermediates: Korean indicine (zebu): Mishima, Taiwan Yellow. Eurasia IV. Zebu: Bos indicus L. Z 1. Mohenjodaro type zebu: Kankrej, Malvi, Tharparkar 2. Shorthorned zebu: Hariana, Nagori, Ongole, Rath 3. Dairy type zebu: Dangi, Gir, Sahiwal, Sindhi 4 Mysore type zebu: Kangayam, Khillari 5 Hill breeds: Afghan, Kumauni 6. Ungrouped zebu: Bermejo, Desi, Gavathi, Indo-Brazilian, zebu V. Modern major crosses A. Taurindicus: Brahman, Droughtmaster, Santa Gertudis B. Taur-sanga: Bonsmara, Drakensberger C. Three-way: derivation: Renitel VI. African humpless A. Kuri type: Kuri B. Longhorn: NDama C. West-African Dwarf Shorthorn: Baoul, Ghana Shorthorn, Lagoon, Muturu VII. African humped A A. African Humped: 1. West-African zebu: Adamawa Gudali, Cameroon Red 2. West-African Shorthorned zebu: Shuwa Arab, Sokoto 3. East-African Shorthorned zebu: Angoni, Boran, Garre, Kajiado, Malawi, (North), Malawi zebu (South), Narok, North Sudan zebu, Tanganikan Shorthorned zebu, Teso00 4. Madagascar zebu: Malagasy 5. Fulani: Foulb, Gobra, Red Bororo, White Bororo 6. East-African zebu-sanga intermediates: Jiddu, Kenana, Nganda B. African Humped (Sanga) S Africander, Angolan Caf Cunene, Angolan Malanje, Angolan Quioenques, Ankole, Barotse, Caprivi-Sanga, Landim, Lesotho, Nguni, Manguni, Mashona, Nguni, Ovambo, Pedi, Sango, Tonga, Tuli (B) Phylogeny  Table S28. Classification and origin of Spanish breeds according to Vallejo (1990) ADDIN RW.CITE{{62 Vallejo, M. 1990/a/f, Fig 2;}}[32]. Group Breeds Mutant form Prehistoric strain I Blanca Cacerea Bos desertorum II Crdena Andaluza Bos primigenius Hahni III Rubia Gallega Bos t. Ibericus or B.t. primigenius Frieiresa Bos primigenius estrepsiceros Limiana Vianesa Caldelana Alistana Sanabresa IV Asturiana Montaa Bos brachyceros and B.t. brachyceros Asturiana Valles Bos brachyceros europeus Cachena Morucha Sayaguesa Figure S1. Phylogeny of Iberian breeds according to (A) Vallejo et al. (1990) ADDIN RW.CITE{{62 Vallejo, M. 1990/a;}}[32]; (B) Fernndez et al. (1998) ADDIN RW.CITE{{2 Fernndez, A. 1998/a;}}[33]. (A) Sayaguesa Morucha Cachena Asturiana Valles Asturiana Montaa Alistana Sanabresa Vianesa Limiana Frieiresa Rubia Gallega Gardena Andaluza Blanca Cacarea (B)  Gallega Cachena Barros Arouquesa Frieiresa Caldelana Limiana Mirandesa Vianesa Maronesa Table S29. Classification of French breeds according to Grosclaude (1990) ADDIN RW.CITE{{20 Grosclaude,F. 1990/a;}}[34]. 1. Northern breeds (Le groupe des races du Nord) Frisonne, Flamande, Maine-Anjou, Shorthorn 2. Central and Southwestern breeds (Le groupe des races du Centre et du Sud-Ouest) Charolais, Ferrandaise, Limousin, Salers, Aubrac, Blonde dAquitaine 3. Western and eastern breeds (Un groupe comprenant la fois des races de lOuest) Eastern: Vosgienne, Montbliarde, Pie-Rouge de Est, Brune, Abondance, Tarine. 4. Normande (la seule race Normande) Table S30. Classification of French breeds and related breeds from other countries according to Gautier et al. (2010) ADDIN RW.CITE{{19 Gautier,M. 2010}}[35]. 1. Northern breeds: Holstein, French Red-Pied Lowland, Maine-Anjou (Rouges des Prs), also related to Angus and Norvegian Red. 2. Central and Southwestern breeds: Aubrac, Blonde dAquitaine, Gasconne, Limousin, Salers, related to Charolais 3. Eastern breeds: Abondance, Montbliarde, Tarentaise, Vosgienne, also related to Brown Swiss 4. Parthenaise (Maraichine) 5. Bretonne Pie Noir 6. Normande The last 3 groups are separate, and intermediate between the Northern group and the others Table S31. Classification according to Alderson (1992) ADDIN RW.CITE{{108 Alderson, L. 1992/a;}}[36]. Group Distribution Typical breeds 1. Podolic Lower/Mid Danube, Balkans, Italy Grey Steppe, Maremmana 2. North-European Poland, Czechoslovakia, Germany, Denmark, Benelux, England Angeln, Danish Red, Flemish, Friesian, Polish Red, Lincoln Red, Sussex Shorthorn 3. Central Europe (a) Swiss group Alpine region, northern Italy Brown Swiss, Garfagnina, Pontremolese (b) Pattern group Germany, Austria, Switzerland, France East-French Red-Pied, Pinzgauer, Spotted Cattle (c)Yellow-Brown Central Germany through France and northern Spain to Portugal Murbodner, German Yellow, Tarentaise, Limousin, Blonde dAquitaine, Aubrac, Parthenaise, Villard-de-Lans, Leonese, Pyrenean, Asturian, Mirandesa, Minhota 4. Western Europe Southwest Iberia, Wales, Scotland, Ireland White Park, Kerry, Welsh Black, Negra Ibrica, Morucha, Retinta, Camargue Table S32. Classification according to Denis (2010) ADDIN RW.CITE{{94 Denis, B. 2010}}[37]. Names of extinct breeds and types in italic. This classification is based on an earlier classification from 1983 with addition of the last group. 1. Race primaire Batave Dutch-Friesian group Franaise Frisonne, Pie Rouge des Plaines, Flamand Durham derived Blue de Nord, Maine-Anjou (Rouge des Prs), Saosnoise, Armoricaine Normande 2. Race primaire Jurassique Jura group Charolaise, Montbliarde, Simmental Franaise, Abondance, Vosgienne, Villard-de-Lans Auvergne group Salers, Ferrandaise Aquitaine group: Limousine, Blonde des Pyrnes, Barnaise, Lourdaise, Bazadaise, Casta 3. Race primaire alpine Alpine group Brune de Alpes, Tarentaise, Aubrac, Gasconne, Mirandaise. Poitevin group Parthenaise, Marachine, Nantaise Breton grou: Bretonne Pie Noir, Froment du Lon, Jersiaise Rustic and semi-wild breeds from South France: Camargue, Corse Table S33. (A) Classification according to Felius (1995) ADDIN RW.CITE{{30 Felius, M. 1995}}[38] and (B) revised classification. Differences between the classifications of 1995 and 2010 are the following: 1. Changes in Group name For instance in Group 1 the name Celtic is now placed between quotation marks, since this would support the unfounded theory that these cattle are introduced by Celts. 2. Extended subgroup names. To subgroup 1A Northern Russia is added because a number of (mainly extinct) breeds from this region are here included, which were not described in the 1995 edition. In several subgroups of Groups 1 and 2 the region where the derived breeds are to be found are mentioned. 3. Change in subgroup names. Subgroup 2B now contains other than black-pied breeds, so the name is shortened to West- and Northeast-European Pied Lowland Dairy breeds. The name of subgroup 13A is shortened, indicating the whole region instead of separate countries. Zebus from subgroup 13A are indicated as large in order to distinguish these from the small zebus in the same region. 4. Change in the order of subgroup interchange. In Group 1, the Subgroups B and C changed places in order to bring closer together the two Scandinavian groups and the British groups. Geographic classification thus overrules the morphological typing (polled and horned cattle groups) as was done in the 1995 publication. 5. New subgroup. To Group 2 a subgroup is added containing a recent Danish multi breed. 6. Splitting subgroups. The first three subgroups in the Group 5 are arranged differently. Several breeds and varieties of Iberian cattle are only recently recognized and added in this classification. This allowed a more refined grouping to region and type. Since more American, Australian and New Zealand breeds could be described than in 1995, the subgroups of Group 16 have been split up. 7. Change in classification. Because since 1995 more zenga breeds have been recognized, all zenga breeds have been moved form Group 13 to Group 14. 8. Modernizing language. In Group 7 the name Middle East is replaced by Southwest Asia. Further, blood is replaced by influence. African zebu sanga types and breeds are now termed zenga. (A) 1995 Group 1 North-European Polled and Celtic breeds Subgroup 1A Polled breeds of Iceland, Scandinavia and the Baltics Subgroup IB Polled breeds of Great Britain and derived breeds Subgroup IC Longhorned dairy breeds of Scandinavia and Scotland Subgroup ID Celtic breeds of Great Britain Group 2 Lowland breeds of West and North Europe Subgroup 2A West-European red lowland breeds and derived breeds in Eastern Europe Subgroup 2B West- and North-European black-pied and red-pied Lowland dairy breeds and derived breeds in central and eastern Europe Subgroup 2C West-European red-pied and blue-pied Lowland dual-purpose and beef breeds and derived breeds in Eastern Europe Subgroup 2D British shorthorned breeds and derived breeds in Eastern Europe Subgroup 2E Lowland breeds of West and South England Subgroup 2F Breeds of the Channel Islands, and Northwestern France Group 3 Shortheaded and broadheaded highland breeds of Western and Central Europe Subgroup 3A Vosges and Black-Forest breeds Subgroup 3B Central-European red highland breeds Subgroup 3C Shortheaded Alpine breeds and derived breeds in Eastern Europe Subgroup 3D Central-European blond and yellow highland breeds Subgroup 3E Western- and Central-European broadheaded red spotted mountain breeds and derived breeds in Eastern Europe Subgroup 3F Charolais Group 4 Solid-colored highland breeds of Western Europe, the Alps and Eastern Europe Subgroup 4A Breeds of central France Subgroup 4B Grey and blond breeds of Southwestern France and the Pyrenees Subgroup 4C North-Italian fawn-brown breeds Subgroup 4D Central-European grey and brown mountain and derived breeds in Eastern Europe Subgroup 4E Illyrian shorthorn breeds of the Balkans and Greece and upgraded breeds Group 5 Breeds of Southwest Europe Subgroup 5A Breeds of the Cantabrian mountains and primitve (semi-feral) breeds of the Pyrenees, the Camargue, Corsica and Sardinia Subgroup 5B Blond-brown northwestern Iberian, Balearic and Canarian breeds Subgroup 5C Chestnut NorthWest-Iberian breeds Subgroup 5D Central and Southern Iberian black breeds Subgroup 5E Central and Southern Iberian red breeds Subgroup 5F Southeastern Iberian breeds Group 6 Podolian breeds of Italy and East Europe Subgroup 6A Large white breeds of Italy Subgroup 6B Podolian breeds of Italy and Croatia Subgroup 6C Podolian grey steppe breeds of Eastern Europe Subgroup 6D Podolian-Illyrian breeds of the Balkan and Anatolia Group 7 Shorthorned breeds of the Caucasus and Southwest Asia Subgroup 7A Humpless breeds of the Caucasus, SW Asia and derived breeds with exotic blood Subgroup 7B Damascus type breeds of the Southwest Asia and derived breeds with exotic blood Group 8 Indo-Pakistani type zebu breeds Subgroup 8A Zebu and zeboid breeds of the central Asia, Afghanistan and northwestern Pakistan and derived breeds with European blood Subgroup 8B Heavily-built breeds with convex forehead and derived breeds with European blood Subgroup 8C Shorthorned grey-white zebu breeds Subgroup 8D Zebu breeds with lyre-shaped horns and derived breeds with taurine blood Subgroup 8E Mysore zebu breeds Subgroup 8F Small zebu breeds of Bangladesh, Northeast and South India, and Sri Lanka and derived breeds with European blood Subgroup 8G Himalayan Hill zebu breeds Group 9 Turano-Mongolian breeds of Central and Northeast Asia, the yak and yak-cattle hybrids Subgroup 9A Turano-Mongolian breeds of Central Asian and derived breeds with European blood Subgroup 9B Breeds of Northeast China, Korea and Japan and derived breeds with European blood Subgroup 9C Yak and yak-cattle hybrids Group 10 Breeds of Central and Southern China and Southeast Asia, and bibovine cattle Subgroup 10A Central-Chinese Yellow breeds Subgroup 10B South-Chinese Yellow and Indo-Chinese zebu breeds and derived breeds Subgroup 10C Indo-Chinese, Philippine, and Indonesian breeds (influenced by banteng) and derived breeds with exotic blood Subgroup 10D Bibovine cattle and hybrids from bibovine cattle taurine or zebu cattle Group 11 North-and West-African taurine breeds Subgroup 11A North-African shorthorned breeds and derived breeds with exotic blood Subgroup 11B Breeds of Lake Chad Subgroup 11C N'Dama, derived breeds Subgroup 11D West-African shorthorned breeds Group 12 West-African zebu breeds Subgroup 12A Shorthorned Sahel zebu breeds Subgroup 12B Medium-horned West-African zebu breeds Subgroup 12C Fulani zebu breeds with long lyre-shaped horns Group 13 East-African zebu breeds Subgroup 13A Zebu and zebu-sanga breeds of Northern Sudan, Eritrea and Northern Ethiopia Subgroup 13B Small zebus of the Arabian peninsula, Somalia and the Abyssinian zebu Subgroup 13C East-African shorthorned zebu breeds Subgroup 13D Small East-African zebu including imported and derived breeds with exotic blood Subgroup 13E Zebu breeds of Madagascar and Mauritius and derived breeds with exotic blood Group 14 African sanga and sanga-zebu breeds Subgroup 14A Sanga and sanga-zebu breeds of Ethiopia and Sudan Subgroup 14B Ankole and Ankole-zebu breeds Subgroup 14C Sanga breeds of Southern Africa and derived breeds with European blood Subgroup 14D Africander and derived breeds with European blood Group 15 American breeds of Iberian descent Subgroup 15A Texas Longhorn, Gulf Coast cattle and Mexican Criollos and derived taurine breeds with exotic taurine blood Subgroup 15B Caribbean Criollo breeds and derived breeds with exotic blood Subgroup 15C Central-American Criollo breeds and derived breeds with exotic blood Subgroup 15D Criollo breeds of the Northern countries of South America and derived breeds with exotic blood Subgroup 15E Sierra Criollo breeds of the High Andes Subgroup 15F South-American Criollo breeds of Spanish-Portuguese descend and derived breeds with exotic blood Group 16 Modern cattle breeds of the Americas, Australia and New Zealand, and the genus Bison Secton 1 Dairy breeds Subgroup 16-1A Authentic American and Australian dairy and dual-purpose breeds Subgroup 16-1B Breeds descended from European dairy breeds, including zebu cross-breeds Section 2 Beef breeds Subgroup16-2A Breeds descended from British beef breeds, including zebu cross-breeds Subgroup 16-2B Continental breeds, including zebu cross-breeds Subgroup 16 2C Indo-Pakistani zebu and African zebu and sanga breeds and derived zebu breeds Section 3 genus Bison Subgroup 16-3A Wisent, bison and bison-cattle hybrids (B) Revised Group 1 Polled and Celtic breeds of North and Northwest-Europe Subgroup IA Polled breeds of Iceland, Scandinavia, the Baltics and Northern Russia Subgroup IB Horned dairy breeds of Scandinavia and Scotland Subgroup IC Polled breeds of Great Britain and derived breeds in Western Europe Subgroup ID Horned Celtic breeds of Great Britain and derived breeds in Western Europe Group 2 Lowland breeds of West, North and Eastern Europe Subgroup 2A West-European red lowland breeds and derived breeds in Eastern Europe Subgroup 2B West- and Northeast-European-pied lowland dairy breeds and derived breeds Subgroup 2C West-European red- and blue-pied dual-purpose and beef breeds and derived breeds Subgroup 2D British Shorthorn breeds and derived breeds in Central and Eastern Europe Subgroup 2E Lowland breeds of West and South England Subgroup 2F Breeds of the Channel Islands and Northwest France Subgroup 2G West-European multi breed Group 3 Short-headed and Broad-headed highland breeds of West and Central Europe Subgroup 3A Vosges and Black-Forest breeds Subgroup 3B Central-European red highland breeds Subgroup 3C Short-headed Alpine breeds and derived breeds in Eastern Europe Subgroup 3D Central European blond and yellow highland breeds Subgroup 3E West-European and Central European broad-headed red spotted mountain breeds, and derived breeds in Western, Central and Eastern Europe Subgroup 3F The Charolais and derived breeds Group 4 Grey and blond to brown breeds of France, Northern Italy, the Alps and the Balkans Subgroup 4A Breeds of Central France Subgroup 4B Grey and blond breeds of Southwest France and the Pyrenees Subgroup 4C North-Italian fawn-brown breeds Subgroup 4D Central European grey and brown mountain and derived breeds in Eastern Europe Subgroup 4E Illyrian Shorthorn breeds of the Balkans and Greece and upgraded breeds Group 5 The Breeds of Southwest-Europe Subgroup 5A Isolated breeds of the Camargue, Corsica and Sardinia and derived breeds in Northwest-Europe Subgroup 5B Feral Pyrenean and Cantabrian breeds (Tronco (Castaa) Cntabrio) Subgroup 5C Galician, Balearic and Canarian Blond breeds (Rojo convexo (turdetano)) Subgroup 5D NorthWest-Iberian Brown breeds (Morenas del Noroeste and Castaa concavo) Subgroup 5E Iberian Black breeds (Negro Iberica) Subgroup 5F Central and South-Iberian Red breeds (Andaluza Rojo convexo) Subgroup 5G Southeast-Iberian breeds (Castao ultraconvexo) Group 6 Podolian breeds of Italy and Eastern Europe Subgroup 6A Large White breeds of Italy Subgroup 6B Podolian breeds of Italy and Istria Subgroup 6C Podolian grey steppe breeds of Eastern Europe Subgroup 6D Podolian-Illyrian breeds of the Balkan Countries and Anatolia Group 7 Shorthorned breeds of the Caucasus, Anatolia, the Levant and Egypt Subgroup 7A Humpless breeds of the Caucasus and Southwest Asia and derived breeds with exotic influence Subgroup 7B Damascus type in West Asia and Egypt and derived breeds with exotic influence Group 8 Indo-Pakistani type zebu breeds Subgroup 8A Zebu and zebod breeds of the Central Asia, Iran, Afghanistan, Northwest Pakistan and derived breeds with exotic influence Subgroup 8B Zebu breeds with convex forehead and derived breeds with taurine influence Subgroup 8C Shorthorned grey-white zebu breeds Subgroup 8D: Zebu breeds with lyre-shaped horns and derived breeds with taurine influence Subgroup 8E: Mysore zebu breeds Subgroup 8F: Small zebu breeds of Bangladesh, Northeast and South India, and Sri Lanka and derived breeds with taurine influence Subgroup 8G: Himalayan Hill zebu breeds and hybrids Group 9 Turano-Mongolian breeds of Central and Northeast Asia, yak and yak-cattle hybrids Subgroup 9A: Central-Asian Turano-Mongolian and derived breeds with European influence Subgroup 9B: Breeds of Northeast China, Korea and Japan and derived breeds with European influence Subgroup 9C: Yak and yak-cattle hybrids Group 10 Breeds of Central and Southern China and Southeast Asia, and Bibovine cattle Subgroup 10A: Central-Chinese Yellow breeds (Huanghuai Group) Subgroup 10B South-Chinese Yellow breeds (Changzhu Group), Indo-Chinese zebu breeds and derived breeds with zebu influence Subgroup 10C Indo-Chinese, Philippine, and Indonesian breeds (influenced by banteng) and derived breeds with exotic influence Subgroup 10D Bibovine cattle and hybrids Group 11 North-and West-African taurine breeds Subgroup 11A North-African Shorthorn breeds and derived breeds with exotic influence Subgroup 11B Lake Chad breeds and populations Subgroup 11C N'Dama, derived taurindicus populations and breeds with exotic influence Subgroup 11D West-African shorthorned breeds, derived taurindicus breeds and populations Group 12 West-African zebu breeds Subgroup 12A Shorthorned Sahel zebu breeds Subgroup 12B Medium-horned West-African zebu breeds and derived taurindicus breeds Subgroup 12C Fulani zebu breeds with long, lyre-shaped horns Group 13 East-African zebu breeds Subgroup 13A Large zebu breeds of Northeast Africa Subgroup 13B Small zebus of the Arabian Peninsula and the Horn of Africa Subgroup 13C East-African shorthorned zebu breeds Subgroup 13D Small East-African zebu and derived breeds with exotic influence Subgroup 13E Zebus of Madagascar, Mauritius and Ocean Islands and derived breeds with exotic influence Group 14 African sanga and zenga breeds Subgroup 14A Sanga and zenga breeds of Northeast Africa Subgroup 14B Central-African Ankole sanga and zenga breeds Subgroup 14C Sanga and zenga breeds of southern Africa and derived breeds with exotic influence Subgroup 14D European, American and Australian purebred populations and derived breeds in Southern Africa Group 15 American breeds of Iberian descent Subgroup 15A Texas Longhorn, Gulf Coast cattle, Mexican Criollos and derived taurine breeds Subgroup 15B Caribbean Criollo breeds and derived breeds with exotic influence Subgroup 15C Central-American Criollo and derived breeds with exotic influence Subgroup 15D Criollo of the northern part of South America and derived breeds with exotic influence Subgroup 15E Sierra Criollo breeds of the High Andes and derived taurine breeds Subgroup 15F South-American Criollo of Iberian descent and derived breeds with exotic influence Group 16 Modern cattle breeds of the Western Hemisphere (Americas, Australia and New Zealand) and the genus Bison Subgroup 16-1A Authentic populations and breeds of the Western Hemisphere the originals Subgroup 16-1B Western hemisphere dairy and dual purpose breeds derived from European breeds Subgroup 16-1Bb Western hemisphere taurindicus dairy and dual-purpose breeds Subgroup 16-2A Western hemisphere beef breeds descended from British beef breeds Subgroup 16-2Ab Western hemisphere taurindicus beef breeds descending from British breeds Subgroup 16-2B Western hemisphere beef breeds, descending continental and Japanese breeds Subgroup 16-2Bb Western hemisphere taurindicus beef breeds descending from continental breeds Subgroup 16-2C Western hemisphere humped breeds of Indo-Pakistani descent, African sanga and zebu Subgroup 16-3A American hybrids, genus Bison and yak Table S34. Source of microsatellite data used in Figure S2. Names and addresses of laboratories are mentioned in the cited publications. N, number of animals; INRA, Institut des Recherches Scientifiques Agronomiques.VHL, Van Haeringen Laboratory (Wageningen). BreedCountryNLaboratoryReferenceAberdeen AngusGreat-Britain50RoslinADDIN RW.CITE{{122 Wiener,P. 2004; 219 European Cattle Genetic Diversity Consortium 2006}}[39,40]AgersoeDenmark41CopenhagueADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006; 225 Withen,K.B. 2011}}[40,41]Alistana-SanabresaSpain50MadridADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Anatolian BlackTurkey49MunichADDIN RW.CITE{{127 Medugorac,I. 2009}}[42]Angeln: Sleswig-Holstein (old and modern), Hessen, Rhine-WestphaliaGermany189Giessen, GottingenADDIN RW.CITE{{142 Bremond, J. 2002; 219 European Cattle Genetic Diversity Consortium 2006}}[40,43]Asturiana MountanaSpain50MadridADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Asturiana VallesSpain50MadridADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]AubracFrance50INRAADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]ADDIN RW.CITE{{233 Baumung, R. 2007}}[44]Avilea Negro IbericaSpain50Madrid/BarcelonaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]AyrshireGreat-Britain48RoslinADDIN RW.CITE{{122 Wiener,P. 2004; 219 European Cattle Genetic Diversity Consortium 2006}}[39,40]BazadaiseFrance47INRAADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Belgian BlueBelgium50Malle/ViterboADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Berrenda (black- and red-pied)Spain100CordobaADDIN RW.CITE{{232 Azor, P. 2007}}[45]BestuzhevRussia66JokioinenADDIN RW.CITE{{131 Li,M.H. 2007/a;}}[46]BetizoaSpain23ZaragozzaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006; 129 Martin-Burriel,I. 2007}}[40,47]Blonde d'AquitaineFrance50INRAADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Bohemian RedCzech Reupblic25INRA/GiessenADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Bretonne Pie-NoireFrance31INRAADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]British HolsteinBritain50RoslinADDIN RW.CITE{{122 Wiener,P. 2004; 219 European Cattle Genetic Diversity Consortium 2006}}[39,40]Bruna PirineusSpain50Madrid/BarcelonaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Busha: Kosovo, Macedonia, AlbaniaBusa146Jokioinen, MunichADDIN RW.CITE{{127 Medugorac,I. 2009/a;}}[42]Byelorussian RedByelorussia20JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]CabanninaItaly26INRA/PiacenzaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]CardenaSpain14CordobaADDIN RW.CITE{{232 Azor, P. 2007}}[45]Carinthian BlondAustria60ViennaADDIN RW.CITE{{233 Baumung, R. 2007}}[44] Casta NavarraSpain50ZaragozzaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]CharolaisFrance50INRAADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]ChianinaItaly36INRA/PiacenzaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]CinesaraItaly30Catania/VHLADDIN RW.CITE{{237 Marletta, D. 2009}}[49]Danish RedDenmark32CopenhagueADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006; 225 Withen,K.B. 2011}}[40,41]DexterGreat-Britain46RoslinADDIN RW.CITE{{122 Wiener,P. 2004; 219 European Cattle Genetic Diversity Consortium 2006}}[39,40]Dutch BeltedNetherlands24INRA/UtrechtADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Dutch FriesianNetherlands34INRA/UtrechtADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Eastern FinncattleFinland31JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]Ennstal-BergscheckAustria41ViennaADDIN RW.CITE{{233 Baumung, R. 2007}}[44]EringerSwitserland50BerneADDIN RW.CITE{{143 Schmid, M. 1999; 219 European Cattle Genetic Diversity Consortium 2006}}[40,50]EvolenardSwitserland15BerneADDIN RW.CITE{{143 Schmid, M. 1999; 219 European Cattle Genetic Diversity Consortium 2006}}[40,50]Finnish AyrshireFinland43JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]GasconneFrance50INRAADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]German Brown (Bavaria, Wqrttemberg)Germany50GiessenADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]German Brown OriginalGermany25GiessenADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]German Original Black-Pied, Western reserveGermany20GiessenADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]German SimmentalGermany50GiessenADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]German YellowGermany50GiessenADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]German, Polish Black-PiedGermany, Poland50Giessen, KrakowaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Grey Gacko BushaBosnia41Gottingen/HannoverADDIN RW.CITE{{127 Medugorac,I. 2009/a;}}[42]Grigia AlpinaItaly28MilanoADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Groningen WhiteheadedNetherlands25INRA/UtrechtADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]GuernseyGreat-Britain50RoslinADDIN RW.CITE{{122 Wiener,P. 2004; 219 European Cattle Genetic Diversity Consortium 2006}}[39,40]HerefordGreat-Britain48RoslinADDIN RW.CITE{{122 Wiener,P. 2004; 219 European Cattle Genetic Diversity Consortium 2006}}[39,40]HinterwlderGermany30GiessenADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Holstein-FriesianSwitserland, Russia80Jokioinen, BerneADDIN RW.CITE{{143 Schmid, M. 1999; 219 European Cattle Genetic Diversity Consortium 2006}}[40,50]Hungarian GreyHungary60Vienna ADDIN RW.CITE{{233 Baumung, R. 2007}}[44]IstobenRussia48JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]IstrianCroatia45INRA/GiessenADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]JerseyGreat-Britain47RoslinADDIN RW.CITE{{122 Wiener,P. 2004; 219 European Cattle Genetic Diversity Consortium 2006}}[39,40]Jutland (from 1950)Denmark44INRA/TjeleADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]KalmykRussia28JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]Kazakh WhiteheadedRussia40JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]KholmogorRussia42JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]LimousinFrance50INRAADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]LonghornEngland44RoslinADDIN RW.CITE{{235 Wiener, P. , Roslin 2010}}[51]Maine-AnjouFrance49INRAADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]MallorquinaSpain28ZaragozzaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006; 129 Martin-Burriel,I. 2007}}[40,47]MenorquinaSpain50ZaragozzaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006; 129 Martin-Burriel,I. 2007}}[40,47]ModicanaItaly37Catania/VHLADDIN RW.CITE{{237 Marletta, D. 2009}}[49]MonchinaSpain50ZaragozzaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006; 129 Martin-Burriel,I. 2007}}[40,47]MontbliardeFrance31INRAADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]MoruchaSpain50Madrid/BarcelonaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]MurbondnerAustria47MunichADDIN RW.CITE{{127 Medugorac,I. 2009/a;}}[42]Murnau-WerdenfelserGermany52MunichADDIN RW.CITE{{127 Medugorac,I. 2009/a;}}[42]N'DamaAfrica30INRAADDIN RW.CITE{{124 Moazami-Goudarzi, K. 2001; 219 European Cattle Genetic Diversity Consortium 2006}}[40,52]NormandeFrance50INRAADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Northern FinncattleFinland26JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]PajunaSpain50CordobaADDIN RW.CITE{{232 Azor, P. 2007}}[45]ParthenaiseFrance37RoslinADDIN RW.CITE{{235 Wiener, P. , Roslin 2010}}[51]Pechora (type of Kholmogory)Russia31JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]PiemonteseItaly49INRA/PiacenzaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]PinzgauerAustria44Dublin/UtrechtADDIN RW.CITE{{233 Baumung, R. 2007}}[44]PirenaicaSpain50Madrid/BarcelonaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]PodolicaItaly50CampobassoADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Polish RedPoland48INRA/GiessenADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]PustertalerAustria44ViennaADDIN RW.CITE{{233 Baumung, R. 2007}}[44]Red Holstein dairy typeGermany25GiessenADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Red-Pied: Meuse-Rhine-Yssel, Red Holstein dual-purposeNetherlands, Germany,139Utrecht, GiessenADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Red SteppeUkraine36JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]RendenaItaly34MilanoADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]RetintaSpain50Madrid/BarcelonaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]RomagnolaItaly32INRA/PiacenzaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Rubia GallegaSpain50Madrid/BarcelonaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Russian Black-PiedRussia30JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]SalersFrance50INRAADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]SayaguesaSpain48MadridADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]ShorthornGreat-Britain43RoslinADDIN RW.CITE{{235 Wiener, P. , Roslin 2010}}[51]Serrana de TeruelSpain44ZaragozzaADDIN RW.CITE{{129 Martin-Burriel,I. 2007}}[47]Simmental/Pezzata Rossa ItalianaGermany, Austria, Italy139Giessen, Vienna, INRA,ADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006; 233 Baumung, R. 2007}}[40,44]South-DevonGreat-Britain44RoslinADDIN RW.CITE{{235 Wiener, P. , Roslin 2010}}[51]SuksunRussia40JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]Swedish Red-PolledSweden32INRA/UppsalaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Swiss BrownSwitserland, Austria93Berne,ViennaADDIN RW.CITE{{143 Schmid, M. 1999; 219 European Cattle Genetic Diversity Consortium 2006; 233 Baumung, R. 2007}}[40,44,50], Swiss SimmentalSwitserland50BerneADDIN RW.CITE{{143 Schmid, M. 1999; 219 European Cattle Genetic Diversity Consortium 2006}}[40,50]TarentaiseFrance43TarentaiseADDIN RW.CITE{{127 Medugorac,I. 2009/a;}}[42]Toro de Lidia (Fighting cattle)Spain44ZaragozzaADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]TudancaSpain50MadridADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006}}[40]Tyrolean Grey CattleAustria48MunichADDIN RW.CITE{{127 Medugorac,I. 2009/a;}}[42]Ukrainian GreyUkrain30JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]Ukrainian WhiteheadedUkrain10JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]Waldviertler BlondAustria60ViennaADDIN RW.CITE{{233 Baumung, R. 2007}}[44]Western FinncattleFinland39JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]Yakutian cattleRussia60JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]YaroslavlRussia44JokioinenADDIN RW.CITE{{131 Li,M.H. 2007; 130 Li,M.H. 2010}}[46,48]Zebu PeulAfrica100INRA/MilanoADDIN RW.CITE{{124 Moazami-Goudarzi, K. 2001; 219 European Cattle Genetic Diversity Consortium 2006}}[40,52]. Figure S2. NeighborNet graph of Reynolds DR genetic distances of 103 European breeds analyzed with 30 FAO-recommended microsatellites. Genetic distances were calculated by the program Microsat (hpgl.stanford.edu/projects/microsat/). The network was constructed with the program Splitstree ADDIN RW.CITE{{224 Huson,D.H. 1998}}[53]. Except for the NW Intermediate breeds, the breed groups are also revealed by model-based clustering by the Structure program ADDIN RW.CITE{{223 Pritchard,J.K. 2000}}[54]. The coloring of the border indicate the four major groups of European cattle, which also are in agreement with model-based clustering: blue, North-European; violet, Central-European; ochre, Iberian; grey, Podolian. Exported (Finnish Ayrshire), recently crossbred (Asturian Valley, Istoben), extremely inbred (Mallorquina, Menorquina) as well as the French Gasconne (spuriously associated with British breeds) and Central-Eastern (Pinzgauer and Pustertaler, converging with Iberian breeds) breeds have not been plotted.  Table S35. Classification of Earasian taurine breeds based on molecular genetic analysis (ADDIN RW.CITE{{7 Edwards,Ceiridwen J. 2011}}[55]; Figures 9 and S2; ADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006; 225 Withen,K.B. 2011; 127 Medugorac,I. 2009; 143 Schmid, M. 1999; 142 Bremond, J. 2002; 238 Lenstra, J.A., www.nas.boku.ac.at/fileadmin/_/H93/H932-NUWI/Kurse-Workshops-Tagungen/ESF-Workshop/Session1/Lenstra.pdf 2008; 7 Edwards,Ceiridwen J. 2011; 129 Martin-Burriel,I. 2007; 129 Martin-Burriel,I. 2007; 130 Li,M.H. 2010; 131 Li,M.H. 2007; 220 Decker,J.E. 2009; 19 Gautier,M. 2010}}[35,40-43,46,47,47,48,50,55-57]). Breed clustering was inferred from model-based clustering and/or genetic distances between breeds. This may assign crossbred breeds either to a specific group or to a rest-group of unassigned breeds, depending on both the degree of crossbreeding and on the molecular dataset. I North-European breeds I.1 British British dairy and beef breeds, including also the Channel-Island breed (Jersey, Guernsey), but not the Shorthorn. Jersey tends to be different from the other breeds in this group and may have apparent affinity to Podolian or Alpine grey breeds. I.2 Nordic Authentic Norwegian, Swedish, Finnish and Baltic breeds, including both polled as longhorned (Dle, Telemark) breeds. I.3 Nordic Ayrshire Imported Finnish Ayrshire and Ayrshire crossbreds: Norwegian Red, Swedish Red-and-White, Ringamla, Vne. I.4 Lowland-Pied Black- and red-pied dairy cattle originating from the north-western lowlands of the European lowlands. Also includes the solid Red Flemish. I.5 Baltic Red Solid red dairy cattle from the Baltic coasts and the German Highland. Also includes the Russian Suksun, Byelorussian Red and Ukrainian Red Steppe. I.6 Northwest Intermediate Cattle from north-western Europe that are not closely related to each other, but are influenced to different degrees by surrounding breeds: Shorthorn, Maine-Anjou (similar to Shorthorn), Bretonne-Pie Noir, Normande, Parthenaise (close to Southern-French breeds), Vosges, Charolais. Charolais clustered with South-French breeds and Vosges with Central-Western breeds in a 50K SNP analysis ADDIN RW.CITE{{220 Decker,J.E. 2009; 19 Gautier,M. 2010}}[35,57]. 1.7 Eastern crossbred Russian breeds heavily influenced by Western breeds: Istoben (influenced by Lowland Pied), Kazakh Whiteheaded (influenced by Hereford), Ukranian Whitehead (influenced by Groningen Whiteheaded), Bestushev (influenced by several breeds). I.8 Eastern Russian and Siberian breeds: Kholmogory, Pechora (both influenced by I.1), Kalmyk, Yaroslav, Yakut. II Central-European cattle II.1 Central-Western Includes four subtypes (II.1.1-3) and Hinterwald; SNP data suggest inclusion of Charolais and Vosges. II.1.1 Central Spotted Central-European spotted dairy cattle with Simmentaler as prototype breed from which several other breeds have been derived. II.1.2 Central Blond Carinthian and Waldviertel Blond, genetically close the Central Spotted. II.1.3 West-Alpine French-Alpine, Swiss Valais (Wallis) and Italian Valdostana breeds, Austrian Tux-Zillertaler. II.1.4 Central Yellow GermanYellow breeds, Murbodner, Portuguese Minhota. II.2 South-French Southern French beef breeds and the Spanish Pyrenaica, which is also influenced by Iberian cattle. II.3 Central Brown Swiss Brown dairy cattle and derived breeds in Germany, Italy and Spain; including Murnau-Wersenfelder. II.3.1 Spanish Brown Spanish breeds derived from Central Brown: Bruna Pirieus, Parda Montana and Serrana de Teruel. II.4 Central Grey Tyrolean Grey, Grigia Alpina. II.5 Central-Eastern Pinzgauer, Pustertaler, Cika. III Iberian cattle Authentic and morphologically diverse Spanish and Portuguese breeds. Relationship with the Mallorquina and Menorquina unclear by the high degree of inbreeding of both Balearic breeds. The feral Betizu is genetically between the Iberian and Southern-French cattle. Contains regional clusters of breeds: III.1.1 Cantabrian Tudanca, Monchina, Betizu. III 1.2 Andalusian Andalusian breeds: Berrenda, Cardena, Marismea, Mostrenca, Pajuna, Fighting cattle (Lidia, Brava, Casta Navarra) III 1.3 Iberian Black Avilea, Morucha, Negra Andaluza, Preta. III 1.4 Morenas Alistana, Barrrosa, Cachena, Frieiresa, Caldelana, Limiana, Marinhoa, Maronesa, Mirandesa, Vianesa. III 1.5 South-Portuguese Red Alentejana, Garvonesa, Mertolenga. IV Podolian cattle Steppe cattle, presumed to originate from the Podolia region. Contains also Ukranian Grey, Turkish Grey and Chianina. V South-Eastern European and SouthWest-Asian taurine Authentic taurine cattle smaller and less developed than most European breeds: Busha, Anatolian and Caucasian cattle. APPENDIX 1. LEXICON OF LATIN DESIGNATIONS The term Bos primigenius was in 1973 proposed for both aurochs and domestic cattle ADDIN RW.CITE{{115 Herre, W. 1973/a;}}[58]. However, the International Commission on Zoological Nomenclature published a Code list with names for mammals ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] in which the name Bos primigenius ADDIN RW.CITE{{145 Bojanus, L.H. 1827;}}[60] was converted into Bos taurus, the name given by Linnaeus ADDIN RW.CITE{{117 Linnus, C. 1758/a;}}[61] in order to consequently apply the first Latin name given to a species. Van Vuure ADDIN RW.CITE{{118 Vuure, C. van 2001/a;}}[62] suggested to use Bos taurus primigenius for the European aurochs and Bos taurus domesticus for (European) domestic cattle ADDIN RW.CITE{{119 Erxleben, J.C.P. 1777}}[63]. However, it is more common, although formally incorrect, to use separate Latin designations for domestic animals. This was defended by Clutton-Brock ADDIN RW.CITE{{146 Clutton-Brock, J. 2001/a;}}[64]: I believe that animals bred under domestication evolve into new species, as a result of reproductive isolation from their wild progenitors combined with natural and artificial selection in association with human societies. Gentry et al. ADDIN RW.CITE{{147 Gentry, A. 1996/a;}}[65] also pleaded for differentiation in names for wild and domestic cattle and proposed to accept Bos primigenius for all aurochs. Today the Latin names Bos primigenius and Bos namadicus are commonly accepted for the two main aurochs species. The names Bos taurus (non-humped) and Bos indicus (true zebus) are then used for the domestic descendants of respectively Bos primigenius and Bos namadicus. Bos taurindicus indicates the several taurine and indicine intermediates ADDIN RW.CITE{{1 Ajmone-Marsan,P. 2010}}[66]. One step further, and certainly incorrect, is the latinizing of breed names, such as Bos taurus taurus ankole, B.t.t. ayrshire, etc. This is, for instance, done on the website Mammals Planet (www.planet-mammiferes.org/drupal/en/node/20 and may suggest prehistoric mutants or stem forms that never have existed as such. Many Latin terms were already introduced by Fitzinger ADDIN RW.CITE{{89 Fitzinger, L.J. 1860/a;}}[67], Werner ADDIN RW.CITE{{37 Werner, A.H. 1892/a;}}[2], Sanson ADDIN RW.CITE{{69 Sanson, A. 1884/a;}}[6] and Keller ADDIN RW.CITE{{41 Keller, C. 1905/a;121 Keller, C. 1902/a;}}[3,68] and were used in 1930 in a review of the descent of cattle ADDIN RW.CITE{{252 Szalay, A.B. 1930}}[69]. Since several of these keep turning up in the scientific literature, we have compiled a list. Several terms are even on the International Commission on Zoological Nomenclature Code list ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59]. Latin designations commonly used are underlined. For literature references that are part of the species designation in the Linnaean format (e.g., Bos taurus Linnaeus, 1758) bibliographic details are not always available. Bos abyssinicus Oken 1838 ADDIN RW.CITE{{252 Szalay, A.B. 1930}}[69] earliest name B. domesticus t. abyssinicus, later as B. t. abyssinicus (q.v.). Bos acutifrons (Lydekker, 1877) ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] for an early Pliocene form, after a two million years old cranium with extreme long horns excavated in the Siwalik Hills in northern India. This so-called Siwalik Ox is presumed to be the ancestor of all species of the group Bovina (common cattle, bibovine cattle, yak and bison). Lydekker thougth it not improbable that a skull from the same deposites described by him as Bos planifrons indicated the female of the same species. Bos africanus Kerr, 1792 for the Sanga, as in Brehms Tierleben ADDIN RW.CITE{{150 Brehm, Alfred Edmund 1893/a;}}[71]. Bos akeratos Arenander, 1989 ADDIN RW.CITE{{66 Arenander, E.O 1898/a;}}[72] for remains of of a polled cattle excavated in Scandinavia, supposed to represent the original European aurochs and to be the ancestor horned cattle ADDIN RW.CITE{{66 Arenander, E.O 1898/a;}}[72]. Bos alpinus Sanson, 1878 ADDIN RW.CITE{{166 Sanson, A. 1878}}[73] for Alpine cattle, renamed B. taurus alpinus ADDIN RW.CITE{{167 Sanson, A. 1893}}[74]. Bos alpium Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67] for Alpine [Braunvieh] type cattle conprising 18 breeds found in Switzerland, Tyrol, Styria and Bohemia, renamed B.taurus alpium. Bos aquitanicus Sanson, 1878 ADDIN RW.CITE{{69 Sanson, A. 1884}}[6] presumed a local aurochs and forefather for blond cattle from southern France and northern Iberian Peninsula. Later regarded as domestic form: B. taurus aquitanicus. Bos asiaticus Sanson, 1878, renamed B. taurus asiaticus ADDIN RW.CITE{{167 Sanson, A. 1893}}[74]. Bos avernensis Sanson, 1878. renamed B. taurus avernensis ADDIN RW.CITE{{167 Sanson, A. 1893}}[74]. Bos balticus Stegmann von Pritzwald, 1924, renamed B.t. balticus ADDIN RW.CITE{{168 Stegmann von Pritzwald, F.P. 1924}}[75], small aurochs-type crania from Danzig, Knigsberg and Copenhague ADDIN RW.CITE{{249 Nehring, A. 1889; 244 Hilzheimer, M. 1925}}[76,77] [Hilzheimer 1925] and identified as the skull of an aurochs cow ADDIN RW.CITE{{65 Leithner, O.F 1927/a;}}[78]. Bos bojanus Grard, 1871 ADDIN RW.CITE{{252 Szalay, A.B. 1930}}[69] other name for B. p. Bojanus. Bos bombifrons Nesti ADDIN RW.CITE{{34 Rtimeyer, L. 1868}}[79] after a Plioncene cranium, by Falconer named Bos (Amphibos) etruscus, which was later shortened to Bos etruscus. Bos brachyceroides Pohling, 1911 ADDIN RW.CITE{{151 Pohlig, H. 1911}}[80] after a skull fragment with short horns excavated in Asti, Italy, dated early Pliocene or the oldest Pleistocene layer, considered a miniature aurochs. One year later the name was changed into Bos mastodontis by Pohlig and later denoted as B.t. brachyceroides. Bos brachyceros Owen, 1830 ADDIN RW.CITE{{188 Owen, R. 1846; 32 Epstein, H. 1971}}[81,82] holotype named after its short horns, first described in a catalogue of the museum of the College of Surgeons in 1830. In 1846 he replaced the name by Bos longifrons (long-head cattle). Rtimeyer ADDIN RW.CITE{{34 Rtimeyer, L. 1868; 32 Epstein, H. 1971}}[79,82] retained the first name for this type of prehistoric cattle, which had been first excavated at Neolithic sites in the Swiss Lake district. He considered this Torfrind (village cattle) as being introduced from Asia, and an older, more important type than primigenius type domestic cattle. For a long time Bos brachyceros was (incorrectly) presumed to be the source of modern shorthorned cattle and is still used to indicate shorthorned cattle of all kinds. Hilzheimer ADDIN RW.CITE{{67 Hilzheimer, M. 1926/a;}}[83] correctly assumed that these cattle had evolved by domestication. Bos brachyceros arnei Amschler, 1939 ADDIN RW.CITE{{187 Amschler, J.W. 1939; 32 Epstein, H. 1971}}[82,84] for a skull fragment excavated at Shah Tep, North Iran and incorrectly regarded as wild ancestral species of all shorthorn cattle of Asia, Africa and Europe, as well as the orthoceros cattle of the Kalmyks and Mongols. Renamed B.t. arnei. Bos brachyceros europaeus Adametz, 1898 ADDIN RW.CITE{{153 Adametz, L.: 1898}}[85]. Initially Adametz did not accept the brachyceros as an introduced Asian form, but as a separate European wild form next to the B.primigenius Bojanus, and incorrectly presumed to be the ancestor of many European cattle breeds. Bos brachyceros ibericus Sanson, 1878 ADDIN RW.CITE{{166 Sanson, A. 1878}}[73] replacing the designations B. ibericus and B. curvidens, for wild forms, found in Algeria and Tunisia ADDIN RW.CITE{{169 Duerst, J.U. 1931}}[86]. Bos brachycerous Cividini & Kume, 2008ADDIN RW.CITE{{154 Cividini, A. 2008/a;}}[87] a putativesmall wild progenitor of all shorthorned Busha type cattle of the Balkans, a rendered out of date theory. Bos brachycerous palustris according to Kume et al. (2008) ADDIN RW.CITE{{155 Kume, K. 2008/a;}}[88] the designation of Rtimeyer for the remains of Neolithic village cattle excavated from the Swiss lake sites. Bos braquiceros Aparicio, 1960 ADDIN RW.CITE{{156 Aparicio, G. 1960/a;}}[89] for brachyceros, in other Spanish literature also written as brachycerus and sometimes with the extension Europeo or Africano. Bos braquicerus africanus Sanchez Belda, 1984 ADDIN RW.CITE{{85 Sanchez Belda, Antonio 1984}}[14] considered to trace to the B.t. ibricus and putative predecessor of the Sayaguesa. Bos caesaris Keferstein, 1834 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] a classic name for the aurochs because of the description of the aurochs by Julius Caesar in De Bello Gallico. Bos curvidens Pomel, 1893 ADDIN RW.CITE{{159 Pomel, A.. 1893}}[90], for remains from Algerian palaeolithic sites for a smaller than Bos primigenius and considered identical to B. opisthonomus and B. p. mauritanicus ADDIN RW.CITE{{252 Szalay, A.B. 1930}}[69]. Bos Dante Link, 1793 for cranium excavated by Link ADDIN RW.CITE{{149 Lydekker, R. 1898; 240 Gray, J.E. 1852}}[70,91]. Rtimeyer ADDIN RW.CITE{{34 Rtimeyer, L. 1868/a;}}[79] stated that the skull had belonged to a polled zebu, named it B. indicus var. Dante, also found as B.taurus dante. Drst ADDIN RW.CITE{{158 Drst, J. Ulrich 1899/a;}}[92] considered it a macroceros type. Bos desertorum hispanico Aparicio, 1960 ADDIN RW.CITE{{156 Aparicio, G. 1960}}[89] after Fitzingers B.t. desertorum, presumed to descend from the African B.p. Hahni and B.t. mauritanicus, a hypothetical Iberian aurochs variant, characterized by high-lyred horns and a light coat and connected with breeds as Barros, Cachena, Blanca Cacerea, and Crdena Andaluza breed. Bos domesticus Johnston, 1651 ADDIN RW.CITE{{38 Duerst, J.U. 1931}}[93]. The Bos species are distinguished as a subclass of horned ruminants for the first time in Johnstons systematic. He mentioned domestic cattle: Bos domesticus besides the wild cattle forms: Urus, Bison, Bonasus, Buffelus and Strepsiceros. Bos ecornis Wagner, 1837 ADDIN RW.CITE{{157 Wagner, A.J. 1837}}[94] probably for polled cattle after the French corner. Bos elatus Pomel, 1853 ADDIN RW.CITE{{149 Lydekker, R. 1898; 159 Pomel, A.. 1893}}[70,90] fossilized skull fragment from Tertiary deposits of Val dArno, Italy, typified as small, possible female form of Bos etruscus (on display in the Museum National dHistoire Naturelle, Paris). Bos etruscus Falconer, 1868 ADDIN RW.CITE{{170 Falconer, Hugh 1868}}[95] from Bos (Amphibos) etruscus ADDIN RW.CITE{{34 Rtimeyer, L. 1868}}[79], the Etruscan Ox for an entire fossilised Late-Pliocene skull excavated in Val dArno, a bovid type considered to have been distributed over Italy and southern France. Alternative names are Bos elatus (Pomel) and Bos stenometopon (Sismonda). Rtimeyer ADDIN RW.CITE{{34 Rtimeyer, L. 1868/a;}}[79] considered it akin to Bos namadicus and the predecessor of gaur, banteng, yak and zebu, and used the name Bos (Bibos) etruscus ADDIN RW.CITE{{34 Rtimeyer, L. 1868}}[79]. However, Lydekker ADDIN RW.CITE{{149 Lydekker, R. 1898/a;}}[70] classified it in the Leptobovine group as subgenus Leptobos ADDIN RW.CITE{{34 Rtimeyer, L. 1868}}[79], a form which became extinct during Early Pleistocene without descendants. Bos europaeus (brachyceros) Adametz, 1927 ADDIN RW.CITE{{28 Adametz, Leopold 1926}}[96] for European shorthorned cattle, after Kerrs Bos europaeus. Bos falconeri Lydekker, 1898 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] Falconers Ox, remains from the Siwalik region, India, akin to the Siwalik Ox, but also to the Etruscan Ox, earlier named Leptobos falconeri ADDIN RW.CITE{{34 Rtimeyer, L. 1868}}[79] and the Pliocene Bos fraseri from the Narbada valley, which was presumed to be closer to the banteng. Bos ferus sylvestris Castello, 1721 ADDIN RW.CITE{{252 Szalay, A.B. 1930}}[69] for aurochs. Bos fossilis Keferstein, 1834 ADDIN RW.CITE{{149 Lydekker, R. 1898; 241 Keferstein 1843; 252 Szalay, A.B. 1930}}[69,70,97] for aurochs remains also named B. t. fossilis (q.v.). Bos frontosus Nilsson, 1849 ADDIN RW.CITE{{149 Lydekker, R. 1898; 77 Nilsson, S. 1849}}[70,98] denoting a broad-headed cranium excavated in Scandinavia, considered as separate aurochs variant that was the founder of the small, often polled cattle of Norway. In 1868 it was identified as the remains of a domestic specimen, and thus renamed Bos taurus frontosus ADDIN RW.CITE{{152 Rtimeyer, L. 1861}}[99]. [Rtimeyer 1861] Bos Galla Salt, 1814 ADDIN RW.CITE{{158 Drst, J.U. 1899; 240 Gray, J.E. 1852}}[91,92], turned out to be a sanga cranium from Abyssinia (Ethiopia), renamed B. taurus galla and by Drst ADDIN RW.CITE{{158 Drst, J. Ulrich 1899}}[92] identified as a macroceros type. Bos giganteus Croizet, 1828 ADDIN RW.CITE{{169 Duerst, J.U. 1931; 149 Lydekker, R. 1898}}[70,86] also Bos giganeus ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70]for a Pliocene aurochs. Bos hybridus Fitzinger 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67] for crossbred cattle. Bos ibericus Sanson, 1878 ADDIN RW.CITE{{166 Sanson, A. 1878}}[73] a so-called nature type, without specifying it as being wild or tame cattle, renamed B.taurus ibericus by Sanson (1893) ADDIN RW.CITE{{167 Sanson, A. 1893}}[74], by Von Leithner ADDIN RW.CITE{{65 Leithner, O.F 1927/a;}}[78] identified as identical to Bos opistonomus and not differing from Bos primigenius. Bos ibricus Pomel, 1893 ADDIN RW.CITE{{159 Pomel, A.. 1893}}[90] considered an aurochs in Portuguese and Spanish literature often mentioned as ancestor of Iberian breeds. Bos indicus Linnus, 1758 ADDIN RW.CITE{{117 Linnus, C. 1758; 149 Lydekker, R. 1898; 116 Wilson, D.E. 1993}}[59,61,70] the taxon including all domestic humped (true) zebu or indicine cattle. Bos indicus var. Dante Rtimeyer, 1867 ADDIN RW.CITE{{34 Rtimeyer, L. 1868/a;}}[79] for the Bos dante cranium identified as from a domestic polled zebu skull. Bos indicus Galla Salt, 1814 ADDIN RW.CITE{{171 Anonymous; 116 Wilson, D.E. 1993}}[59,100] former Bos Galla, also asB. taurus galla, for theN.W. Ethiopian Raya-Azeb breed, in the past named Galla breed. Bos indicus namadicus Falconer, 1859 ADDIN RW.CITE{{170 Falconer, H. 1868; 34 Rtimeyer, L. 1868; 149 Lydekker, R. 1898}}[70,79,95] Falconers term for domestic zebu, initially indicated as B. namadicus. Bos indicus nanus ADDIN RW.CITE{{171 Anonymous}}[100] dwarf zebu, also B.t. indicus minor. Bos indicus planus Sasaki, 1934 ADDIN RW.CITE{{172 Sasaki, K. 1934}}[101] the Formosa zebu, a subtype of South-Korean cattle considered a strain descending the Indian zebu. Bos indicus var. Pusio Rtimeyer, 1867 ADDIN RW.CITE{{34 Rtimeyer, L. 1868/a;}}[79] for the Bos Pusio cranium identified as from a domestic longhorned zebu. Bos intermedius Serres, 1829 ADDIN RW.CITE{{252 Szalay, A.B. 1930}}[69] for a fossil excavated in Lunel-Viel [probably Lunville in Meurthe-et-Moselle]. In the table Palaeontologische Reihe as well as Morphologische Reihe Rtimeyer ADDIN RW.CITE{{34 Rtimeyer, L. 1868/a;}}[79] placed Bos intermedius next to the diluvial Bos primigenius with a cautionary note. Bos italiae Pohlig, 1911 ADDIN RW.CITE{{151 Pohlig, H. 1911; 252 Szalay, A.B. 1930}}[69,80] also B.p. Italiae, skull excavated in Lombardia ADDIN RW.CITE{{151 Pohlig, H. 1911}}[80] belonging to a diluvial aurochs ADDIN RW.CITE{{65 Leithner, O.F 1927/a;}}[78]; term used occasionally for indicating a local aurochs as presumed ancestor of white Italian cattle. Bos Larteti Indre, 1866 ADDIN RW.CITE{{169 Duerst, J.U. 1931}}[86] after a skull fragment from Monte Sacro, Rome (Museum, Paris) considered a small aurochs (B.p. minutis) ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70], but by Duerst ADDIN RW.CITE{{169 Duerst, J.U. 1931/a;}}[86] identified as a female aurochs. Bos latifrons Fischer, 1830 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] for the late Pliocene aurochs from northern Indias Shivalik Hills, possibly a later form of the Bos acutifrons / Bos planifrons. Bos leucoprymnus Wagner, 1837 ADDIN RW.CITE{{157 Wagner, A.J. 1837}}[94] Rtimeyer presumed it to be a hybrid form between banteng and European cattle. Bos longifrons Owen, 1846 ADDIN RW.CITE{{149 Lydekker, R. 1898; 188 Owen, R. 1846}}[70,81] After the term brachyceros had been for the wild Bubalus brachyceros ADDIN RW.CITE{{32 Epstein, H. 1971}}[82], Owen ADDIN RW.CITE{{188 Owen, R. 1846}}[81]changed Bos brachyceros into B. longifrons, meaning long-headed, which fitted the type as well ADDIN RW.CITE{{169 Duerst, J.U. 1931; 32 Epstein, H. 1971}}[82,86]. Although Rtimeyer ADDIN RW.CITE{{152 Rtimeyer, L. 1861/a;}}[99] did not agree, this name became used by several authors. Rtimeyer ADDIN RW.CITE{{152 Rtimeyer, L. 1861/a;}}[99] and Owen ADDIN RW.CITE{{149 Lydekker, R. 1898; 188 Owen, R. 1846}}[70,81] agreed that this small Neolithic type or Celtic shorthorn was domestic, thus should be named Bos taurus, var longifrons. Rtimeyer ADDIN RW.CITE{{152 Rtimeyer, L. 1861/a;}}[99] postulated an Asian origin ADDIN RW.CITE{{32 Epstein, H. 1971/psee ;}}[82], but Hilzheimer ADDIN RW.CITE{{67 Hilzheimer, M. 1926/a;}}[83] showed that it was not a single, basic domestic type, let alone an aurochs form, so a Bos longifrons never existed as such. Bos macroceros Drst, 1899 ADDIN RW.CITE{{158 Drst, J. Ulrich 1899}}[92] presented this African type together with B. akeratos and B. brachyceros as one of the three basic modified types of domestic cattle, later indicated as B.p. macroceros / B.t. macroceros. Bos mastodontis Pohlig, 1911 ADDIN RW.CITE{{151 Pohlig, H. 1911}}[80] replacing after one year the term Bos brachyceroides for a small type of aurochs, after small aurochs skulls, later turned out to have belonged to female aurochsen. Bos namadicus Falconer, 1859 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] according to the current view representing the Asian aurochs as a separate species, from which the Bos indicus is the domestic form. Remains of the namadicus are very scant and its taxonomy has been disputed for decades. Rtimeyer ADDIN RW.CITE{{34 Rtimeyer, L. 1867}}[79] incorrectly presumed this to be the ancestor of Bos primigenius. Lydekker ADDIN RW.CITE{{173 Lydekker, R. 1912}}[102] referred to it as the Narbada Ox and supposed that this form was probably related to the banting (banteng) or a close allied species, that was the ancestor of the zebu as well as ancient Egyptian cattle and modern Hungarian longhorned cattle. Adametz ADDIN RW.CITE{{28 Adametz, L. 1926}}[96] stated that his student Liebscher observed a considerable variation in these skulls, some resembling the Bos primigenius Bojanus, others Bibovine type cattle. Bos (Urus) namadicus Falconer, 1865 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] synonym for Bos namadicus. Bos opisthonomus Pomel, 1893 ADDIN RW.CITE{{159 Pomel, A.. 1893}}[90] for a North-African Wild Race identical to B. primigenius mauritanicus and B. curvidens ADDIN RW.CITE{{252 Szalay, A.B. 1930}}[69] and also not differing from the Spanish Bos ibericus or the Bos primigenius ADDIN RW.CITE{{65 Leithner, O.F 1927/a;}}[78]. Bos opisthonotus, according to ADDIN RW.CITE{{169 Duerst, J.U. 1931}}[86] miswriting of opisthonomus by Troussart. Bos palaeogarus Rtimeyer, 1867 ADDIN RW.CITE{{34 Rtimeyer, L. 1868}}[79] for a skull that according to Lydekker ADDIN RW.CITE{{173 Lydekker, R. 1912}}[102] belonged to a banting [banteng]. Bos planifrons Lydekker, 1898 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] Pliocene fossil that may have belonged to a female Bos acutifrons found in the same depostis of the Siwalik Hills in northern India. Bos primigenius Bojanus, 1827 ADDIN RW.CITE{{145 Bojanus, L.H. 1827}}[60] Bojanus introduced this name for a skeleton in the museum of Jena, considering it as a prehistoric (antediluvialis) species, not aware that it in fact was the historic aurochs, by Linnaeus designated as Bos taurus. Bos primigenius estrepsicerus Fernndez, 1998 ADDIN RW.CITE{{2 Fernndez, A. 1998/a;}}[33] postulated aurochs variant, with B.p. ibericus and B.p. hahni the supposed ancesters of a cluster of NorthWest-Iberian breeds (see also ADDIN RW.CITE{{174 de Lima, B. 1873}}[103,104]). Bos primigenius ferus Pira, 1926 ADDIN RW.CITE{{176 Pira, A. 1926}}[105] regarded as predecessor of brachyceros type cattle of which skeletal remains were excavated from a Stone Age site on the island of Stora Karls, of the southwest coast of Gottland, Sweden. Bos primigenius f.d., proposed for domestic humpless European cattle of Europe (f.d.: forma domesticata) Bos primigenius f.d. indicus Zeuner, 1963 ADDIN RW.CITE{{43 Zeuner, F.E. 1963}}[106] for domestic zebu. Bos primigenius frontosus Keller, 1905 ADDIN RW.CITE{{41 Keller, C. 1905}}[3] after Rtimeyers morphological type B.primigenius var. frontosus ADDIN RW.CITE{{34 Rtimeyer, L. 1868/a;}}[79] for cattle typified by heavy horns, flattened at the root, growing downward, and then turning sideways and finally upward, including Frieburg and Simmental strains as well as the English Longhorn. Keller presumed these breeds traced to a mixture of primigenius type and frontosus type domestic cattle. Bos primigenius hahni Hilzheimer, 1917 ADDIN RW.CITE{{177 Hilzheimer, M. 1917}}[107] a diluvial cranial fragment excavated in 1910 in Fajum (Egypt), by Hilzheimer identified as an important aurochs subtype, named Bos primigenius Hahni nova subspecies. By Adametz ADDIN RW.CITE{{161 Adametz, L. 1920/a;}}[108] used for Egyptian aurochs, but in 1926 shortened to Bos primigenius var Hahni Hilzheimer. According to Von Leithner ADDIN RW.CITE{{65 Leithner, O.F 1927/a;}}[78] identical to Bos trochoceros. Bos primigenius hollandicus Keller, 1905 ADDIN RW.CITE{{41 Keller, C. 1905}}[3] for the heavy lowland breeds of Holland, Schleswig-Holstein and Oldenburg, considered a primigenius form. Bos primigenius Italiae Pohlig, 1911 ADDIN RW.CITE{{151 Pohlig, H. 1911}}[80], also Bos Italiae (q.v.), Italian aurochs postulated on the basis of a cursory description of a skull fragment ADDIN RW.CITE{{65 Leithner, O.F 1927}}[78]. Bos primigenius macroceros Duerst, 1903 ADDIN RW.CITE{{178 Duerst, J.U. 1903;179 Antonius, O. 1922;}}[109,110] earlier described as Bos macroceros, also as B. macrceros. Bos primigenius maureticanus Pomel for an early aurochs fossil found in Algeria (probably miswriting of mauritanicus) ADDIN RW.CITE{{65 Leithner, O.F 1927/a;}}[78]. Bos primigenius mauritanicus Thomas, 1881 ADDIN RW.CITE{{180 Thomas, Phil. 1881; 149 Lydekker, R. 1898}}[70,111] for the North-African Wild Race or aurochs and identical to B. opisthonomus and B. curvidens ADDIN RW.CITE{{252 Szalay, A.B. 1930}}[69]. Bos primigenius minutus Von Malsburg, 1911 ADDIN RW.CITE{{169 Duerst, J.U. 1931; 250 La Baume, W. 1909; 256 Von Malsburg, K. 1911}}[86,112,113] for two skull fragments from the Mrkischen Museum (Danzig) and two crania excavated in West Prussia, identified as the first domestic cattle ADDIN RW.CITE{{244 Hilzheimer, M. 1925}}[77] but identified in 1927 as female aurochs [70]. Bos primigenius namadicus Falconer, 1859 by Epstein and Mason ADDIN RW.CITE{{181 Mason, I.L 1984}}[114] accepted as designation for the Asian aurochs as subspecies of Bos primigenius. Bos primigenius opisthonomus Epstein and Mason, 1984 ADDIN RW.CITE{{181 Mason, I.L 1984}}[114] name for the African type of aurochs as subspecies, after Pomels Bos opisthonomus. Bos primigenius podolicus Wagner, 1837 ADDIN RW.CITE{{157 Wagner, A.J. 1837}}[94] for Grey Steppe cattle of south-eastern Europe, southern Siberia, and also found in Italy and considered to show pure aurochs morphology. Bos primigenius primigenius distinguishing European aurochs from two other possible aurochs varieties: B.p. opisthonomus from North Africa and B.p. orthoceros from Central Asia. Bos primigeniu primigeniu possibly a miswriting of B.p .primigenius ADDIN RW.CITE{{83 Miranda do Vale, J. 1907}}[115]. Bos primigenius (recentiorum) Storer, 1877 ADDIN RW.CITE{{183 Storer, J. 1877}}[116] presumably indicating the Holocene aurochs to be distinguished from the Pleistocene type aurochs. Bos primigenius Siciliae Pohlig, 1911 ADDIN RW.CITE{{151 Pohlig, H. 1911}}[80], also Bos Siciliae ADDIN RW.CITE{{252 Szalay, A.B. 1930}}[69] postulated on the basis of cursory description of a skull fragment from Sicily ADDIN RW.CITE{{65 Leithner, O.F 1927}}[78]. Bos primigenius var. trochoceros Rtimeyer, 1861 ADDIN RW.CITE{{152 Rtimeyer, L. 1861; 34 Rtimeyer, L. 1868}}[79,99], initially considered an early wild form Bos trochoceros, ADDIN RW.CITE{{152 Rtimeyer, L. 1861}}[99] later identified as a domestic type: B.taurus var. Trochoceros ADDIN RW.CITE{{34 Rtimeyer, L. 1868/pSee ;}}[79]. Bos priscus Bojanus, 1827 ADDIN RW.CITE{{145 Bojanus, L.H. 1827}}[60] also named Urus priscus, before its identification as Bison priscus. Bos pusio Swainson, 1835 ADDIN RW.CITE{{149 Lydekker, R. 1898; 242 Swainson, W. 1835}}[70,117] after a longhorned zebu cranium, by Rtimeyer ADDIN RW.CITE{{34 Rtimeyer, L. 1868/a;}}[79] identified as a domestic form B. indicus var. Pusio. Bos scoticus Swainson, 1835 ADDIN RW.CITE{{149 Lydekker, R. 1898; 242 Swainson, W. 1835}}[70,117] for Chillingham and other white park cattle, also B.t. scoticus or Urus scoticus. Bos scythicus Johnston, 1661 by Zimmerman (1780) ADDIN RW.CITE{{230 Zimmerman, E.A.M. 1780}}[118] mentioned as a type of humped cattle (Bckelochse), illustrated in the form of a bull with a dromedary-like hump. Bos scythicus gibbosus Charleton, 1668 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] presumably for polled humped cattle; cattle of the Scythians were considered to be polled and gibbus means hump. Bos siciliae Pohlig, 1911 ADDIN RW.CITE{{151 Pohlig, H. 1911; 252 Szalay, A.B. 1930}}[69,80] also as B. p. Siciliae (q.v.). Bos silvestris Charleton, 1668 ADDIN RW.CITE{{38 Duerst, J.U. 1931}}[93] for wild bovids as opposed to Bos domesticus for domestic cattle. Bos sondaicus Blyth, 1842 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] for Bali cattle, domestic form of the Java banteng (Bos javanicus). Keller ADDIN RW.CITE{{121 Keller, C. 1902/a;}}[68] used this name for the banteng, which he incorrectly assumed to be the ancestor of all Asian cattle. Therefore he used sondaicus as prefix for a number of presumed stem forms. Bos sondaicus africanus Keller, 1905 ADDIN RW.CITE{{41 Keller, C. 1905}}[3] name for the sanga (by then known from Abyssinia, towards the upper Nile and Lake Chad). The name is possibly taken from Bos taurus brachyceros Africanicus ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2]. Keller did not believe in an African aurochs and was convinced that all cattle were imported into that continent. Bos sondaicus akeratos Keller, 1905 ADDIN RW.CITE{{41 Keller, C. 1905}}[3] for hornless cattle found in several places in Europe, Asia and Africa. The taxon refers to the domestic Bos akeratos of Arenander ADDIN RW.CITE{{66 Arenander, E.O 1898/a;}}[72], though Keller stated the type was of Asian origin, developed separate from small-horned cattle in several places. Bos sondaicus brachyceros Keller, 1905 ADDIN RW.CITE{{41 Keller, C. 1905}}[3] According to Keller more appropriate for Bos brachyceros, as he postulated the origin of the wild form was found in the Asian zebu, which in turn was assumed to descend from the banteng as already proposed by Rtimeyer ADDIN RW.CITE{{34 Rtimeyer, L. 1868}}[79]. According to Keller this type of cattle, present in East Asia, West Asia and North Africa was already known in the time of the pharaos in the Nile Valley and included the prehistoric Torfrind, Braunvieh, Albanian cattle, Polish Red and the Jersey. Bos sondaicus brachycephalus Keller, 1905 ADDIN RW.CITE{{41 Keller, C. 1905}}[3] Keller was convinced this domestic form had developed by selection on European soil. It was typified by a short head, very broad forehead and strong rounded horns and included the breeds Eringer (Hrens), Duxer (Tuxer), Pustertaler, Egerlander, Voigtlnder, Devon, Sussex and Hereford. Bos sondaicus indicus Keller, 1905 ADDIN RW.CITE{{41 Keller, C. 1905}}[3] for the Indian zebu and German colonial East-African zebus. Bos sondaicus longicornis Keller, 1905 ADDIN RW.CITE{{41 Keller, C. 1905}}[3] for an ancient form of Sanga, depicted in earliest Egyptian cultures and receded to central African Lake district, where it is known as Watusi cattle. Keller believed this form had entered Africa from Asia, as he did not believe in an African aurochs. Bos stenometopon Sismonda, 1861 ADDIN RW.CITE{{34 Rtimeyer, L. 1868}}[79] for a Pliocene skull excavated near Asti, Italy, by Rtimeyer ADDIN RW.CITE{{34 Rtimeyer, L. 1868/a;}}[79] identified as Bos (Amphibos) etruscus ADDIN RW.CITE{{170 Falconer, H. 1868/psee Bos etruscus ;}}[95]. Bos synophrys Fischer, 1829 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] no description given. Bos sylvestris Bonaparte, 1845 ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] wild white cattle in Great Britain. According to John Leslie, Bisshop of Ross (1598) ADDIN RW.CITE{{57 Hengeveld, G.J. 1865}}[119] Bos sylvestris roamed the woods of Scotland in large numbers, especially in the Sylva Caledonia, while the chronicler Fritz-Stephen around 1174 refers to tauri sylvestres (Uri sylvestres) in the forest near London ADDIN RW.CITE{{173 Lydekker, R. 1912}}[102]. The B. sylvestris of Belonius mentioned in 1780 by Shaw ADDIN RW.CITE{{185 Shaw, T. 1780/a;}}[120] might also indicate wild cattle ADDIN RW.CITE{{57 Hengeveld 1865/a;}}[119]. Bos taurus Linnaeus, 1758 ADDIN RW.CITE{{117 Linnus, C. 1758}}[61] for European domestic cattle as well as their wild predecessor, the urus as described by Caesar. Currently the most common zoological name for domestic, humpless or taurine cattle. Linnaeus seemed uncertain about the extinction of the wild form in Poland. Bos Taurus, by Fitzinger 1860 applied for the Thallandrind ADDIN RW.CITE{{89 Fitzinger, L.J. 1860/a;}}[67], meaning valley type cattle as found in the lower parts and hills of Switzerland, Germany, Austria, France, England and Spain. Bos taurus abyssinicus Gmelin, 1788 ADDIN RW.CITE{{252 Szalay, A.B. 1930}}[69] for B. t. domesticus abyssinicus (q.v.) and identical to B. macroceros Drst. Bos taurus aceratos Hilzheimer, 1926 ADDIN RW.CITE{{67 Hilzheimer, M. 1926; 116 Wilson, D.E. 1993}}[59,83] possibly B.t. akeratos Bos taurus adelensis Boddaert, 1785 ADDIN RW.CITE{{116 Wilson, D.E. 1993; 253 Boddaert, P. 1784}}[59,121] might indicate white cattle. Bos taurus aegyptiacus Lydekker, 1904 ADDIN RW.CITE{{173 Lydekker, R. 1912; 116 Wilson, D.E. 1993}}[59,102] after Fitzingers Bos aegyptiorum. Bos taurus akeratos Arenander, 1898IC by only two authors ADDIN RW.CITE{{66 Arenander, E.O 1898}}[72] considered as aurochs. Keller ADDIN RW.CITE{{41 Keller, C. 1905/a;}}[3] accepted the polled Bos sondaicus akeratos as a separate domestic type. Bos taurus albus Sundevall, 1848 ADDIN RW.CITE{{38 Duerst, J.U. 1931}}[93], with Urus, gaur, gayal and banteng subspecies of Boves proprii (propercattle). Bos taurus alpestris Wagner, 1837 ADDIN RW.CITE{{157 Wagner, A.J. 1837; 116 Wilson, D.E. 1993}}[59,94] possibly the same as Sansons B.t. alpinus. Bos taurus alpinus Sanson, 1878 ADDIN RW.CITE{{166 Sanson, A. 1878; 116 Wilson, D.E. 1993}}[59,73] name for cattle of the presumed ethnic Alpine Dolichocphale type: Swiss, Tarentaise and Gasconne. In 1893 Sanson added taurus to most of his names for taurine cattle. Bos taurus alpium Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860; 171 Anonymous; 116 Wilson, D.E. 1993}}[59,67,100] former B.alpium Bos taurs akeratos Arenander, 1898 ADDIN RW.CITE{{66 Arenander, E.O 1898; 171 Anonymous; 116 Wilson, D.E. 1993}}[59,72,100] former B. akeratos Bos taurus aquitanicus Sanson, 1878 ADDIN RW.CITE{{166 Sanson, A. 1878; 171 Anonymous; 116 Wilson, D.E. 1993}}[59,73,100] former B. aquitanicus, also as B.t. aquitanus, indicated as cattle of the presumed ethnic Aquitaine Dolichocphale type: Blonde dAquitaine, Limousin and Lourdaise. Bos taurus arnei Amschler, 1939 ADDIN RW.CITE{{68 Amschler, J.W. 1956; 171 Anonymous; 116 Wilson, D.E. 1993}}[59,100,122] former B.brachyceros arnei. Bos taurus arvernensis Sanson, 1878 ADDIN RW.CITE{{166 Sanson, A. 1878; 116 Wilson, D.E. 1993}}[59,73] for the Brachycphale type breeds Salers and Ferrandaise. Bos taurus asiaticus Sanson, 1878 ADDIN RW.CITE{{166 Sanson, A. 1878; 116 Wilson, D.E. 1993}}[59,73] former Bos asiaticus, for cattle of the presumed ethnic Asiatic Brachycphale type, and fantasized origin of the cattle of the Camargue. Bos taurus atlanticus Sanson, 1878 ADDIN RW.CITE{{166 Sanson, A. 1878}}[73] one of the twelve presumed basic troncos (strains or types, ADDIN RW.CITE{{83 Miranda do Vale, J. 1907}}[115]). Bos taurus balticus Stegmann von Pritzwald, 1924 ADDIN RW.CITE{{168 Stegmann von Pritzwald, F.P. 1924; 116 Wilson, D.E. 1993}}[59,75] former B.balticus. Bos taurus batavicus Sanson, 1878 ADDIN RW.CITE{{166 Sanson, A. 1878; 116 Wilson, D.E. 1993}}[59,73] for the presumed ethnic Lowland Dolichocphale type, Dutch and Flemish cattle. Bos taurus brachiceros Brehm, 1864 ADDIN RW.CITE{{186 Brehm, A.E. 1875; 116 Wilson, D.E. 1993}}[59,123] misspelling of B.brachyceros. Bos taurus brachycephalus Wilckens, 1876 ADDIN RW.CITE{{160 Wilckens, M. 1876; 116 Wilson, D.E. 1993}}[1,59] also found as B.t. brachicephalus for short-headed type cattle, a cranial type excavated in Italian soil, dating back to the Roman period, and presumed the foundation for several short-headed breeds of Alpine cattle. Keller changed the name to Bos sondaicus brachycephalus. Bos taurus brachycephalus africanicus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for North-African Algerian Brown Atlas cattle. Bos taurus brachycephalus alverniensis Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67] after Sansons B. arverniensis for Salers and related breeds from Auvergne. Bos taurus brachycephalus aquitanicus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] after Sansons B. aquitanicus for Limousin and former Garonne and related Southwest-French blond breeds, which in 1961 were amalgamated into the Blonde dAquitaine. Bos taurus brachycephalus britannicus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] after Sansons B. britanicus for West-English breeds (Devon, Sussex, Hereford). Bos taurus brachycephalus celticus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for breeds of French Brittany (Bretonne Pie-Noir and related breeds), Ireland (Kerry), and Wales (Welsh Black and former related strains). Bos taurus brachycephalus ibericus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] after Sansons B. ibericus for unspecified Iberian breeds. Bos taurus brachycephalus isolanus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for cattle from Sicily, Corsica and the Camargue. Bos taurus brachycephalus Italicus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for North-Italian Grey Alpine and former varieties. Bos taurus brachycephalus licestriensis Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67] for the Longhorn (former Leicester). Bos taurus brachycephalus pyreneus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for the many Pyrenean strains of cattle. Bos taurus brachycephalus salisburgensis Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for Pinzgauer, Pustertal, Ennstal Spotted and former related strains. Bos taurus brachycephalus tauricus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for breeds of the Vosges Mountains, Wallis and Tyrol. Bos taurus brachycephalus teutonicus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for a number of (now mainly extinct) South- and Southwestern-German red-brown, red white-headed, red white-backed and red-pied breeds. Bos taurus brachyceroides Pohlig, 1911 ADDIN RW.CITE{{151 Pohlig, H. 1911; 116 Wilson, D.E. 1993}}[59,80] former B. brachyceroides Bos taurus brachyceros Owen, 1843 ADDIN RW.CITE{{188 Owen, R. 1846; 116 Wilson, D.E. 1993}}[59,81] former B. brachyceros, short-horned domestic cattle of which remains from 6750 yr ago were unearthed in North Africa and from 6000 years ago in Iran, Asia and Europe (Neolithic Shorthorn or Celtic Shorthorn) Adametz ADDIN RW.CITE{{64 Adametz, L 1895/a;}}[124] noted similarities with modern, small shorthorned Balkan cattle, for which he introduced the term Illyrian and is most probably is the nearest in type to the early shorthorned domestic stock. Keller ADDIN RW.CITE{{121 Keller, C. 1902/a;}}[68] assumed that these shorthorned cattle were introduced from Asia (Bos sundaicus brachyceros) and in 1997 Payne ADDIN RW.CITE{{105 Payne, W.J.A. 1997/a;}}[125] still refers to ideas of a separate shorthornded domestication for this type. It is now accepted that short-horned cattle were the result of degeneration due to the domestication of Bos primigenius, already put forward by Dawkins ADDIN RW.CITE{{189 Dawkins, W.B. 1866/a;}}[126] and Nehring ADDIN RW.CITE{{203 Nehring, A. 1888/a;}}[127]. Bos taurus brachyceros Africanicus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] after B. africanus, for Sanga cattle and considered a separate type that had not changed over thousands of years. By Morse ADDIN RW.CITE{{199 Morse, E.W. 1912/a;}}[128] and Newbold ADDIN RW.CITE{{200 Newbold, D. 1928/a;}}[129] used to designate ancient Egyptian cattle, regarded as a local race of the primigenius type. Bos taurus brachyceros palustris Cardas, 1936 ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] palustris for living in the marshes as the former Polesian cattle from the Pripet Marshes of Byelorussia and Ukraine. Bos taurus brachyceros polonicus Cardas, 1936 ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] after Adametz B. t. longifrons (brachyceros) polonicus. Bos taurus britannicus Sanson, 1878 ADDIN RW.CITE{{166 Sanson, A. 1878; 116 Wilson, D.E. 1993}}[59,73] also found as B. britanicus for breeds from Great Britain, indicated as cattle of the presumed British Dolichocphale type. Bos taurus bunelli Frick, 1937 ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] no description given. Bos taurus burdeyalensis ADDIN RW.CITE{{171 Anonymous}}[100] for Bordelaise cattle. Bos taurus caledoniensis Sanson, 1878 ADDIN RW.CITE{{166 Sanson, A. 1878; 116 Wilson, D.E. 1993}}[59,73] for cattle of the presumed ethnic Scottish Brachycphale type. Bos taurus chinensis Swinhoe, 1870 ADDIN RW.CITE{{190 Swinhoe, R. 1870; 45 Wilson, J. 1909; 171 Anonymous}}[100,130,131] as B.t. indicus chinensis. Bos taurus collicerus Rostafinski, 1933 ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] no description given. Bos taurus communis S.D.W., 1836ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] probably indicates common cattle (taurine and zebu). Bos taurus curvidens Pomel, 1893 ADDIN RW.CITE{{159 Pomel, A.. 1893; 116 Wilson, D.E. 1993; 171 Anonymous}}[59,90,100] domestic form of the B. curvidens. Bos taurus dante Link, 1794 ADDIN RW.CITE{{171 Anonymous; 116 Wilson, D.E. 1993}}[59,100] and as B.t. indicus dante, former Bos Dante (q.v.), identified as a polled zebu skull by Rtimeyer ADDIN RW.CITE{{34 Rtimeyer, L. 1868/a;}}[79]. Bos taurus desertorum Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860; 116 Wilson, D.E. 1993}}[59,67] for Steppe cattle found from Mongolia to Eastern Europe and southern Italy. Bos taurus ditophus Fischer, 1829, 1914 ADDIN RW.CITE{{149 Lydekker, R. 1898; 171 Anonymous; 116 Wilson, D.E. 1993}}[59,70,100] no description given. Bos taurus domesticus Erxleben, 1777 ADDIN RW.CITE{{119 Erxleben, J.C.P. 1777; 116 Wilson, D.E. 1993}}[59,63] for domestic cattle. Bos taurus domesticus abyssinicus Pennant, 1771, later Bos taurus abyssinicus Gmelin 1788 and Bos abyssinicus Oken 1838 ADDIN RW.CITE{{252 Szalay, A.B. 1930}}[69], brachyceros type skull with gigantic long horns ADDIN RW.CITE{{177 Hilzheimer, M. 1917}}[107] and identical to Bos macroceros Drst ADDIN RW.CITE{{252 Szalay, A.B. 1930}}[69]. Bos taurus dunelmensis Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860; 116 Wilson, D.E. 1993}}[59,67] for the Durham (Shorthorn) breed. Bos taurus ecornis Wagner, 1837 ADDIN RW.CITE{{157 Wagner, A.J. 1837; 116 Wilson, D.E. 1993}}[59,94] former Bos ecornis. Bos taurus europaeus Kerr, 1792 ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] presumed wild European ancestor of domestic cattle. Bos taurus ferus for wild cattle from Chillingham park ADDIN RW.CITE{{152 Rtimeyer, L. 1861/a;}}[99], also named Bos urus scoticus ADDIN RW.CITE{{157 Wagner, A.J. 1837}}[94] and White Urus ADDIN RW.CITE{{149 Lydekker, R. 1898; 254 Cuvier, M. 1827}}[70,132]. Bos taurus forma domesticata Zeuner & Mourant, 1963 ADDIN RW.CITE{{148 Zeuner, F.E. 1963}}[133] proposed for domestic cattle. Bos taurus fossilis Baer ADDIN RW.CITE{{252 Szalay, A.B. 1930}}[69] Bos fossilis (q.v.). Bos taurus friburgensis Fitzinger, 1860 ADDIN RW.CITE{{171 Anonymous; 89 Fitzinger, L.J. 1860}}[67,100] for the extinct Fribourg (Freiburger) breed. Bos taurus frisius Wagner, 1837 ADDIN RW.CITE{{157 Wagner, A.J. 1837; 116 Wilson, D.E. 1993}}[59,94] for straightbacked Friesian-type cattle. Bos taurus frisius Wagner, 1837 ADDIN RW.CITE{{157 Wagner, A.J. 1837; 169 Duerst, J.U. 1931; 116 Wilson, D.E. 1993}}[59,86,94]. Bos taurus frontosus Nilsson, 1849 ADDIN RW.CITE{{77 Nilsson, S. 1849; 116 Wilson, D.E. 1993}}[59,98] Initially considered a separate Scandinavian aurochs (Bos frontosus) and the ancestor of small, often polled Norwegian cattle. By Rtimeyer (1867) ADDIN RW.CITE{{34 Rtimeyer, L. 1868/a;}}[79] considered a domestic form: Bos taurus var. Frontosus, in 1892 definitive identified as remains of a domestic specimen. Bos taurus frontosus burgundicus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for Swiss Fleckvieh (Fribourg and Simmental) and related breeds of central Europe and France (Montbliarde). Bos taurus frontosus franconicus Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67] for Franconian and other German yellow breeds and the French Charolais. Bos taurus frontosus goticus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for the Smland-Gotland breed. Bos taurus frontosus noricus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for Austrian Blond breeds. Bos taurus frontostus piemontanus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for Piemontese and related strains. Bos taurus galla Salt, 1814 ADDIN RW.CITE{{171 Anonymous}}[100] former Bos Galla, also as B. indicus Galla. Bos taurus garumnensis ADDIN RW.CITE{{171 Anonymous}}[100] French Garonnaise breed. Bos taurus germanicus Sanson, 1878 ADDIN RW.CITE{{166 Sanson, A. 1878; 167 Sanson, A. 1893}}[73,74] former B. germanicus, indicating cattle of the presumed ethnic Dolichocphale cattle of Normandy. Bos taurus giganteus Owen, 1846 ADDIN RW.CITE{{188 Owen, R. 1846; 149 Lydekker, R. 1898; 173 Lydekker, R. 1912}}[70,81,102] description of huge skulls from Ilford, kept in the British Museum . Bos taurus grandifrons Kaltenegger, 1904 ADDIN RW.CITE{{79 Kaltenegger, Ferdinant 1904}}[8] large-head cattle, alternative of frontosus (broad-headed) for several Austrian breeds such as the Pinzgauer. Bos taurus hamiticus Adametz, 1920 ADDIN RW.CITE{{161 Adametz, L. 1920}}[108] for domestic longhorned cattle of North-African (Egyptian) descent. Bos taurus hibernicus Sanson, 1878 ADDIN RW.CITE{{166 Sanson, A. 1878; 116 Wilson, D.E. 1993}}[59,73] for cattle presumed to descent (Cornish) Irish Dolichocphale type cattle, the Channel Island breeds, Bretonne, Froment du Lon, and Jersiaise. Bos taurus hollandicus Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860; 116 Wilson, D.E. 1993}}[59,67] for Dutch-Friesian cattle. Bos taurus hungaricus Gray, 1846 ADDIN RW.CITE{{240 Gray, J.E. 1852}}[91] for Hungarian Steppe cattle ADDIN RW.CITE{{192 Vasey, G. /a; 169 Duerst, J.U. 1931}}[86,134]. Bos taurus hypselurus Wagner, 1937 ADDIN RW.CITE{{157 Wagner, A.J. 1837; 171 Anonymous; 116 Wilson, D.E. 1993}}[59,94,100] synonymous for B.t. taurus for with their back sloping up from withers to hindquarters, contrary to the straight backed type Taurus Frisius ADDIN RW.CITE{{38 Duerst, J.U. 1931}}[93]. Bos taurus ibericus Sanson, 1878 ADDIN RW.CITE{{166 Sanson, A. 1878; 116 Wilson, D.E. 1993}}[59,73] former B. ibericus, according to Sanson ancestor of Brachycphale breeds: Corse, Basque and Barnais; in Spanish-Portuguese literature as B.t. ibricus, predecessor of brown and black, elegant cattle in central and south Spain and adjoining Portugal. Bos taurus indicusIC ADDIN RW.CITE{{171 Anonymous; 116 Wilson, D.E. 1993}}[59,100] disputed name for Bos indicus, just as B.T indicus var. major (large zebu) and B.t. indicus minor (small zebu). Bos taurus indicus abessinicus Kerr, 1792 ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] also found as: Bos taurus indicus abessynicus Fischer, 1829 ADDIN RW.CITE{{169 Duerst, J.U. 1931; 116 Wilson, D.E. 1993}}[59,86] presumably for Ethiopian cattle. Bos taurus indicus aegyptiorum Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860; 116 Wilson, D.E. 1993}}[59,67] presumably for Egyptian cattle. Bos taurus indicus aethiopicus Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860; 116 Wilson, D.E. 1993}}[59,67] originally as B. Zebu africanus aethiopicus. Bos taurus indicus africanus Kerr, 1792 ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] for African cattle. Bos taurus indicus brookii Smith, 1827 ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] no description given. Bos taurus indicus chinensis Swinhoe, 1870 ADDIN RW.CITE{{190 Swinhoe, R. 1870; 116 Wilson, D.E. 1993}}[59,130] also as Bos taurus chinensis, for Chinese cattle. Bos taurus indicus dante Link, 1794 ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] originally named Bos Dante (q.v.). Bos taurus indicus galla Salt, 1814 ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] originally as Bos Galla (q.v.). Bos taurus indicus gibbosus Blyth, 1860 ADDIN RW.CITE{{193 Blyth, E. 1860}}[135] ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] originally as Zebu gibbosus, no description given. Bos taurus indicus harveyi de Rochebrune, 1882 ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] no description given. Bos taurus indicus hottentottus Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860; 116 Wilson, D.E. 1993}}[59,67] former Bos Zebu africanus hottentottus, cattle of the South-African Khoi herders (Hottentot). Bos taurus indicus hybridus Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860; 116 Wilson, D.E. 1993}}[59,67] presumably for zebu taurine crossbred cattle. Bos taurus indicus madagascariensis Kerr, 1792 ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] for Madagascar zebu. Bos taurus indicus major Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860; 116 Wilson, D.E. 1993}}[59,67] presumably for large Indian zebu type. Bos taurus indicus medius Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860; 116 Wilson, D.E. 1993}}[59,67] presumably for medium large Indian zebu type. Bos taurus indicus pusio Swainson, 1835 ADDIN RW.CITE{{242 Swainson, W. 1835; 116 Wilson, D.E. 1993}}[59,117] originally as Bos pusio (q.v.). Bos taurus indicus sanga Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860; 116 Wilson, D.E. 1993}}[59,67] presumably for sanga. Bos taurus indicus triceros Rochebrune, 1882 ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] no description given. Bos taurus indicus zebu Boddaert, 1785 ADDIN RW.CITE{{116 Wilson, D.E. 1993; 253 Boddaert, P. 1784}}[59,121] for domestic zebu. Bos taurus inermis Boddaert, 1785 ADDIN RW.CITE{{116 Wilson, D.E. 1993; 253 Boddaert, P. 1784}}[59,121] possibly indicates hornless cattle (inermis = unarmed) Bos taurus jurassicus Sanson, 1878 ADDIN RW.CITE{{166 Sanson, A. 1878; 116 Wilson, D.E. 1993}}[59,73] former B. jurassicus, for the Brachycphale type breeds, identical to the B. frontosus: Pie-rouge de lest, Montbliarde, Abondance and Charolais Bos taurus latifrons Kalternegger, 1904 ADDIN RW.CITE{{79 Kaltenegger, F. 1904}}[8] for several Austrian cattle breeds of the shorth-headed type, by Wilckens ADDIN RW.CITE{{160 Wilckens, M. 1876}}[1] named brachycephalus. Kaltenegger, however, used the name broad-headed in stead of shorth-headed. However, Fischer (1830) ADDIN RW.CITE{{149 Lydekker, R. 1898;}}[70] already had used the term Bos latifrons for Pliocene Bos remains excavated in the Siwalik range of India. Bos taurus ligeriensis Sanson, 1878 ADDIN RW.CITE{{166 Sanson, A. 1878; 116 Wilson, D.E. 1993}}[59,73] former B. ligeriensis, for the Brachycphale type breeds: Parthenaise and Aubrac. Bos taurus longifrons ADDIN RW.CITE{{188 Owen, R. 1846}}[81] ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] for Bos longifrons, indicating domestic cattle rather than an aurochs species. Bos taurus var. longifrons Lydekker, 1885 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] synonym for B.t. longifrons. Bos taurus longifrons alpestris Wagner, 1837 ADDIN RW.CITE{{157 Wagner, A.J. 1837}}[94] with the proposed varieties B.t.l. var. brunneus (Swiss, German, Austrian Brown, etc.), B.t.l. var. griseus (Tyrol Grey, Grey Alpine, etc.) and B.t.l. var. flavus (German and Austrian Yellow). Bos taurus longifrons illyricus Adametz, 1895 ADDIN RW.CITE{{64 Adametz, L 1895}}[124] for Illyrian Shorthorn cattle (Busha) of the Balkans. Bos taurus longifrons isolanus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for Channel Island cattle (Jersey, Guernsey, former Alderney). Bos taurus longifrons ligeriensis Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] after Sansons B.t. ligeriensis for breeds and strains of the Vende (Parthenaise, Nantaise, Marachine). Bos taurus longifrons polonicus Adametz, 1895 ADDIN RW.CITE{{64 Adametz, L 1895}}[124] for extinct Polish Brown, Busha-like cattle from the West Carpathians. Bos taurus longifrons vasconiensis Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for the Gasconne, Bazadaise and related grey lowland and highland breeds. Bos taurus macroceros Drst, 1899 ADDIN RW.CITE{{158 Drst, J.U. 1899; 116 Wilson, D.E. 1993}}[59,92] collective name for the zoological types Bos Galla, Bos Dante and Bos triceros, which represented several East- and West-African cattle. Bos taurus major Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860; 171 Anonymous}}[67,100] presumably B.t. indicus major Bos taurus mastodontis Pohlig, 1911IC ADDIN RW.CITE{{151 Pohlig, H. 1911}}[80] former Bos mastodontis (q.v.). Bos taurus mauritanicus Thomas, 1881 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] after B.p. mauritanicus for domestic cattle of African descent. Bos taurus minor Owen, 1846 ADDIN RW.CITE{{188 Owen, R. 1846; 116 Wilson, D.E. 1993}}[59,81] for a hypothetic small type of aurochs. Bos taurus minutis Von der ADDIN RW.CITE{{256 Von Malsburg, K. 1911}}[113]Malsburg, 1911 ADDIN RW.CITE{{149 Lydekker, R. 1898; 116 Wilson, D.E. 1993; 256 Von Malsburg, K. 1911}}[59,70,113] for a hypothetic small type of aurochs, which turned out to be female aurochs remains. Bos taurus orthoceros Stegmann von Pritzwald, 1906; 1912 ADDIN RW.CITE{{194 Stegmann von Pritzwald, F.P. 1906;195 Stegmann von Pritzwald, F.P. 1912/a; 116 Wilson, D.E. 1993}}[59,136,137] for Kalmuk-Kirgiz (Turano-Mongolian type) cattle from central and eastern Asia, typified by peculiar upright direction of the horns that influences the cranial conformation as well as a ridge along the forehead from the poll downwards. Initially Stegmann von Pritzwald ADDIN RW.CITE{{194 Stegmann von Pritzwald, F.P. 1906/a;}}[136] assumed a descent from Bali cattle. In 1912 he as well as Kuleschow et al. ADDIN RW.CITE{{195 Stegmann von Pritzwald, F.P. 1912}}[137] considered it as a taurine zebu crossbred, but Adametz ADDIN RW.CITE{{28 Adametz, Leopold 1926/a;}}[96] postulated a separate wild form. Bos taurus planifrons Kaltenegger, 1904 ADDIN RW.CITE{{79 Kaltenegger, F. 1904}}[8] synonym for the flat forehead primigenius type breeds of Austria. However, Lydekker ADDIN RW.CITE{{149 Lydekker, R. 1898/a;}}[70] already had use the designation Bos planifrons for a Pliocene skull from the Siwalik range of northern India. Bos taurus podolicus Wagner, 1837 ADDIN RW.CITE{{157 Wagner, A.J. 1837; 116 Wilson, D.E. 1993}}[59,94] renamed B. primigenius podolicus for Steppe cattle after the region Podolia in Southwest Ukraine. Bos taurus primigenius Lydekker, 1885 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] used for both aurochs and longhorned domestic cattle, presumed to trace directly to the aurochs (as all cattle). Bos Taurus var. Primigenius Rtimeyer, 1868 ADDIN RW.CITE{{34 Rtimeyer, L. 1868/a;}}[79], for domestic cattle tracing to the Bos primigenius. Bos taurus primigenius dacicus Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67] for steppe cattle from the lower Danube. Bos taurus primigenius ferus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for feral British White Park cattle. Bos taurus primigenius germanicus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] after Sansons B.t. germanicus for pied and unicolored red lowland breeds from the Netherlands (B.t.germanicus var. frisisus); N.W. Germany (B.t.g. var. saxonicus), Belgium (B.t.g. var. flandricus), Normandy (B.t.g. var normanus) and Britain (B.t.g. var. anglosaxonius). Bos taurus primigenius hungaricus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for Austrian-Hungarian Steppe type cattle. Bos taurus primigenius montanus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2].for intermediate steppe-shorthorn cattle from the Balkan Mountains. Bos taurus primigenius nomas Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for Kalmyk and Kirgiz cattle. Bos taurus primigenius norvegicus Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67] for Norwegian Mountain (Blacksided Trondheim). Bos taurus primigenius podolicus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] earlier B.p. podolicus for Podolian Steppe cattle. Bos taurus primigenius romanicus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for Italian Romagnola and related breeds. Bos taurus primigenius sarmaticus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] for North-Russian horned breeds. Bos taurus primigenius scoticus Werner, 1912 ADDIN RW.CITE{{37 Werner, A.H. 1912}}[2] after earlier Urus scoticus, B. urus scoticus, B. scoticus, B. t. scoticus, or B.t. var. scoticus. for Scottish Highland cattle, by also used for both the tame Scottish Highland and feral English park cattle ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67] Bos taurus primigenius suecicus Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67] for Fjell (Swedish Mountain) breed. Bos taurus priscus(Keferstein, 1834 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] after Urus priscus ADDIN RW.CITE{{145 Bojanus, L.H. 1827}}[60] for supposed aurochs remains that later were identified as originating from the steppe wisent Bison priscus. Bos taurus scythicus for Russian polled cattle, presumably after Charletons Bos scythicus., Several scholars maintained that polled cattle of North Russia, Scandinavia and Iceland were descended from the hornless cattle owned by the ancient Scythians described by Herodotus. Bos taurus scoticus Smith, 1827 ADDIN RW.CITE{{171 Anonymous; 116 Wilson, D.E. 1993}}[59,100] after Urus scoticus by Smith (1827) ADDIN RW.CITE{{149 Lydekker, R. 1898; 254 Cuvier, M. 1827}}[70,132], also as Bos scoticus. Bos taurus var. scoticus Bell and Alston, 1847 ADDIN RW.CITE{{149 Lydekker, R. 1898; 243 Bell,T. 1874}}[70,138] after Urus scoticus ADDIN RW.CITE{{149 Lydekker, R. 1898; 254 Cuvier, M. 1827}}[70,132] or Swainsons Bos scoticus ADDIN RW.CITE{{242 Swainson, W. 1835}}[117] for Chillingham and other feral park cattle. Bos taurus silvestris Fischer, 1829 [Szalay] for Bos silvestris Charleton, 1668 or B. primigenius. Bos taurus tinianus Boddaert, 1785 ADDIN RW.CITE{{116 Wilson, D.E. 1993}}[59] possibly designating the same as: Bos taurus tinianensis Fischer, 1829 ADDIN RW.CITE{{169 Duerst, J.U. 1931; 116 Wilson, D.E. 1993}}[59,86], white cattle with black ears from the Pacific Tinian island (Northern Mariana Islands, VS). Bos taurus var. Trochoceros Rtimeyer, 1861 ADDIN RW.CITE{{152 Rtimeyer, L. 1861}}[99] domestic form of the wild B. p. var. trochoceros and one of the races (Trochoceros Race) of the current Frontosus-Rind ADDIN RW.CITE{{34 Rtimeyer, L. 1868}}[79]. Bos taurus turdetanus Snchez Belda, 1981 ADDIN RW.CITE{{84 Sanchez Belda, Antonio 1981}}[13] postulated ancestor of blond and red, rounded French, Iberian, German, Austrian and British breeds. Bos taurus typicus Lydekker, 1898 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] for the Swedish domestic cattle, considered a typical representative of Bos taurus ADDIN RW.CITE{{173 Lydekker, R. 1912/a;}}[102]. Bos (Taurus) Urus Smith, 1827 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] aurochs. Bos taurus vulgaris Wagner, 1837 ADDIN RW.CITE{{157 Wagner, A.J. 1837; 116 Wilson, D.E. 1993}}[59,94] no description given Bos triceros Rochebrune, 1880 ADDIN RW.CITE{{158 Drst, J. Ulrich 1899}}[92] identified as a Senegalian type (B.t. macroceros) ADDIN RW.CITE{{158 Drst, J. Ulrich 1899}}[92]. Bos trochoceros Meyer, 1835 ADDIN RW.CITE{{163 Meyer, H. von 1835; 34 Rtimeyer, L. 1868; 179 Antonius, O. 1922}}[79,110,139] a diluvial (Pleistocene) form excavated in Arezzo. According to Rtimeyer ADDIN RW.CITE{{34 Rtimeyer, L. 1868;}}[79] Meyer based the existence of the trochoceros species on one female skull that was a primigenius variant (B. primigenius var. trochoceros) and was found in Swiss and German Stone Age cattle as well as in Wild White cattle from Lyme park. Early domestic cattle with similar skulls was denoted by B.taurus var.Trochoceros ADDIN RW.CITE{{34 Rtimeyer, L. 1868;}}[79]. Leithner (1927) ADDIN RW.CITE{{65 Leithner, O.F 1927}}[78] identified Bos trochoceros as being identical to Bos primigenius hahni. Bos trochoceros hahni Leithner, 1927 ADDIN RW.CITE{{65 Leithner, O.F 1927}}[78] combination of B. trochoceros and B. primigenius hahni and indicating the same type. Bos turano-mongolicus Kolesnik, 1936 ADDIN RW.CITE{{164 Kolesnik, N.N. 1936}}[140] a presumed basic type as ancestor of several breeds of central Asia: Kalmuck, Mongolian, Yakut and Kirgiz cattle. Bos urus Boddaert 1785 ADDIN RW.CITE{{116 Wilson, D.E. 1993; 253 Boddaert, P. 1784}}[59,121] for aurochs ADDIN RW.CITE{{254 Cuvier, M. 1827; 255 Fleming, J. 1928; 189 Dawkins, W.B. 1866}}[126,132,141]. Urus is the name Caesar gave to wild cattle living in Germanic woods. Cuvier ADDIN RW.CITE{{196 Cuvier, M. 1823/a;}}[142] used this name for a cranium, which later turned out to be from a wisent ADDIN RW.CITE{{57 Hengeveld, G.J. 1865}}[119]. A drawing from 1827 of an wild ox entitled Bos urus was copied from a painting on a panel from the 16th century, which showed the word Thur (aurochs) in golden German characters in one of the corners ADDIN RW.CITE{{254 Cuvier, M. 1827}}[132]. Reynolds ADDIN RW.CITE{{197 Reynolds, S.H. 1939/a;}}[143] used urus as the common name for B. primigenius. Bos urus (antiquorum) Storer, 1877 ADDIN RW.CITE{{183 Storer, J. 1877}}[116] presumably indicating the Pleistocene aurochs. Bos urus Gesneri, also for Bos primigenius ADDIN RW.CITE{{57 Hengeveld 1865}}[119] after the print of an aurochs by Gessner in a text of Jonston (1660). Bos urus minutus von der Malsburg, 1911 ADDIN RW.CITE{{169 Duerst, J.U. 1931; 256 Von Malsburg, K. 1911; 244 Hilzheimer, M. 1925; 67 Hilzheimer, M. 1926; 245 Grigson, C. 1978}}[77,83,86,113,144] for a skull in the Antwerp Museum, excavated in superficial formations of the Lower Rhine region, supposed to belong a juvenile aurochs, just as skulls from musea in Bruxelles and Danzig (ADDIN RW.CITE{{250 La Baume, W. 1909; 248 La Baume, W. 1922}}[112,145], see B.p. minutus), but identified as female aurochs crania ADDIN RW.CITE{{65 Leithner, O.F 1927/a;}}[78]. Bos urus primigenius Melnyk, 1927 ADDIN RW.CITE{{165 Melnyk, O. 1927}}[146] for the ancestor of East-European domestic cattle of primigenius type with a large and proposed to have been influenced by Bos namadicus or banteng. Bos urus priscus Schlotheim, 1820 ADDIN RW.CITE{{252 Szalay, A.B. 1930}}[69] for B. primigenius. Bos urus scoticus Wagner, 1837 ADDIN RW.CITE{{157 Wagner, A.J. 1837}}[94] after Urus scoticus for Chillingham cattle. Bos velanus Robert, 1930 ADDIN RW.CITE{{169 Duerst, J.U. 1931}}[86] ADDIN RW.CITE{{252 Szalay, A.B. 1930}}[69] for a supposed diluvial aurochs subspecies. Bos vulgaris Wagner, 1937) ADDIN RW.CITE{{157 Wagner, A.J. 1837}}[94] no description given. Bos zebu Boddaert, 1785 ADDIN RW.CITE{{116 Wilson, D.E. 1993; 253 Boddaert, P. 1784}}[59,121] early name for humped cattle, also used by Blandford ADDIN RW.CITE{{198 Blanford, W.T. 1870/a; 149 Lydekker, R. 1898}}[70,147]. Bos Zebu africanus Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67] for East- and West-African zebus. Bos Zebu africanus aethiopicus Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67] for Ethiopian zebu. Bos Zebu africanus hottentottus Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67] for cattle owned by the South-African Khoi herders (Hottentots). Bos Zebu africanus Sanga Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67] for the Raya-Azeb breed, formerly named Galla breed, synonym of Bos sondacus africanus and Bos sondacus longicornis. Bos Zebu Indicus major Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67] for large Indian zebu. Bos Zebu indicus medicus Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67] for medium sized Indian zebu. Bos Zebu indicus minor Fitzinger, 1860 ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67] for miniature zebu. Uri sylvestres Bos sylvestres (q.v.). Uro ibrico, postulated subtype of the B.p. primigenu ADDIN RW.CITE{{83 Miranda do Vale, J. 1907}}[115] as ancestor of the Maronesa breed. Urus Smith, 182) ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] for aurochs, already mentioned by Johnston (1651) ADDIN RW.CITE{{38 Duerst, J.U. 1931}}[93]. Urus colossus Keferstein, 1834 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] also for aurochs remains. Urus fossilis Keferstein, 1834 ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] also for aurochs remains. Urus nostras Bojanus, 1827 ADDIN RW.CITE{{145 Bojanus, L.H. 1827}}[60] or Uro nostrate for an assumed cattle species from the pre world, having thrived in the same period as the Elephas primigenius and Rhinoceros antiquitatis, and not having influenced modern taurina (cattle). Urus priscus Bojanus, 1827 ADDIN RW.CITE{{145 Bojanus, L.H. 1827}}[60] as Bos taurus priscus ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] identified later as the extinct steppe wisent. Urus scoticus Smith, 1827 ADDIN RW.CITE{{116 Wilson, D.E. 1993; 243 Bell,T. 1874; 254 Cuvier, M. 1827; 149 Lydekker, R. 1898}}[59,70,132,138] for White urus, British white park cattle supposed to be the probable remains of the genuine urus .However, Lydekker ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70] already recognized that the feral park cattle descended from domestic cattle. Zebus, Zebus gibbosus Blyth, 1860 ADDIN RW.CITE{{193 Blyth, E. 1860; 149 Lydekker, R. 1898}}[70,135] for zebu as a separate genus, besides the genus Bos. First mentioned as Zebus by Blyth, who supposed the zebu had its origins in Africa and came to Asia as domestic cattle, but was still found as a wild species in India, Ceylon and Africa. APPENDIX 2. ALPHABETICAL LIST OF AUTHORS WITH THEIR THEORIES Adametz, L. (1895) ADDIN RW.CITE{{64 Adametz, L. 1895}}[124] Teacher at the Austrian Hochschule fr Bodenkultur in Vienna. He stated that early shorthorned cattle (Torfrind) as described by Rtimeyer represented a genuine European wild form and gave it the name Bos europaeus (brachyceros). He also accepted the polled Akeratos type of Arenander as a separate type and emphasized the strict separation of European and Asian types of the Taurina. In 1920 ADDIN RW.CITE{{161 Adametz, L. 1920}}[108] he compared crania of Egyptian Apis bulls with those of modern cattle and applied the term Bos primigenius var. Hahni Hilzeheimer for this type, which he believed to have entered Africa as domestic cattle from Asia and to be the ancestor not only of North, East- and South-African domestic cattle, but also of several European breeds from Spain, France, Great Britain, Switzerland and Austria. Alderson, L. (1977, 1992) Prominent in the rare breeds movement, Technical Advisor to the Rare Breeds Survival Trust and author of several books on the subject. He designed a chart with three branches (Iberian, Scandinavian and Germanic) to clarify a putative origin of British and related breeds, related to pre-historic and historic immigrations of peoples and their cattle (ADDIN RW.CITE{{86 Alderson, L. 1977}}[148]; cf. ADDIN RW.CITE{{78 McKenny Hughes, T. 1896}}[149]). In 1992 he categorized European breeds into four main groups: Podolic, Northern European, Central European and Western European ADDIN RW.CITE{{108 Alderson, L. 1992}}[36]. Amschler, J. (1956) classified on the basis of the hypothesis of two main stem forms (B.t. primigenius and B.t. brachyceros / B.t. longifrons), with mutual influence of the descendants of both types ADDIN RW.CITE{{68 Amschler, J.W. 1956}}[122]. Antonius, O. (1922) ADDIN RW.CITE{{179 Antonius, O. 1922}}[110] Teacher at the same high school as Adametz. He stated that systematic morphological comparison of domestic animals species reveals the groups in which one has to look for the wild progenitor. He recognized two branches of taurine cattle, and as Rtimeyer believed that the aurochs formed the basis of only part of the domestic cattle classified as B t. primigenius, while B.t. brachyceros was the oldest type of domestic cattle in Europe. He observed that zebu occurs over a wide area and with various degrees of crossbreeding with taurine cattle. For him zebu is not the same as Buckelrind [humped cattle] but denotes all cattle with a zebu type skull with or without a hump. Aparicio, G. (1960) designed a phylogenetic tree of Spanish breeds in which each cluster of breeds is supposed to originate from hypothetical aurochs variant, such as B.t. ibericus, B.t. desertorum hispanico, B.braquiceros Europeo and B.braquiceros Africano ADDIN RW.CITE{{156 Aparicio, G. 1960}}[89]. Aristotle (350 BC) distinguished species by habitat and means of reproduction and divided animals into higher and lower classes ADDIN RW.CITE{{24 Aristoteles 2002}}[150]. Baker, C.M.A. & Manwell, G. (1980) published two seminal papers on a biochemical classification of 196 breeds ADDIN RW.CITE{{22 Baker,C M. 1980; 21 Manwell,C. 1980}}[30,31]. Baron, R. (1928) Professor in zootechnics at the cole dAlfort in France. He applied three criteria: form, color, production type . He designed a diagram according to profile of body, head and type of horns as tool for classification ADDIN RW.CITE{{140 Baron, R. 1928}}[9]. The Coordonnes baroniennes [Barons arrangement] were followed in the Larousse Agricole ADDIN RW.CITE{{81 Chacrin, E. 1921}}[11]. Bojanus, L. (1827), supposed that only one wild form of cattle had lived in Europe, the wisent and that by the name urus Linnaeus had meant the wisent ADDIN RW.CITE{{145 Bojanus, L.H. 1827}}[60]. After an unindentified skeleton in the museum of Jena, Germany, mentioned by Goethe in 1822, Bojanus thought to describe a new prehistoric (antediluvialis) species which he named Bos primigenius, not aware this was the historic aurochs, classified as Bos taurus by Linnaeus. Buffon, G.-L. Leclerc, Comte de (1749) French naturalist, mathematician, cosmologist and encyclopedic author. Buffon made the suggestion that species may have been both improvedand degenerated after dispersing from a centre of creation. In volume 14 he argues that all quadrupeds had evolved from an original set of just thirty-eight quadrupeds, qualifying as preDarwinina transformist ADDIN RW.CITE{{206 de Buffon, G.L.L. 1749/h;}}[151]. He described the Urus and Bison as two different species. Cheng, P (1986) designed a classification for Chinese cattle breeds according to regions and climatic zones ADDIN RW.CITE{{104 Cheng, P. 1986}}[152]. Curson and Thornton (1936) introduced the term sanga for African cattle characterized by a cervico-thoracic placed hump and long, lyre-shaped horns ADDIN RW.CITE{{214 Curson, H.H. 1934}}[153]. Cuvier, G.L.C.F.D. (1823) Founder of paleontology, a student of Lamarck and in 1795 his successor as professor of comparative anatomy at the Jardin des Plantes. He used anatomy to classify animal species by skeletal structures. Cuvier observed little difference between fossilized skulls of the pre-world Bos primigenius and those of modern cattle and stated that all domestic cattle are descending one Urus (aurochs) ADDIN RW.CITE{{196 Cuvier, M. 1823}}[142]. However he confused the European bison (wisent) with the aurochs. Darwin, C.R. (1859) Founder of the evolution theory ADDIN RW.CITE{{76 Darwin, C. 1859}}[154]. On the authority of Blyth, Darwin believed that zebu descended from a different source than European cattle. Blyth, who observed the different habits, voice, constitution and structure of humped Indian cattle, later classified the zebu in a separate genus besides genus Bos: Zebus gibbonus. Dechambre, P. (1913), professor at the same agronomic school at Grignon as Sanson and at Lcole vtrinaire dAlfort. He followed the classification developed by his colleague veterinarian Raoul Baron (q.v.). As second criterion for classification Dechambre used coat color ADDIN RW.CITE{{80 Dechambre, P. 1913}}[10]. Diffloth, P. (1914) Ingnieur agronome and professeur spcial dAgriculture at the veterinary school in Brussels. He adopted Sansons brachycphale / dolichocphale typing, recognizing twelve European cattle populations strictly according to geographic region ADDIN RW.CITE{{114 Diffloth, P. 1914}}[7]. Doutresoulle, G. (1947) Veterinarian and Chef du Service de l Elevage du Sudan. [director breeding services in French Sudan]. Doutresoulle described and classified the cattle breeds of colonial French West Africa according to region and climate zones and classed cattle in two main groups: the taurine breeds and short-horned, long-horned and sanga-type zebus (intermixed with taurine cattle) ADDIN RW.CITE{{217 Doutresoulle, G. 1947}}[23]. Drst, J.U. (1899) examined a large number of ancient and modern crania from Europe, Egypt and Mesopotamia. In his Inaugural-Dissertation ADDIN RW.CITE{{158 Drst, J.U. 1899}}[92] he recognized two additional basic forms which were thought to have been evolved by modification from the earliest cattle of the world: the polled Bos taurus akeratos as introduced by Arenander ADDIN RW.CITE{{66 Arenander, E.O 1898}}[72], and longhorned B.t. macrceros for African and Iberian cattle which he believed to be related. He agreed with Dawkins ADDIN RW.CITE{{189 Dawkins, W.B. 1866}}[126] and Nehrling ADDIN RW.CITE{{203 Nehring, A. 1888}}[127] that brachyceros cattle represented an impoverished form of the large aurochs and advocated the monophyletic domestic cattle descent theory. He proposed that the burden of the horn weight determines the shape of the skull and presented five different outlines of basic types. Furthermore he recognized three Pleistocene aurochs varieties: B. namadicus, B. opithonomus and B. primigenius. Epstein, H. initiated in 1933 the classification of African cattle, worked out further by Curson and Epstein (1934) ADDIN RW.CITE{{214 Curson, H.H. 1934}}[153]. With Mason ADDIN RW.CITE{{182 Epstein, H. 1984}}[155] he did in 1984 not recognize the namadicus as a separate species and emphasized the geographical range is the basis of the classification. They followed Duerst (1931) in distinguishing three aurochs subspecies according to continental origin: B. p. primigenius for the European Aurochs; B. p. opisthonomus for the North-African aurochs and B. p. namadicus for the Asian aurochs ADDIN RW.CITE{{182 Epstein, H. 1984}}[155]. European Cattle Genetic Diversity Consortium (from 2002). Participants and collaborators of the EU project Towards a strategy for the conservation of the genetic diversity of European cattle (19992002), coordinated by J.A. Lenstra, Faculty of Veterinary Medicine, Utrecht University. This consortium is listed as coauthor of several publications on the genetic diversity of cattle ADDIN RW.CITE{{219 European Cattle Genetic Diversity Consortium 2006; 87 Lalo, D. 2010; 3 Groeneveld,L.F. 2010; 225 Withen,K.B. 2011; 7 Edwards,Ceiridwen J. 2011}}[40,41,55,156,157]. Falconer, H. (1868) In the Catalogue of the Fossil Remains of Vertebratra in the Museum of the Asiatic Society of Bengal, Calcutta, the name Bos namadicus Falconer appears first. Falconer published in Palaeontological Memoirs (1868) on fossils excavated in the Siwalik Hills of northern India, where the oldest remains of B. namadicus were found ADDIN RW.CITE{{170 Falconer, H. 1868}}[95]. Fernndez, A. et al. (1998) analyzed 11 blood proteins of 10 different breeds from Galicia and northern Portugal ADDIN RW.CITE{{2 Fernndez, A. 1998}}[33]. Some of the breeds were also include in Vallejos survey, but with different outcomes. Like Vallejo ADDIN RW.CITE{{62 Vallejo, M. 1990}}[32] the breed clusters according to the molecular markers were linked to prehistoric forms denoted by Latin names. Fitzinger, L.J. (1860) Naturalist who proposed that domestic cattle consisted of at least seven geographic based forms ADDIN RW.CITE{{89 Fitzinger, L.J. 1860}}[67]. Felius, M. (1995) based the classification of cattle breeds from all over the world according to geographical, historical and morphological data by combining the information by early authors as well as later prominent authors on classifying cattle breeds, by personal communications and by travelling ADDIN RW.CITE{{30 Felius, M. 1995}}[38]. After classifying first 470 breeds into 16 groups ADDIN RW.CITE{{218 Felius, M. 1985}}[158] the 1995 classification emphasizes more the continental and geographic location. The coverage was extended to 700 breeds and supported by pictures. Hansen, J. (1927) described breeds according to region, per region into external type as well as after purpose: high productive dairy, beef and dual-purpose; lowland or highland; or low productive triple-purpose land cattle type ADDIN RW.CITE{{93 Hansen, J. 1927}}[21]. This led to a classification into 14 main groups, the first of which consisting of German, Dutch, Danish and Swiss cattle. Hengeveld, G.J. (1865) Veterinarian who was the first to described Dutch cattle in detail, stating that the cattle of the Netherlands belong to one breed only ADDIN RW.CITE{{57 Hengeveld, G.J. 1865}}[119]. He classified the various locally adapted varieties of the Dutch breed according to type of soil on which they were found: rich, medium or poor soil, after which he described the local types per province. Hilzheimer, M. (1909, 1917, 1926) Division director of the Marker Museum in Berlin. Considered the Bos primigenius as the one and only aurochs, though consisting of several subspecies, such as the B.p. Hahni (Hilzheimer, 1917) ADDIN RW.CITE{{208 Hilzheimer, M. 1909; 177 Hilzheimer, M. 1917; 67 Hilzheimer, M. 1926}}[83,107,159]. Hodges, J. & Payne, W.J.A. (1997) classified breeds according to the continent of their recent origin, within continents on a regional basis and within regions on a breed type basis. They differentiated humpless (Bos taurus), humped (Bos indicus) breeds, taurindicine crossbred and Southeast-Asian bibos breeds. The crossbred breeds have been further subdivided into three groups, stabilized, intermediate and recent crossbreds, respectively ADDIN RW.CITE{{46 Hodges, J. 1997}}[160]. Another subdivision is according to type of horn, purpose or region. Holecek Holleschowitz, C. (1939) recognized two main stem forms from which domestic cattle originated, the genuine Ur (Bos primigenius) and the smaller, shorthorned species (Bos europaeus brachyceros). However, he recognizes three basic types of European cattle: a western European variety, a northern subtype and the genuine aurochs ADDIN RW.CITE{{113 Holecek Holleschowitz, C. 1939}}[4]. He then also presented four groups of breeds, tracing to the aurochs, shorthorned cattle, short-headed cattle and polled cattle, respectively. Joshi, N.R. & Phillips, R.W. (1953) adopted the classification of cattle of India and Pakistan of Olver ADDIN RW.CITE{{102 Olver, A. 1938}}[161] and recognized six morpholical/regional groups ADDIN RW.CITE{{103 Joshi, N.R. 1953}}[27]. Joshi, N.R., Phillips, R. & McLaughlin, E.A.M. (1957) arranged African cattle geographically and then morphologically ADDIN RW.CITE{{97 Joshi, N.R. 1957}}[25]. Keller, C. (1902, 1905) Professor in zoology in Zrich. He recognized a number of basic domestic cattle forms according to cranium and type of horns, but also considered coat coloration and geographic origin ADDIN RW.CITE{{121 Keller, C. 1902; 41 Keller, C. 1905}}[3,68]. He developed breed groups and classified according to their supposed origin: Primigenius type cattle descending from European aurochs and shorthorned cattle of Oriental (Asian) origin. He was of the opinion that the brachyceros type traced to the Asian zebu (which in turn descended from the banteng or Bali cattle) but had lost its hump, and therefore named it Bos sondaicus brachyceros. Linnaeus, C. (1735, 1758) Swedish naturalist, catalogued eighteen thousand species and classified individual species, which he then arranged according to their similarities ADDIN RW.CITE{{25 Linnaeus 1735; 117 Linnus, C. 1758}}[61,162]. He introduced the binary nomenclature (genus name plus species name) and a definitive concept of species in biology. Linnaeus introduced the name Bos taurus for humpless cattle (aurochs and domestic) and Bos indicus for the zebu. Linnaeus also used the name Urus. Lydekker, R. (1898), British naturalist, dismissed the common believe that the half-wild Chillingham and other British park cattle are authentic wild animals. He proposed the name Bos taurus var. longifrons for domestic taurine cattle. Lydekker believed that the namadicus is closely allied to the European wild ox, initially presuming it to be the ancestor of the bibovine group. He noted that nothing was known about the ancestry or original habitat of humped cattle ADDIN RW.CITE{{149 Lydekker, R. 1898}}[70]. In 1912 he described the history of the extermination of the aurochs ADDIN RW.CITE{{173 Lydekker, R. 1912}}[102]. Mason, I.L. classified in 1951 the breeds of West Africa according to Doutresoulle, but in a more specified way as he divided the taurine cattle into longhorned Chad cattle and small (NDama) cattle; and humped cattle into short-horned, medium-horned, lyre-horned and long-lyre-horned zebus ADDIN RW.CITE{{95 Mason, I.L 1951}}[24]. Original author of the standard work Masons world dictionary of livestock breeds, types and varieties, which lists names and synonyms of farm animals ADDIN RW.CITE{{231 Porter, V. 2002}}[163]. With Maule he refined in 1960 their classification of West-African and East- and South-African humped cattle, respectively, recognizing the form and place of the hump and the form and length of the horns and arriving at 14 Sanga, 9 zebu and 5 intermediate breed types ADDIN RW.CITE{{100 Mason, I.L. 1960/h;}}[164]. Maule, J.P. (1990) classified tropical cattle into five groups: zebu, sanga, humpless, humped humpless and Bibovine cattle, further subdivided according to locality (region, continent or country) or to breed and regional type ADDIN RW.CITE{{106 Maule, J.P. 1990}}[29]. McKenny Hughes, T. (1896) proposed in an extensive theory on the origin of several British breeds. He considered Kerry cattle as the most typical Celtic short-horn of the British Isles, the Chillingham as the most close to the cattle introduced by the Romans, Highland and Welsh cattle as a mixture of the Roman breed with the Celtic shorthorns and the longhorns as offspring from the breeds imported from Holstein and the Low Countries in the late Middle Ages ADDIN RW.CITE{{78 McKenny Hughes, T. 1896/h;}}[149]. Miranda Do Vale, M.J. (1907/1949) adhered to the twelve basic ethnic types troncos of Sanson (1893), such as Bos taurus aquitniucus, Bos taurus ibricus and B.t. atlanticus ADDIN RW.CITE{{83 Miranda do Vale, J. 1907}}[115]. Mller, W. (1957) divided Austria into three different historical zones: Rheatian, Norik and Pannonica ADDIN RW.CITE{{82 Mller, W. 1957}}[12]. To each zone an ethnic group was considered endemic, each owning a specific type of cattle: Western unicolored, Central pied group and Eastern unicolored breeds. Olver, A. (1938) Animal husbandry expert of the Imperial Council of Agricultural Research in British India. Olver related the different types of zebus on the Indian subcontinent to migrations of peoples into India, in pre-historic times, as along the various migration routes characteristic zebu types are to be found, some of which must have been in existence prior to these invasions ADDIN RW.CITE{{102 Olver, Arthur 1938/h;}}[161]. Ramm, E. (1901) Teacher at the Agricultural Academy in Bonn am Rhein. He applied a more practical inventory than a rigorous zoological classification, a geographical classification according to both country or region and level (highland vs lowland) ADDIN RW.CITE{{56 Ramm, E. 1901}}[19]. Rtimeyer, L. (1861, 1868) Professor of zoology in Basel and considered as the founder of domestic animal archaeozoology. He founded a classification within the Bovini tribe according to basic types of fossil as well as modern crania ADDIN RW.CITE{{34 Rtimeyer, L. 1868}}[79]. By examining extensive material from a Neolithic Swiss settlements he stated that shorthorned Neolithic domestic cattle, Bos taurus brachyceros represented the oldest form of cattle from which all other forms had evolved. He believed that B.t. brachyceros descended from an Asian ancestor and was the progenitor of most European domestic cattle. In 1868 he recognized in the European Bos primigenius both the wild animal as its domestic (longhorned) taurine progeny, while in the Indian Bos namadicus (Falconer, 1859) ADDIN RW.CITE{{170 Falconer, H. 1868}}[95] he recognized all forms of humped cattle. He was not sure on the predecessor of the domestic zebu (Bos indicus) and suggested a close relationship to banteng and yak. He further proposed that Bos namadicus was the predecessor of Bos primigenius. Snchez Belda, A. (1981, 1984) combined skull, coat color and region of Spanish cattle and recognized three branches of Iberian cattle and a fourth group, directly related to North-African Atlas cattle ADDIN RW.CITE{{84 Sanchez Belda, A. 1981; 85 Sanchez Belda, A. 1984}}[13,14]. Sanson, A. (1878, 1893) Prof. in zoology and zootechnics at the National Agricultural School, Grignon and the National Agronomic Institute, Paris. Sanson introduced a scientific method for animal ethnology founded on the skull measurement theories of the anthropologists Retizius and Broca. He classified cattle according to their cranium, the form of the poll and horn implant, and the length and form of the horns ADDIN RW.CITE{{166 Sanson, A. 1878; 69 Sanson, A. 1884; 167 Sanson, A. 1893}}[6,73,74]. He linked these types to those of human migrants: Dolichocphale (longheaded) and Brachycphale (short-headed) people. In 1878 Sanson introduced many Latin Bos names for basic types, to which in 1893 the term, taurus was added in order to indicate their domestic status (e.g., Bos aquitanicus became Bos taurus aquitanicus). Simon, D.L. & Buchenauer, D. (1993) classified cattle breeds in 10 main groups, based mainly on coat color, but with subgroups according to color patterns, geographical origin and genetic relationship ADDIN RW.CITE{{55 Simon, D.L. 1993}}[17]. Vallejo, M. et al. (1990) applied a biochemical approach to Iberian cattle, linking breed clusters of breeds to presumed origin and supposed prehistoric forms (ADDIN RW.CITE{{62 Vallejo, M. 1990}}[32] cf. ADDIN RW.CITE{{156 Aparicio, G. 1960; 84 Sanchez Belda, A. 1981; 85 Sanchez Belda, A. 1984}}[13,14,89]). Van Leithner, O. (1927) compared and measured a large number of aurochs crania. He recognized large (pre-world) diluvial aurochsen (Bos trochoceros) and smaller alluvial aurochsen (Bos primigenius), each containing of several subtypes ADDIN RW.CITE{{65 Leithner, O.F 1927}}[78]. He also concluded that African aurochs crania (such as B p. mauritanicus ADDIN RW.CITE{{180 Thomas, P. 1881}}[111], B. opisthonomus (Pomel, 1893) B p. Hahni ADDIN RW.CITE{{177 Hilzheimer, M. 1917}}[107] were not to be distinguished from the crania of the European Bos primigenius. He delivered convincing evidence of the pronounced sexual dimorphism in the aurochs, which excluded the idea of a small type of aurochs as proposed by Pohlig ADDIN RW.CITE{{151 Pohlig, H. 1911}}[80]. Werner, H. (1912) Teacher at the Agricultural Highschool in Berlin. In his Practical Handbook he elaborated upon the classification of Wilckens by a detailed regional subdivision in Rasse (Types) and Unterrasse (Subtypes) ADDIN RW.CITE{{37 Werner, A.H. 1912/h;}}[2]. Wilckens, M. (1876) Teacher at the Highschool for Soil Science in Vienna. He rejected the idea that Bos primigenius was the ancestor of domestic cattle ADDIN RW.CITE{{29 Wilckens,M. 1905}}[165]. He proposed the first classification of domestic cattle breeds, based on measurements of crania: B. primigenius, B.t. brachyceros, B.t. frontosus and B. t. brachycephalus. Wilson, J. (1909) Professor at the Royal College of Science Dublin. He followed McKenny Hughes ADDIN RW.CITE{{78 McKenny Hughes, T. 1896}}[149] with even stronger, though equally unfounded ideas on the relation between different ethnics which entered the British Isles in course of time and the color of the cattle they introduced: Black Celtic, White Roman, Red Anglos-Saxon, Dun (polled) Scandinavian and flecked Dutch. ADDIN RW.CITE{{45 Wilson, J. 1909/h;}}[131]. Youatt, W. (1834) Veterinarian, the first to describe British breeds in detail. Classified the British breeds on basis of their horns while disregarding the rest of the cranium ADDIN RW.CITE{{207 Youatt, W. 1834}}[166]. Zeuner, F.E. (1963) believed that the so called Torfrind (Bos longifrons) descended from the Bos primigenius and believed that in several modern breeds the primigenius or longifrons type was relatively well kept, although most current breeds are of mixed origin ADDIN RW.CITE{{148 Zeuner, F.E. 1963; 43 Zeuner, F.E. 1963}}[106,133]. References ADDIN RW.BIB1. Wilckens, M. Die Rinderrassen Mittel-Europas; Wilhelm Braumller: Wien, Austria, 1876. 2. Werner, A.H. Die Rinderzucht; Paul Parey: Berlin, Germany, 1912. 3. Keller, C. Naturgeschichte Der Haustiere; Paul Parey: Berlin, Germany, 1905. 4. Holecek Holleschowitz, C. Angewandte Tierzucht Auf Rassenbiologischer Grundlage; Julius Springer: Vienna, Austria, 1939. 5. Haring, F.; Hammond, J.; Johannson, I. Handbuch Der Tierzchtung; Paul Parey: Hamburg-Berlin, Germany, 1961. 6. Sanson, A. 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